Allocladius fortispinatus (Edwards)

Mauad, Melina, Siri, Augusto & Donato, Mariano, 2013, New species of Pseudosmittia Edwards, 1932 and new records of Allocladius Kieffer, 1913 (Diptera: Chironomidae, Orthocladiinae) from South America, Zootaxa 3694 (5), pp. 445-460 : 454-455

Allocladius fortispinatus (Edwards)

Spaniotoma (Smittia) fortispinata Edwards, 1931: 296 .

Bryophaenocladius fortispinata (Edwards) ; Ferrington & Saether (1995: 57). Bryophaenocladius fortispinatus (Edwards) ; Spies and Reiss (1996: 75).

Allocladius fortispinatus (Edwards) ; Andersen et al. (2010: 10); Ferrington & Saether (2011: 50).

Material examined. ARGENTINA: Holotype male, Río Negro, Bariloche, 23–28.x.1926, F.W. Edwards (BM 1927–63, BMNH); 2 males, Río Negro, P.N. Nahuel Huapi, Mallín La Heladera, 41º 00' 06.4'' S, 71º 48' 40.3'' W, 878 m a.s.l., 7–i / 4.ii.2007, Malaise trap, A. Garre and F. Montes de Oca (MLP); 5 males, same data except 15.xii.2006, sweep net.

Diagnosis. Emended from Andersen et al. (2010) and Ferrington & Saether (2011). The combination of supraalar absent; long costal extension; parallel-sided anal point; superior volsella slightly developed as basal swelling fused with long, narrow and adpressed inferior volsella without accessory lobe; and pars ventralis absent distinguishes A. fortispinatus from the rest of the species of the genus.

Male (n = 8, except when otherwise stated). Total length 1.54–2.17 [2.17] mm. Wing length 1.05–1.33 [1.33] mm. Total length/wing length 1.44–1.71 [1.63]. Wing length/length of profemur 2.44–3.00 [3.00]. Coloration uniformly dark brown.

Head. Antennae with 13 flagellomeres, AR 0.43–0.48 [0.48] (3), ultimate flagellomere 180–230 [230] (3) long. Temporal setae 0–2 [not observed] divided in 0–1 [not observed] (6) inner verticals, 1–2 [not observed] outer verticals and 0–2 [2] (6) postorbitals. Clypeus with 9–10 [8] setae. Tentorium 98–120 [120] long and 15–22 [not observed] wide. Stipes 81–100 [100] (6) long and 5–7 [not observed] (6) wide. Palpomere lengths (1–5): 15–22 [20]; 22–34 [34]; 46–64; 51–73; 66–78 (6).

Thorax. Antepronotum with 1 lateral setae [not apparent]. Dorsocentrals 6–10 [8]; acrostichals 2–4 [2]; prealars 3 [2]; supraalars absent. Scutellum with 1–2 [2].

Wing. VR 1.4 [1.4]. C extension 88–110 [90]. Brachiolum without setae (5); all veins bare. R4+5 ending distal to M3+4.

Legs. Spur of front tibia 35–42 [not observed] (6) long; spurs of mid tibia: 12–14 [14] (6) and 15–17 [18] (6) long; of hind tibia: 12–15 and 34–42 (6) long. Width at apex of front tibia 24–32 [not observed]; of midd tibia 24– 34 [not observed]; of hind tibia 29–42 [not observed]. Comb with 9–11 [not observed] setae. Lengths and proportions of legs in Table 4 .

Hypopygium. Anal point parallel-sided with rounded apex,17–28 [19] long, 4–8 [5] wide. Tergum IX with 4– 12 [12] setae, laterosternite IX with 3–5 [4–6] setae. Phallapodeme 30–58 [58] long; transverse sternapodeme 46– 60 [60] long, with well developed oral projections. Without virga. Gonocoxite 134–154 [146] long; length from base of gonocoxite to apex of inferior volsella/length of gonocoxite 0.15–0.20 [not observed]. Gonostylus 46–54 [54] long; megaseta 6–12 [12] long. HR 2.84–3.14 [2.70]; HV 3.16–4.21 [4.02].

Tables

TABLE 4. Lengths (in µm) and proportions of legs of Allocladius fortispinatus (male) (n = 8, except when otherwise stated). Measurements of the holotype in square brackets. For abbreviation explanations see Table 1.

fe	ti	ta1	ta2	ta3
P1 400–490 [443]	420–540 [451]	260–320 (4)	180–220 (4)	120–150 (4)
P2 400–500 [451] (6)	390–470 [451]	170–210 [197] (6)	110–120 [123] (6)	70–90 [90] (6)
P3 430–520 [467]	440–560 [517]	230–300 [287] (5)	120–140 [164] (5)	110–150 [139] (4)
ta4	ta5	LR	BV	SV
P1 60–80 (4)	50–60 (4)	0.45–0.64 (4)	2.49–3.68 (4)	2.97–4.42 (4)
P2 40–50 [49] (6)	40–50 (6)	0.43–0.54 [0.44] (6)	3.17–3.85 (4)	3.44–4.71 (5)
P3 50–60 [49] (4)	50–60 (4)	0.46–0.54 [0.56] (3)	3.05–3.54 (3)	4.17–3.75 (3)