Arcyphysa angusta

Wells, Alice & Contents, Arturs Neboiss Table Of, 2018, Australian Diplectroninae reviewed (Insecta: Trichoptera), with description of 21 new species, most referred to a new genus, Zootaxa 4415 (1), pp. 1-44 : 25-27

Arcyphysa angusta Group

( Figs 104–105 , 146, 158–159 , 179–180)

Arcyphysa angusta (Banks)

Diplectrona angusta Banks 1939 , 495; Mosely & Kimmins 1953, 337.

Material examined. Holotype ♂, Dorrigo, NSW ( ANIC, ex MCZ 22087); 1♂, 2♀, NSW, Styx River, 12 km S of Ebor, 17.x.1973, A. Neboiss ( NMV [PT-1005]).

Diagnosis. This species resembles Arcyphysa diamontona sp. nov. in having the gonopods in ventral view widely bowed, but is distinguished from Arc. diamontona by having dorsal processes clearly arising on abdominal tergite IX and lanceolate, not club shaped, and by lacking sharp spines apicolaterally on sternite IX; harpagones are leaf-like, rounded apically, not acute as in Arc. diamontona .

Description. Length of each forewing: ♂ 10.0 mm (n = 2), ♀ 10.4 mm (n = 2); forewing median cell length about 5x maximum width.

Male. Abdomen with internal reticulate-walled internal sacs all elongate oval, about length of segments.

Male genitalia ( Figs 104–105 , 158–159 ): Segment IX short, apicolaterally expanded distally to form subtriangular lobe; apicolateral processes elongate, lanceolate, about twice length of tergite X, in ventral view with convergent club-shaped lobes densely clothed in short setae; tergite X compressed, taller than long; phallic apparatus without parameres, in ventral view wide at base, in lateral view with apex curved downward; gonopods each with coxopodite stout, strongly curved in ventral view, width almost equal throughout length, harpagones leaf-shaped, about 1/3 length of coxopodites, curved gently, tapered to rounded apices.

Female ( Figs 179–180 ). Abdomen terminating bluntly; distally with pair of cerci and 2 pairs of small terminal protuberances; sternite VIII formed by 2 ventrolateral plates (valves) covering inferior appendage receptacle grooves, fully divided mesally, apicolateral angles produced, broadly rounded; segment IX with rounded sclerotic cavity, its inner surface granular.

Distribution. Collected only from north-eastern New South Wales.

Remarks. Banks (1939, 495) interpreted the paired apicolateral structures in Arc. angusta as part of the ‘superior plate’. Here these structures are interpreted as apicolateral processes on abdominal tergite IX. The arrangement in Arcyphysa angusta contrasts with the arrangement in Arc. diamontona, Arc. hystricosa , and Arc. fraserensis in which abdominal segment IX and tergite X appear to be fused, with the result that the apicolateral lobes appear to arise from the fused structure.

Figures

FIGURES 102–109. Arcyphysa gen. nov. spp., diagnostic features.102–103, Arc.diamontona sp. nov., Vic., Diamond Creek, 7 km SE of Gembrook [PT-1006], ♂: 102, right fore- and hind wings, dorsal, 103, maxillary palp. 104–109, ♂ genitalia. 104– 105, Arc. angusta (Banks 1939; NSW, Styx R. [PT-1005]): 104, ventral, and 105, left lateral. 106–107, Arc. diamontona sp. nov. [PT-1006]:106, ventral, and 107, left lateral.108–109, Arc.fraserensis sp. nov. (Qld, Fraser Island, Pile Valley, nr Central Station [PT-1125]):108, ventral,and 109, leftlateral.Scale bars: Fig.102 = 1 mm; Fig.103 = 0.5 mm.Abbreviations:ac= anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; cox = coxopodite; fI, fII, fIII, fIV, fV = forks I–V; mc= median cell; har = harpago; ph = phallic apparatus; IX, X = abdominal segments IX, X

FIGURES 146–163. Arcyphysa gen. nov. spp., ♂ diagnostic features. 146–150, heads, dorsal: 146, Arc.angusta (Banks 1939; [PT-1005]); 147, Arc. carnarvona sp. nov. (Qld, Carnarvon Gorge); 148, Arc. satana (Mosely 1953; N Qld, Crystal Cascades, Trib. Crystal Creek); 149, Arc. diamontona sp. nov. [PT-1047]; 150, Arc. hystricosa (Neboiss 1979; Qld, North Stradbroke Island [PT-512]). 151, Arc. diamontana (Vic., Diamond Creek, 7 km SE of Gembrook), ♂ right fore- and hind wings, dorsal. 152–153, Arc. hystricosa [PT-512]: 152, antennal segments; 153, maxillary palp. 154–163, ♂ genitalia. 154–155, Arc. hystricosa [PT-512]: 154, ventral, and 155, left lateral; 156–157, Arc. satana (N Qld, Crystal Cascades, Trib. Crystal Creek): 156, ventral, and 157, left lateral; 158–159, Arc. angusta (NSW, Styx R. [PT-1005]): 158, ventral, and 159, left lateral; 160– 161, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]): 160, ventral, and 161, left lateral; 162–163, Arc. diamontona [PT-1047]:162, ventral, and 163, left lateral. Scale bar = 2 mm. Abbreviations: ac = anal cell; alp = apicolateral process; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; dalw = divided anterolateral wart; cox = coxopodite; mc = mediancell; har = harpago; ph = phallic apparatus; pw = posterior wart; X = abdominal segment X.

FIGURES 179–187. Arcyphysa gen. nov. spp., diagnostic features. 179–182, ♀ terminal abdominal segments: 179–180, Arc. angusta (NSW, Styx R. [PT-1005]): 179, ventral, and 180, left lateral; 181–182, Arc. diamontona sp. nov. [PT-1006]: 181, ventral, and 182, left lateral. 183–187, ♂ reticulate-walled internal sacs of abdominal segments VI and VII: 183, Arc. tortula sp. nov. (Qld, Bellenden Ker Range, Cable Tower No. 3 [PT-1031]), ventral; 184–185, Arc. acmea sp. nov. (Qld, Mt Bartle Frere,0.5 km Nof South Peak [PT-999]):184, rightlateral, and 185, ventral;186, Arc.anaplasis sp.nov. (Qld, 25 km along Mt Lewis Road, SW of Mossman [PT-1003]), ventral; 187, Arc. candela sp. nov. (NSW, Brown Mountain [PT-1017]), ventral. Abbreviation:VIII = abdominal segment VIII.