Pupina paviei Morlet, 1883

Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak, 2022, Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia, ZooKeys 1119, pp. 1-115 : 1

Pupina paviei Morlet, 1883

Figs 37F-I , 38F, G , 39A, B

Pupina paviei Morlet, 1883: 107, 108, pl. 4, fig. 4. Type locality: La chaîne de l’Éléphant et les forêts non inondées qui la bordent, particulièrement, près des rapides de Kamchay et aux environs de Kampot [The Elephant Range and the non-flooded forests that border it, particularly near the Kamchay rapids and around Kampot; currently Preah Monivong Bokor National Park, Kampot Province, Cambodia]. Morlet 1889: 152. Fischer 1891: 107. Fischer and Dautzenberg 1904: 431. Morlet 1904: 370, 371, pl. 20, fig. 13, 13a. Fischer-Piette 1950: 153. Fischer 1973: 48. BEDO 2017: 92.

Pupina (Tylotoechus) paviei - Kobelt 1902: 319.

Type material examined.

Paralectotypes MNHN-IM-2000-35837 (4 shells; Figs 37F , 38F ) from Chaîne de l’Eléphant, Kampot, Cambodge. Paralectotypes RBINS 525404 (76 shells; Figs 37G , 38G ) from Kampot et forêts de la chaîne de l’Eléphant, Cambodge et Kamchay.

Material examined.

NHMUK ex. Dautzenberg coll. (1 shell; Figs 37I , 39A ) from Kampot, Cambodge. CUMZ 12134 (129 shells; Figs 37H , 39B ) from Lalu, Ta Phraya District , Sa Kaeo Province, 24 Nov. 2006 .

Diagnosis.

Shell globose to ovate; last whorl ca. three quarters of shell height. Apertural lip slightly thickened but not expanded. Parietal tooth triangular, not thickened to slightly thickened, covering posterior canal but not extending beyond apertural lip; columellar tooth fin-shaped, slightly thickened, located next to slit-like anterior canal.

Differential diagnosis.

Pupina paviei is similar to P. tongupensis in a globose shell shape, but differs in having a triangular parietal tooth that is either not thickened or slightly thickened, and a fin-shaped, slightly thickened columellar tooth that is located next to a slit-like anterior canal.

Distribution.

Cambodia ( Morlet 1883) and Sa Kaeo Province, eastern Thailand.

Remarks.

As the original description did not explicitly state that the description of this species was based on a single specimen (nor could this be inferred), the designation of a holotype by Fischer-Piette (1950) in fact constitutes a lectotype designation ( ICZN 1999: Art. 74.6).

Figures

Figure 37. Shells of Pupina aureola species group from mainland Southeast Asia. A-C Pupina tchehelensis: specimens A SMF 109947 / 6 B CUMZ 12135, and C CUMZ 12136 D, E Pupina latisulci sp. nov. D holotype CUMZ 12146 / 1 and E paratype CUMZ 12146 / 2 F-I Pupina paviei F paralectotype MNHN-IM- 2000 - 35837 G paralectotype RBINS 525404, and specimens H CUMZ 12134 and I NHMUK ex. Dautzenberg coll. J Pupina tongupensis, syntype NHMUK 1906.4.4.38 K Pupina lowi, syntype MNHN-IM- 2000 - 35846 L Pupina anceyi, syntype MNHN-IM- 2000 - 35833 M Pupina laffonti, syntype MNHN-IM- 2000 - 9656 N Pupina solidula, lectotype SMF 109915 / 1 O Pupina douvillei, holotype MNHN-IM- 2000 - 35532. Photo: M. Caballer, P. Maestrati, MNHN (F, K-O), F. Trus, RBINS (G), J. Ablett, H. Taylor, NHM (J).

Figure 38. A-E Pupina aureola: specimens A CUMZ 12121 from Tham Thong Panara Temple, Nakhon Sri Thammarat B CUMZ 12124 from Talot Cave, Nakhon Sri Thammarat C CUMZ 12126 from Khao Huai Haeng Temple, Trang D CUMZ 12130 from Sra Morakot, Krabi, and E CUMZ 12133 from Khantiphol Cave, Satun F, G Pupina paviei: paralectotypes F MNHN-IM- 2000 - 35837 from Chaine de l'Elephant, Kampot, Cambodge and G RBINS 525404 from Kampot et forets de la chaine de l'Elephant, Cambodge et Kamchay. Photo: P. Maestrati, MNHN (F), F. Trus, RBINS (G).

Figure 39. A, B Pupina paviei: specimens A NHMUK ex. Dautzenberg coll. from Kampot, Cambodge and B CUMZ 12134 from Lalu, Sa Kaeo C, D Pupina tchehelensis: specimens C SMF 109947 / 6 from Bukit Pondong, Perak and D CUMZ 12136 from limestone mountain, Phang Nga E, F Pupina dorri isanensis ssp. nov. E holotype CUMZ 12140 / 1 and F specimen CUMZ 12137 from Khao Wang Pha, Nong Bua Lam Phu.