Scaphinotus (Pseudonomaretus) merkelii (G. H. Horn) 1890

Kavanaugh, David H. & Angel, Kathleen W., 2015, A Taxonomic Review of the Northwestern Nearctic Subgenus Pseudonomaretus Roeschke 1907, including Description of the New Species Scaphinotus hoodooensis (Coleoptera: Carabidae: Cychrini) from the Bitterroot Mountains of Montana, Proceedings of the California Academy of Sciences 62 (14), pp. 381-396 : 391-393

Scaphinotus (Pseudonomaretus) merkelii (G. H. Horn) 1890

Figures 2b, 3b , 5 , 9

Cychrus merkelii G. H. Horn 1890:71 . Holotype, a female, in MCZ, labeled: “Ida”/ “Merkel”/ “ C. merkelii Horn ”/ “ Cychrus merkelii Horn ” [red label]/ “MCZ Holotype 34934” [red label]. Type locality: Coeur d’Alene, Idaho (designated by Lindroth, 1961:20).

Cychrus idahoensis Webb 1901:133 . Lectotype, here designated, a male, in WSU, labeled: “Moscow Mt. Idaho ”/ “July 99”/ “Type I” [red label]/ “ LECTOTYPE Cychrus idahoensis Webb , design. by D. H. Kavanaugh and K. W. Angel 2015”. We examined one male and two female paralectotypes with same data as lectototype, except each labeled with a different “Type” number (“II”, “3” and “IV”, respectively), and one additional female paralectotype (labeled: “Collins, Idaho July 24, 1898.”/ “Type 5”, all also in WSU. Each of these paralectotypes now also bears the following label: “ PARALECTOTYPE Cychrus idahoensis Webb 1901 , design.by D. H. Kavanaugh and K. W. Angel 2015” [yellow label]. Synonymy established by Gidaspow (1973). Type locality: Moscow Mountain, Latah County, Idaho.

Pseudonmomaretus merkeli (G. H. Horn) . Casey, 1914:30.

Scaphinotus (Pseudonomaretus) merkeli (G. H. Horn) . Roeschke 1907: ##; Hatch 1953:45; Lindroth, 1961:20; Gidaspow 1973:74.

Scaphinotus (Pseudonomaretus) merkelii (G. Horn) . Bouquet 1993; Bergdahl 2014:4.

NOTES ON NOMENCLATURE AND TYPES.— In his original description of Cychrus idahoensis, Webb (1901) noted that he had seen a total of 14 specimens, including 11 from “Cedar Mountain” [now know as Moscow Mountain] and three from Collins, Idaho. This suggests that there are nine additional specimens, which we have not seen, that should also be considered paralectotypes of this name .

RECOGNITION.— Adults of this species can be distinguished from those of other species of Pseudonomaretus by the following combination of character states: SBL = 12.5 mm or less (males 9.9–1.0 mm, females 10.7–12. 2 mm); elytra moderately shiny, with microsculpture shallowly impressed, without or with faintly suggested bronze metallic reflection; head with a distinct transverse impression at level of posterior margin of eyes clearly delineating frontal and occipital regions; labrum ( Fig. 2b) with lateral lobes markedly long and narrow; antennomere 4 ( Fig. 3b ) as pubescent as antennomere 5 or nearly so, apical half of antennomere 3 also but more sparsely pubescent; pronotum with basolateral pair of setiferous pores present; male protarsomere 1 ( Fig. 5b ) with pad of adhesive setae ventrally on apical two-thirds only, protarsomere 4 also with a pad; elytra with 13 to 15 striae, with none to all of them interrupted by punctures in and between striae, elytral intervals 4 and 8 without discal setiferous pores; median lobe of male aedeagus ( Fig. 9) slen- der, with ventral margin evenly arcuate in lateral aspect ( Fig. 9b), widest and slightly flaired laterally just distal to midpoint of shaft and gradually tapered to a broadly rounded apex in dorsal aspect ( Fig. 9a).

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GEOGRAPHICAL DISTRIBUTION.— This species is known from southeastern British Columbia, northwestern Montana and northern and central Idaho ( Gidaspow 1973; Bergdahl 2014).

Figures

FIGURE 3. Digital photographs of right antenna, antennomeres 1 to 5, of Scaphinotus (Pseudonomaretus) species, dorsal aspect, with antennomeres identified by number; a, S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana); b, S. merkelii (G. Horn) (St. Joe National Forest, Idaho); c, S. regularis (LeConte) (Hayden Creek, Kootenai County, Washington); d, S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho); e, S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 1.0 mm.

FIGURES 4–7. Digital photographs of male left protarsus of Scaphinotus (Pseudonomaretus) species, with tarsomeres identified by number; a, dorsal aspect; b, ventral aspect; as = pad of adhesive setae. Fig. 4. S. hoodooensis Kavanaugh and Angel sp. nov. (near Hoodoo Pass, Mineral County, Montana). Fig. 5. S. merkelii (G. Horn) (9 km SE of Lowell, Idaho County, Idaho). Fig. 6. S. relictus (G. Horn) (Selway River at O’Hara Campground, Idaho County, Idaho). Fig. 7. S. mannii Wickham (Steptoe Canyon, Whitman County, Washington). Scale lines = 1.0 mm.