Landouria tonywhitteni, Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 45-47

publication ID

https://doi.org/ 10.5852/ejt.2019.526

publication LSID

lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC

DOI

https://doi.org/10.5281/zenodo.11237934

persistent identifier

https://treatment.plazi.org/id/59260748-3438-411F-82B4-14A7847AFA58

taxon LSID

lsid:zoobank.org:act:59260748-3438-411F-82B4-14A7847AFA58

treatment provided by

Plazi

scientific name

Landouria tonywhitteni
status

sp. nov.

Landouria tonywhitteni View in CoL sp. nov.

urn:lsid:zoobank.org:act:59260748-3438-411F-82B4-14A7847AFA58

Figs 39 View Figs 29–47 , 52 View Figs 52–55 , 60 View Figs 60–62 , 63 View Figs 63–71 ; Tables 3–4 View Table 3 View Table 4

Diagnosis

Landouria tonywhitteni sp. nov. is characterized by having a strongly depressed conical, brownishcorneous, sharply keeled shell and a penis that does not become distinctly broader towards is proximal end, the lack of a coecum at the proximal end of the penis and the presence of a node at the base of the flagellum.

Etymology

Landouria tonywhitteni sp. nov. is named in honour of the late British naturalist Tony Whitten (1953– 2017), who dedicated his life to the conservation of karst habitats and especially the snails that inhabit these habitats.

Material examined

Holotype

INDONESIA • holotype (det. anat.); Central Java, Grobogan, Taruman, near Sendang Sedayu ; 7°00′52″ S, 110°51′29″ E; 20 m a.s.l.; 16 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; measurements: D = 17.6 mm, H = 7.6 mm; MZB 19214 .

GoogleMaps

Paratypes

INDONESIA – Central Java • 9 spec.; same data as for holotype; MZB 19217 5 spec.; same data as for holotype; ZMH 133434 View Materials GoogleMaps 9 spec. (det. anat.); Grobogan , Taruman, near Sendang Sedayu ; 7°00′52″ S, 110°51′29″ E; 23 m a.s.l.; MZB 19215 GoogleMaps 1 spec.; same data as for preceding; MZB 21268 GoogleMaps 5 spec.; same data as for preceding; ZMH 133436 View Materials GoogleMaps .

GoogleMaps

Description

SHELL ( Fig. 60 View Figs 60–62 ; Tables 3–4 View Table 3 View Table 4 ). Depressed conical, with 5.75–6 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct spiral lines, umbilical side with indistinct incised spiral lines; with radial scaly processes; tuberculate around umbilicus; brownishcorneous; body whorl sharply keeled; aperture rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 32–41% of shell diameter, hardly obscured by columellar edge.

GENITALIA ( Figs 39 View Figs 29–47 , 63 View Figs 63–71 ; Table 4 View Table 4 ). Atrium short; penis long, consisting of a long, cylindrical distal part covered by an additional sheath, which is connected by muscular tissue to proximal part of epiphallus and a dilated proximal part containing a short penial papilla; without penis coecum; penial retractor runs from diaphragm to distal part of long, narrow epiphallus; flagellum short, broadest at its base, where it is equipped with a globular node, tapering towards its proximal end, not distinctly crenated; vas deferens narrow and long; vagina long; peduncle of bursa copulatrix subdivided into a slightly broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.

Remarks

Landouria nodifera sp. nov., L. tonywhitteni sp. nov., L. pacitanensis sp. nov., L. sewuensis sp. nov. and L. intumescens share strongly depressed, brownish-corneous, sharply keeled shells. These species cannot be reliably distinguished by shell characters. Landouria tonywhitteni sp. nov. shares a node at the base of the flagellum with L. nodifera sp. nov., from which it differs in the lack of a coecum at the proximal end of the penis and in having a penis that does not become broader towards its proximal end.

Distribution

Landouria tonywhitteni sp. nov. is endemic to Sukolilo Karst (Fig. 52), where it occurs sympatrically with L. sukoliloensis sp. nov.

MZB

Museum Zoologicum Bogoriense

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