Scaphisoma furcigerum, Ivan Löbl & Ryo Ogawa, 2016

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1423-1424

5.2.19.9. Scaphisoma furcigerum nov.sp. (Figs 13, 204-206)

T y p e m a t e r i a l: Holotype 6: Burungkot, Upi, Cotabato Prov. Mindanao Elev,1500 ft., I,1-9, 1947 / Lot 154 taken on polypores / CNHM-Philippine Zool. Exped. (1946-47) F.G.Werner leg. (FMNH). Paratypes: Mindanao, with the same data as the holotype, 5 66 (FMNH, MHNG); Mindanao, South Cotabato, Motoklot, 650 m, Maitum, 11.VIII.1985, leg. M. Sakai, 2 66 (EUMJ, MHNG).

D e s c r i p t i o n: Length 1.10-1.23 mm, width 0.75-0.83 mm. Head and most of body dark reddish-brown to blackish, apical abdominal segments yellowish, femora ochreous, tibiae, tarsi and antennae yellowish. Pronotum and elytra not microsculptured and not iridescent. Length/width ratio of antennomeres as: III 11/6: IV 26/5: V 48/6: VI 40/6: VII 50/9: VIII 43/7: IX 50/10: X 45/10: XI 55/11. Pronotal punctation fairly finely and densely near base, very finely and sparsely on centre, densely and comparatively coarsely on lateral areas; pronotum with lateral contours weakly rounded, lateral margin carinae hardly visible in dorsal view, lateral striae coarsely punctate. Exposed tip of scutellum minute. Elytra with lateral margin carinae entirely visible in dorsal view, apical margins truncate, inner apical angles not prominent, situated somewhat posterior level of outer angles; sutural margin not raised, sutural striae deep, starting posterior level of scutellum, parallel toward apical third of sutural length, converging toward apices, coarsely punctate; adsutural areas narrow, flat, densely and finely punctate, lateral and epipleural striae distinctly punctate. Elytral disc with punctation dense and coarse from bases to apical fourth, puncture rows fairly distinct, puncture intervals mostly about as large to twice as large as puncture diameters; apical fourth fairly finely and sparsely punctate. Hind wings fully developed. Hypomera and mesanepisterna smooth. Mesepimera not quite twice as long as intervals to mesocoxae, about four times as long as wide. Median part of metaventrite convex, with shallow apicomesal impressions, lacking distinct microsculpture; with punctation fairly coarse and dense in apicomedian impressions, very fine and dense in middle, sparse and very fine on sides; antecoxal puncture rows present, in impressed lines. Submesocoxal areas 0.02 mm, about as fifth of intervals to metacoxae, submesocoxal lines parallel, coarsely punctate. Metanepisternum convex, hardly narrowed anteriad, inner margin rounded, impressed below margin of metaventrite. Tibiae straight. Abdomen with strigulate microsculpture, very finely punctate; sternite 1 with submetacoxal areas 0.04 mm, about as third of intervals to apical margin of sternite, submetacoxal lines convex, coarsely punctate.

M a l e: Protarsomeres 1 to 3 weakly widened. Aedeagus ( Figs 204-206 ) 0.66-0.75 mm long, symmetrical, strongly sclerotized. Apical process of median lobe robust, inflexed, with almost oblique ventral side, blunt apex. Articular process robust, not prominent. Parameres with strongly widened bases and each with large, curved lobe, in dorsal view evenly wide and almost evenly curved, excepted in basal section. Internal sac with a pair of conspicuous, strongly sclerotized rods joined by V-shaped apophysis and complex basal sclerites, membranous denticulate structures present in basal part only.

E t y m o l o g y: The species epithet is a Latin adjective meaning bearing a fork and refers to the strongly sclerotized aedeagal rods.

C o m p a r a t i v e n o t e s: This species is similar to S. furcatum in external characters, thought it may be readily distinguished by the punctation on lateral parts of the pronotum. It differs drastically from the latter by its aedeagal characters. The genital characters suggest close relationships with S. furcillatum described below. These two species share robust, short apical processes of the median lobe, parameres bearing large ventral lobes and internal sac possessing long, strongly sclerotized and mesally joined apical rods.

Figures

Figs 201 - 206: Aedeagi of Scaphisoma; (201, 202), S furcatum nov. sp., in dorsal and lateral views, scale = 0.2 mm; (203), ditto, internal sac, scale = 0.1 mm; (204, 205), S. furcigerum nov. sp., in dorsal and lateral views, scale = 0.2 mm; (206), ditto, internal sac, scale = 0.2 mm.