Exocelina erteldi (Balke, 1998)

Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo & Balke, Michael, 2019, Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 878, pp. 73-143 : 73

7. Exocelina erteldi (Balke, 1998) Figs 14 , 20

Copelatus (Papuadytes) erteldi Balke, 1998: 330; Nilsson 2001: 76 (catalogue).

Papuadytes erteldi (Balke, 1998): Nilsson and Fery 2006: 56 (comb. nov.).

Exocelina erteldi (Balke, 1998): Nilsson 2007: 33 (comb. nov.).

Type locality.

Indonesia: Papua Province: Pegunungan Bintang Regency, Borme, ca. 04°24'S, 140°25'E, 1200 m a.s.l.

Type material studied.

Holotype: male "IRIAN JAYA Zentralmassive 140°25'E 04°24'S ", "14./17.8.1992 Borme, 1900m leg. Balke (11)", " Copelatus erteldi Balke des. 1997" [red], “HOLOTYPUS” [red] (NHMW). Paratypes: 1 female with the same label as the holotype and additionally with a red label "Paratypus Copelatus erteldi Balke des. 1997" (NHMW). 54 males, 35 females "IRIAN JAYA Zentralmassive 140°25'E 04°24'S ", "Borme, 1800m 16.8.1992 leg. Balke (12, 12 A)", one of the males with two additional labels "M.Balke 3273" [green] and "M.Balke 6404 DNA" [green text], another male with an additional green label "M.Balke 3273" (CGW, NHMW). Note: in the original description ( Balke 1998), number of specimens of the locality (12, 12 A) is erroneously given as "43 males, 46 females". 1 female "12./18.8.1992 Borme, 100m leg. Balke (7)", "Paratypus Copelatus erteldi Balke des. 1997", this paratyte does not belong to species of E. erteldi but to E. bifida Shaverdo et al. 2012.

Additional material.

1 male "IRIAN JAYA Zentralmassive 140°25'E 04°24'S ", "Borme, 1800m 16.8.1992 leg. Balke (12, 12 A)", "Paratypus Copelatus fume Balke des. 1997" [red] (NHMW).

Diagnosis.

For complete description, see Balke (1998: 330). Beetle small (TL-H 3.45-3.75 mm), oblong-oval; brown to piceous, usually with paler pronotal sides; dorsally more or less shiny, with fine but conspicuous punctation and weakly impressed microreticulation; pronotum with distinct lateral bead ( Fig. 14 ); male protarsomere 4 with anterolateral seta thin, weakly curved, smaller than more laterally situated large seta; male protarsomere 5 ventrally with anterior band of ca. 70 and posterior row of 6 relatively long setae ( Fig. 20D ); median lobe in lateral view evenly attenuated to elongate, thin apex, which slightly pointed in ventral view; paramere slightly concave on dorsal side and with distinct, long, dense, uniform setae, situated along dorsal margin ( Fig. 20 A–C ).

Affinities.

The species can be distinguished from the species co-occurring in the same area ( E. ascendens , E. aipomek , E. takime , the E. ekari group: E. eme Shaverdo and E. bifida , the E. danae group: E. damantiensis and E. danae , the E. okbapensis group: E. ketembang , E. talaki , and E. okbapensis , and all species of the E. aipo group) by body size and colouration, presence of pronotal bead, fine but conspicuous dorsal punctation, and shape and setation of its median lobe, paramere, and male protarsomere 4. For the affinities within the group, see the “Key”.

Distribution.

Indonesia: Papua Province: Pegunungan Bintang Regency, Borme ( Fig. 25 ). The species is known only from the type material.

Figures

Figure 25. Map of the eastern part of New Guinea showing distribution of the species of the E. bacchusi group.

Figures 12 - 17. Habitus and colouration 12 Exocelina oiwa sp. nov. 13 E. akameku sp. nov. 14 E. erteldi (Balke, 1998) 15 E. oksibilensis sp. nov. 16 E. bacchusi (Balke, 1998) 17 E. bacchusi herzogensis sp. nov.

Figures 20, 21. 20 Exocelina erteldi (Balke, 1998) 21 E. oksibilensis sp. nov. A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view.