Phortica (Ashima) halimunensis Toda, 2020

Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru, 2020, Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species, Zootaxa 4789 (1), pp. 1-54 : 24

21) Phortica (Ashima) halimunensis Toda , sp. nov.

( Fig. 6 )

Diagnosis. Apical piece of aedeagal, median rod small and flat plate ( Fig. 6E,F ).

Description (not repeating characters common to P. tanabei ). Supracervical setae 8–11. Dorsomedial, tentorial apodeme slightly shorter than basal, parallel portion of dorsolateral, tentorial apodeme. Arista with arb = 1–2/1; apical, leaf-like expansion apically somewhat roundish and with short, awn-like projection ( Fig. 6A ). Cibarial, medial sensilla 5–6 per side; posterior sensilla approximately 5 per side. Fore and hind tarsomere Is as long as 3 succeeding tarsomeres together; mid one slightly longer than rest tarsomeres together.

Measurements (holotype /range of 4♂ paratypes, in mm): BL = 2.90/2.46–2.74, ThL = 1.39/1.15–1.37, WL = 2.11/1.83–2.11, WW = 0.95/0.86–0.98.

Indices (holotype /range of 4♂ paratypes, in ratio): FW/HW = 0.46/0.45–0.47, ch/o = 0.07/0.07–0.08, prorb = 1.13/1.11–1.29, rcorb = 0.66/0.47–0.69, vb = 0.31/0.33–0.46, dcl = 0.44/0.37–0.47, presctl = 0.41/0.45–0.51, sctl = 1.00/1.00–1.07, sterno = 0.88/0.85–0.99, orbito = 1.84/2.00–2.39, dcp = 0.22/0.21–0.26, sctlp = 1.08/0.93–1.12, C = 2.31/2.06–2.20, 4c = 1.44/1.43–1.70, 4v = 2.87/2.62–3.21, 5x = 1.30/1.04–1.28, ac = 3.72/3.34–4.03, M = 0.70/0.59–0.80, C3F = 0.54/0.61–0.72.

Holotype. ♂ ( MZB), “ Cikaniki , Mt. Halimun, West Java, Indonesia, 6°44’42.5” S 106°32’14.5” E, 1,051 m a.s.l., 4.xi.2009, ex around eyes, M.J. Toda ”.

Paratypes. Indonesia: 1♂, same data as the holotype, except 5.xi.2009 ( MZB) ; 1♂, ditto, except ex tree trunk ( SEHU) ; 2♂, same data as the holotype, except 9.xi.2009 ( SEHU) .

Distribution. Java (West Java).

Remarks. This species is coupled with P. tanabei (BP = 72%; Fig. 1 ), sharing the aedeagal, median rod distally diffentiated into one asymmetric sclerite piece (ch.58-3; Fig. 6E,F ), but can be distinguished from it by the diagnostic character.

Etymology. Referring to the type locality.

Figures

FIGURE 1. The strict consensus tree resulting from the parsimony analysis I (PAUP* v4.0a166) of the data matrix of 40 spp. × 66 morphological characters (Appendix 1) for the genus Phortica (especially focusing on the subgenus Ashima). Synapomorphies (solid circle: nonhomoplastic; open circle: homoplastic) inferred from both ACCTRAN and DELTRAN character optimization are indicated on each internal branch along with support values (bootstrap frequency %).

FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C.