Proeryon charbonnieri, Audo & Schweigert, 2018

Audo, Denis & Schweigert, Günter, 2018, Large polychelidan lobsters with a rounded carapace from the Middle Jurassic La Voulte-sur-Rhône Lagerstätte: taxonomic clarifications, Geodiversitas 40 (9), pp. 183-194 : 188

Proeryon charbonnieri n. sp.

( Figs 3-5 )

urn:lsid:zoobank.org:act:BE1BD92F-4B39-4B6A-B222-AD80ECB698E1

Eryon giganteus Van Straelen, 1923: 91 , fig. 8 (pro parte). — Balss 1924: 175.

Coleia gigantea – Van Straelen 1925: 145-147, fig. 66, pl. 3, figs 4 (non fig. 3), pl. 4, figs 1-2. — Roman 1928: 109, fig. 18, pl. 3, fig. 4 (non fig. 1). — Glaessner 1929: 127. — Charbonnier 2009: 158-159, 163, figs 236, 244, 224-229, 393, 394, 399-401. — Charbonnier et al. 2010: 115-117, figs 3c, 4c, 11a-c (pro parte).

Cyclocaris giganteus – Pinna 1968: 103.

Cycleryon gigantea (sic) – Fischer 2003: 241, fig. 27 (genus also misspelled Glycerion and Cyclerion).

Proeryon giganteus – Schweitzer et al. 2010: 44 (pro parte). — Audo et al. 2014c: 505-509, figs 2E, 6, 7.

“ Coleia ” gigantea – Charbonnier et al. 2014: 375, table 1, fig. 4F.

ETYMOLOGY. — The specific epithet honors the French palaeontologist Sylvain Charbonnier, who recently rediscovered the lectotype of E. giganteus and published the first comprehensive study of the La Voulte-sur-Rhône Lagerstätte.

TYPE MATERIAL. — Holotype: UJF-ID 14023 ( Fig. 3 A-C).

Paratypes: MSNM.i20703 ( Fig. 3D ); UJF-ID.11547, 11548-11549 (part and counterpart, Fig. 4B ), 11550, 11552, 14020, 14051, 14052 ( Fig. 4C ); FSL 170603, 170607-170609 (coll. Marin), 710080; MNHN.F.A50714, A50716-A50719, A50726, A50741 (coll. Fischer, Fig. 4A ), B11759 ( Fig. 4D ) ; MNHGr.PA.10203 ( Fig. 4E ), 10276; UPMC-248.

ADDITIONAL SPECIMENS. — FSL 170521, 170610 (coll. Caillet), 71081 (coll. Charbonnier),MNHN.F.A50713, A50727 (coll. Fischer), R03515 (cast) ; UJF-ID.14026, 14047-14049 (part and counterpart), 15050.

TYPE LOCALITY. — Ravin des Mines, near La Boissine, La Voultesur-Rhône Lagerstätte, Ardèche, Auvergne-Rhône-Alpes, France.

TYPE AGE. — Early Callovian, Gracilis ammonite Zone.

DESCRIPTION

Carapace (cephalothoracic shield) outlines ( Fig. 5 ) Dorsoventrally flattened carapace, pyriform in outline in dorsal view; slightly concave frontal margin; anterolateral angle forming a small spine oriented diagonally; spiny lateral margin, cut by ocular, cervical and hepatic incisions; hemicircular ocular incision, opening laterally; spiny and convex anterolateral margin, oblique compared to longitudinal axis; anterolateral spine as large as the other anterolateral spines; cervical and hepatic incisions opening in the anterior half of the lateral margin; marked, shallow cervical incision; straight mediolateral margin, slightly longer than anterolateral margin and fringed by a few small spines; subtriangular hepatic incision, shallower than cervical incision; curved posterolateral margin fringed with small spines; rounded posterolateral angle contiguous with pleon; slightly concave posterior margin wider than anterior margin.

Figures

FIG. 3. — Proeryon charbonnieri n. sp. from the La Voulte Lagerstätte: A-C, holotype UJF-ID 14023 (former paralectotype of S. giganteus n. comb.); A, dorsal view, cross-polarized light; B, third maxillipeds and pereiopods, 3D model from CT-scan data, modified from Audo et al. (2014c); C, interpretative line-drawing; D, paratype MSNM i20703, dorsal view, cross-polarized light. Abbreviations: a, branchiocardiac groove; a1, antennula; a2, antenna; b1, hepatic groove; bc, branchial carina; bi, hepatic incision; c, postcervical groove; d, gastro-orbital groove; e1e, cervical groove; ei, cervical incision; lc, lateral carina; mdb, mandible; mxp3, third maxilliped; o, ocular incision; P1-P5, pereiopods 1 to 5; pc, postcervical carina; po, postorbital carina; s1-s5, pleonites 1 to 5. Scale bars: 50 mm. Images: D. Audo.

FIG. 4. — Proeryon charbonnieri n. sp. from the La Voulte Lagerstätte:A, paratype MNHN.F.A50741, dorsal surface, natural light (A1) and tail-fan (A2); B, paratype UJF-ID.11548-49 (former paralectotype of S. giganteus n. comb.), partial tail-fan,counterpart,cross-polarized light; C, paratype MNHN.F.B11759, dorsal surface, natural light; D, paratype UJF-ID.14052 (former paralectotype of S. giganteus n. comb.), dorsal surface, cross-polarized light, small-sized specimen; E, paratype MHNGr.Pa.10203, dorsal surface, natural light. Abbreviations: a, branchiocardiac groove; b1, hepatic groove; ba, uropodal basipod; di, uropodal diaeresis; e1e, cervical groove; ei, cervical incision; ex, uropodal exopod; lc, lateral carina; o, eye; oph, ophiuroid; P1, pereiopod 1; s5-s6, pleonites 5 to 6; t, telson. Scale bars: 20 mm. Images: A1, A2, Peter Massicard (E-Recolnat project); B-D, D. Audo; E, Noël Podevigne.

FIG. 5. — Sketch ofcarapace outline,grooves and carinae ofProeryon charbonnieri n. sp. Abbreviations: a, branchiocardiac groove; ala, anterolateral angle; b1, hepatic groove; bc, branchial carina; bi, hepatic incision; c, postcervical groove; d, gastro-orbital groove; e1e, cervical groove; ei, cervical incision; lc, lateral carina; o, ocular incision; pc, postcervical carina; pla, posterolateral angle; po, postorbital carina; pr, postrostral carina. Not to scale.