Vetosmylus tentus, Ma & Shih & Ren & Wang, 2020

Ma, Yiming, Shih, Chungkun, Ren, Dong & Wang, Yongjie, 2020, A new genus of lance lacewings from the Middle Jurassic of Inner Mongolia, China, Acta Palaeontologica Polonica 65 (2), pp. 363-369 : 365-366

Vetosmylus tentus sp. nov.

Fig. 1 .

Zoobank LSID: urn:lsid:zoobank.org:act:30620AD1-7A74-4989-8F8A-81660E7449A5

Etymology: From Latin tentus , outstretched; refers to the spreaded four wings of the new species.

Holotype: CNU-NEU-NN2019001P/C, a completely preserved male specimen with fully spreaded wings and visible structures ( Fig. 1A 1 , A 2 ). Type locality: Daohugou Village , Shantou Township, Ningcheng County, Inner Mongolia, China.

Type horizon: Jiulongshan Formation, Aalenian–Bajocian boundary, Middle Jurassic.

Diagnosis.—Forewing with many scattered fuscous spots; CuP about three-fourths of CuA, forming neatly arranged pectinate branches; anal region well-developed, 2–3 cross-veins between A2 and A3; A3 single; hind wing speckless; cross-veins in radial sector arranged irregularly, only the outer gradate series present; A1 well-developed, parallel with posterior margin, pectinately branched.

Description.—Head laterally compressed and deformed; antenna filiform and incompletely preserved;presence of thorax and abdomen well-developed; four wings spreaded. Forewing about 28.18 mm long as preserved, 9.53 mm wide. Hind wing about 28.36 mm long as preserved, 8.59 mm wide.

Forewing ( Fig. 1A 3 , A 4): Elongate with rather rounded apex; nygmata undetected; pterostigma fuscous; trichosors well-defined along the entire margin; subcostal veinlets forked distally, becoming denser towards trichosors; crossvein sc-ra single and oblique; cross-veins ra-rp numerous, forming many square cells; RP with more than 15 branches, each broadly curved toward margin, with complicated distal forks; M forked close to the wing base, MA and MP forming several simple branches distally; CuA forked near the middle of the wing; CuP with 7–9 pectinate branches; A1 long, with 8–10 pectinate branches; A2 with 7 distal pectinate branches.

Hind wing ( Fig. 1E ): Without any large fuscous spots; trichosors well-developed along the entire margin; subcostal veinlets simple, with few forks; sc-ra single; presence of sigmoid vein joined the base of RP; the region between MA and MP expanded obviously, with one row of cells; MA forming simple distal branches; MP with 4–5 distal pectinate branches; CuA long, forming 9 pectinate branches; CuP with 10 pectinate branches; A2 and A3 poorly preserved.

Remarks.— Vetosmylus tentus is separated from the other species by the following characters: forewing with many fragmentary fuscous spots, and the hind wing immaculate rather than distributed with large spots; the outer gradate series present in forewing; MP forming several pectinate distal branches instead of simple branches in forewing.

Stratigraphic and geographic range.— Jiulongshan Formation, Middle Jurassic; Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.

Figures

Fig. 1. Lance lacewing Vetosmylus tentus gen. et sp. nov. (holotype, CNU−NEU−NN2019001P/C) from Aalenian–Bajocian boundary (Middle Jurassic) of Jiulongshan Formation of Daohugou, China. Photographs of part (A1) and counterpart (A2). Line drawings of left forewing (A3), right forewing (A4), left hind wing (A5). Abbreviations: A, anal veins; CuA/P, anterior/posterior cubitus; MA/P, anterior/posterior branch of media; RA/P, anterior/posterior branch of radius; Sc, subcosta.

Fig. 2. Lance lacewing Vetosmylus maculosus gen. et sp. nov. (holotype, CNU−NEU−NN2019002P/C) from Aalenian–Bajocian boundary (Middle Jurassic) of Jiulongshan Formation of Daohugou, China. Photographs of part (A1) and counterpart (A2). Line drawings of forewing (A3), hind wing (A4). Abbreviations: A, anal veins; CuA/P, anterior/posterior cubitus; MA/P, anterior/posterior branch of media; RA/P, anterior/posterior branch of radius; Sc, subcosta.

Fig. 3. Female lance lacewing Vetosmylus maculosus gen. et sp. nov. (paratype, CNU−NEU−NN2019003P/C) from Aalenian–Bajocian boundary (Middle Jurassic) of Jiulongshan Formation of Daohugou, China (A) and extant Parosmylus tibetanus Yang, 1988 (B). A. Photographs of part part (A1) and counterpart (A2); terminalia in lateral view, a combination of photos of part and counterpart under dry condition (A3) and line drawing (A4), sternite 8 and gonapophysis 9 cannot be clearly observed. B. Line drawing of terminalia in lateral view.