Uduba lakroa, Griswold & Ubick & Ledford & Polotow, 2022

Griswold, Charles, Ubick, Darrell, Ledford, Joel & Polotow, Daniele, 2022, A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar, Proceedings of the California Academy of Sciences 67, pp. 1-193 : 56-57

Uduba lakroa , new species

Figures 48 A–C , Maps 12, 21.

Type material. Holotype male ( CASENT9064782 ) from pitfall traps at 875 m elevation along Andrauorena River on the NE slope Mt. Papango in Parc National de Midousy, 23°50.1ʹS, 46°57.8ʹE, Fianarantsoa Prov., Madagascar, collected 26–31 October 2003 by L. E. Olsen [FMNHLEO128], deposited in FMNH. Three paratype males, same data, two deposited in FMNH ( CASENT9062274 and CASENT9062280 ) and one in CAS ( CASENT9062275 ).

Etymology. The species epithet is from a Malagasy word for cross, lakroa; a noun in apposition.

Diagnosis. Uduba lakroa are one of the Unclassified species ( Map 21 ), which have a divided cribellum. Males of U. lakroa can be distinguished from those of other Uduba by having the palpus with a swollen, dark lobe near the base of TA2 and swollen, unsclerotized lobe near the base of TA3 and the conductor ( Figs. 48 A–C ). Females are unknown.

Description. Male (Holotype): Total length 13.80. Markings typical of Uduba . Carapace 8.10 long, 5.60 wide, 3.40 high; clypeus 0.40 high. Eye diameters: AME 0.24, ALE 0.28, PME 0.26, PLE 0.28. Chelicerae 3.70 long; sternum 3.40 long, 2.60 wide; labium 1.70 long, 1.20 wide; palpal coxae 2.60 long, 1.00 wide. Cribellum divided. Spination (paratype, CASENT9062275): palpus–femur 1-1-0-1, p0-0-2, patella p0-1-0, tibia p1 [elongate, stout spine]-0-0; leg I–femur d1-1-1, p0-1-2, r0-1-1-1, patella p1, r1, tibia d0-1-1, p0-1-1-0, v2-2-2-2, r0-1-1-0, metatarsus p01-1-1, v2- 2-2, r1-1-0; leg II–femur d1-1-1, p0-1-2, r1-1-1-1, patella p1, r1, tibia d1-1-1, p0-1-1-0, v2-1-2-2- 2, r0-1-1-0, metatarsus p1-1-1, v2-2-2, r1-1-1; leg III–femur d1-0-1-1, patella p1, r1, tibia d1-1-1, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2; leg IV–femur d0-1-1-1, patella p1, r1, tibia d0-1-1, p0-1-1-0, v2-2-2, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1-1-2. Scopulae: cymbium, apicodorsal; tarsi I–IV, ventral. Leg measurements (combination of males CASENT9062274 and CASENT9062275): I: 6.70 + 2.80 + 6.20 + 6.10 + 3.60 = 25.40; II: 6.00 + 2.70 + 5.40 + 5.60 + 3.20 = 22.90; III: 5.60 + 2.30 + 4.10 + 5.50 + 3.00 = 20.50; IV: 6.70 + 2.50 + 5.70 + 7.30 + 3.40 = 25.60; palpus: 3.00 + 1.10 + 1.10 + NA + 3.30 = 8.50. Leg formula 4123. Male palp (paratype, CASENT9062275): palpal tibia 0.30 cymbial length, RTA very broad ( Fig. 48 C ), RTA length 0.27 tibia length, length 0.57 times width, with blunt, curved apex, VTA small, slender, curved ( Fig. 48 A ), length 1.60 times width, length 0.33 tibia width; tibia with an elongate, stout prolateral spine; tegulum convex, without ridge ( Figs. 48 B, C ), tegulum length 0.92 times width; TA1 extends to tegulum apex and slightly past TA3 apex; TA3 and TA2 both thick, curved, TA2 crosses in front of TA3, TA3 extends apicad of TA2 by 1.05 times tegulum length ( Fig. 48 B ), MA transverse, bilobed, origin at 0.47 times tegulum length, tegulum apex at 0.32 tegulum length from MA apex, MA length (including apical processes) 0.72 times width, MA large but narrowly transverse ( Figs. 48 B, C ), length 0.28 tegulum length, width 0.35 tegulum width; conductor fan entire. Female: Unknown.

Variation. Male (N= 4): Total length 13.00 to 14.50; ratios of carapace length / width = 1.40– 1.48, carapace height / width = 0.49–0.61, PER / carapace width = 0.37–0.40, PER / OAL = 3.03– 3.48, PER / AER = 1.32–1.39, OQP / OQA = 1.03–1.07, clypeal height / diameter AME = 1.25– 1.67, cheliceral length /clypeal height = 9.25–12.33, sternum length / width = 1.31–1.46; length femur I / carapace width = 1.25–1.37, metatarsus I /carapace width = 1.07–1.27, femur IV length / carapace width = 1.26–1.35, cymbium length / carapace width = 0.58–0.63, palpal tibia length / palpal patella length = 0.83–1.00, cymbium length / palpal tibia length = 2.75–3.00, cymbium length / palpal femur length = 1.00–1.11. Female variation is unknown: only males are recognized.

M aterial examined. MADAGASCAR: Fianarantsoa Province: Parc National de Midousy, NE slope Mt. Papango, along Andrauorena River , 2 km SW Befotaka, 23°50.1ʹS, 46°57.8ʹE, elev. 875m, camp #1, pitfall trap line #3, 26–31 October 2003, L. E. Olsen, [ FMNH # LEO128 ] (Holotype, CASENT9064782 , 1♂, FMNH), (paratype, CASENT9062274 , 1♂, FMNH), (paratype, CASENT9062280 , 1♂, FMNH), (paratype, CASENT9062275 , 1♂, CAS) .

Natural history. The type locality is in a region of rainforest. Collection data reveal that males of the cribellate Uduba lakroa wander, as they were collected in pitfall traps.

Distribution. Uduba lakroa are known only from the type locality in far southeastern Madagascar (Maps 12, 21).

Figures

FIGURE 48. Uduba lakroa, new species, paratype (CASENT9062275), from Mt. Papango in Parc National de Midousy, left male palpus. A. Prolateral. B. Ventral. C. Retrolateral. White arrow (in figs. A, B) points to bump on tegulum. Illustrations by Daniele Polotow. TA1, proapical apophysis of the tegulum; TA2, proapical apophysis of the tegulum; TA3, retroapical apophysis of the tegulum. Scale bar = 1.0 mm.

MAP 21. Distribution of Unclassified species: Uduba sarotra, U. ida, U. lakroa and Uduba madagascariensis (Vinson, 1863).