Episcaphula (Cosmoscaphula) tamburinea Heller, 1920

Skelley, Paul E., Leschen, Richard A. B. & Liu, Zhenhua, 2021, Nomenclatural notes for some Australian Erotylinae (Coleoptera: Erotylidae), Zootaxa 4966 (1), pp. 69-76 : 71-72

publication ID

https://doi.org/ 10.11646/zootaxa.4966.1.7

publication LSID

lsid:zoobank.org:pub:31606F03-C533-4329-BBD4-596B58A491E6

DOI

https://doi.org/10.5281/zenodo.4729461

persistent identifier

https://treatment.plazi.org/id/F76FA665-5061-FF90-F192-F98EFDCAF803

treatment provided by

Plazi

scientific name

Episcaphula (Cosmoscaphula) tamburinea Heller, 1920
status

 

Episcaphula (Cosmoscaphula) tamburinea Heller, 1920

Episcaphula (Cosmoscaphula) tamburinea Heller 1920: 105–106 . Chûjô & Chûjô 1988: 161.

Episcaphula (Cosmoscaphula) clatrata Heller 1920: 106 , new synonymy. Chûjô & Chûjô 1988: 161.

Episcaphula rufolineata Wilson 1921: 35 , new synonymy.

Episcaphula (Episcaphula) rufolineata Wilson 1921: 35 . Chûjô & Chûjô 1988: 165 (subgenus combination).

Comments. Heller (1920) published a large work on the Erotylidae of the Indo-Australian region with an emphasis on the Philippines. The Australian taxa described in this work have been overlooked by previous Australian workers. Some need clarification and nomenclatural corrections.

The subgenus Cosmoscaphula Heller (1920: 105–106) is recognized as an Episcaphula Crotch, 1876 with an enlarged lobe or callus between the eye and antennal base. Heller (1920) placed two species in the subgenus: E. (C.) tamburinea Heller, 1920 and E. (C.) clatrata Heller, 1920 , stating that E. clatrata may be a “var.?” of E. tamburinea . Both species were from Tamborine Mountain, near Brisbane, Queensland, Australia. The following year, Wilson (1921) described Episcaphula rufolineata , also from Tamborine Mountain, noting the same lobe in front of the eyes, sexual dimorphism, and variation in color pattern.

While Heller (1920) only discussed one specimen for E. clatrata (SDEI, Figs. 3–4 View FIGURES 1–7 ), he mentioned two depositories for E. tamburinea . Since his types are generally deposited in SDEI, the male specimen (SDEI, Figs. 1–2 View FIGURES 1–7 ) with the following label data is here designated as the lectotype for Episcaphula (Cosmoscaphula) tamburinea Heller, 1920 : “[white paper, hand-written] S. Queensland / Mt. Tambourine ” // “[red paper, typed] Syntypus ” // “[white paper, hand-written] Coll. Hacker ” // “[white paper, hand-written] Episcaphula / ( Cosmoscaphula ) / tambourinea n. / [typed] Det. K M Heller 1918” // “[lavender paper, black border, hand-written] tam- / bourinea / Hell.” // “[white paper, typed] SDEI Coleoptera / # 302836” // “[red paper, typed] LECTOTYPE ♂ / Episcaphula / ( Cosmoscaphula ) / tamburinea Heller / des. Skelley, / Leschen, Liu 2020” . Paralectotypes may exist but were not studied.

Wilson (1921) discussed multiple specimens of E. rufolineata from multiple localities, but stated “Type in author’s collection”, explicitly indicating that only a single specimen was used to describe the species. The Museums Victoria website hosts an image of this holotype ( Figs. 5–7 View FIGURES 1–7 ; Hoath 2020a, MVMA).

Examination of these images clearly show Heller’s types represent a male and female with extremes of the color pattern variation as discussed by Wilson (1921). Males of the species have a notched clypeal margin, dilated protarsomeres, a slightly more elongated pronotum, and more strongly curved tibiae.

Because they are from the same locality and share all morphological characters, it is clear they are conspecific and are here synonymized. With Heller considering E. clatrata as a possible variety of E. tambruinea and since evidence is lacking to invoke prevailing use rules to maintain E. rufolineata ( ICZN 1999: Article 23.9.1), we use the name E. tamburinea for this species.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Erotylidae

Genus

Episcaphula

Loc

Episcaphula (Cosmoscaphula) tamburinea Heller, 1920

Skelley, Paul E., Leschen, Richard A. B. & Liu, Zhenhua 2021
2021
Loc

Episcaphula rufolineata

Wilson, F. E. 1921: 35
1921
Loc

Episcaphula (Episcaphula) rufolineata

Chujo, M. & Chujo, M. 1988: 165
Wilson, F. E. 1921: 35
1921
Loc

Episcaphula (Cosmoscaphula) tamburinea

Chujo, M. & Chujo, M. 1988: 161
Heller, K. M. 1920: 106
1920
Loc

Episcaphula (Cosmoscaphula) clatrata

Chujo, M. & Chujo, M. 1988: 161
Heller, K. M. 1920: 106
1920
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