Hellyethira malleoforma Wells

Wells, Alice & Johanson, Kjell Arne, 2015, Review of New Caledonian species of Oxyethira Eaton, with description of 17 new species, and new records for Hydroptila Dalman and Hellyethira Neboiss (Trichoptera, Hydroptilidae), ZooKeys 530, pp. 37-90 : 73-75

Taxon classification Animalia Trichoptera Hydroptilidae

Hellyethira malleoforma Wells Fig. 113

Hellyethira malleoforma Wells, 1979: figs 41-45; Wells 1995: 232.

Diagnosis.

Males of this species are distinguished by their complex asymmetrical genital structures, including multilobed gonopods, and females by the sclerotised annulus formed by abdominal segment VIII (see Wells 1979).

Material examined.

2 males, females, Province Sud, Sarraméa, 220 m, forest stream, loc 10, 21°37.883'S 165°51.958'E, Malaise trap, 18-21.xi.2001, leg. K.A. Johanson, T. Pape & B. Viklund(NHRS); 1 male, 4 females, Province Sud, Sarraméa, 2907 m, stony forest stream, loc 13, 21°37.097'S 165°49.351'E, Malaise trap, 18-21.xi.2001, leg. K.A. Johanson, T. Pape & B. Viklund(NHRS); numerous males, females, Province Nord, Amoa River, 23 m, loc 20, 12 km W Poindimié, 22°58.092'S, 165°11.804'E, light trap, 26.xi.2001, leg. K.A. Johanson, T. Pape & B. Viklund(NHRS); 1 male, 4 females, Province Sud, Monts Kwa Ne Mwa, on road between Noumea and Yaté, 2.0 km E Pic Mouirange, 22°12.356'S, 166°40.798'E, 220 m, 7-16.xi.2003, Malaise trap, loc#014, leg. K.A. Johanson (NHRS); 2 males, 17 females, Province Sud, Mt Dzumac, source stream of Ouinne River, near crosspoint to mountain track, 22°02.439'S, 166°28.646'E, 805 m, 18. xi– 4.xii.2003, Malaise trap, loc#029, leg. K.A. Johanson (NHRS); males, females, Province Sud, Tamoa River, 700m S road RT1 between Noumea and La Foa, 22°04.518'S, 166°16.592'E, 19.xi.2003, light trap, loc#033, leg. K.A. Johanson (NHRS); males, females, Province Sud, stream at Refuge de Farino, 4.0 km W Grand Couli village, 21°38.934'S, 165°46.845'E, 260 m, 25.xi.2003, light trap, loc#044, leg. K.A. Johanson (NHRS); 4 males, 2 females, St. Vincent, Bongou Stream, at bridge on road to Tribu de Bangou, 700 m N RT1 Noumea–Tontoutu road, 22°03.477'S, 166°15.718'E, 26.xi.2003, light trap, loc#050, leg. K.A. Johanson (NHRS); numerous males, females, Province Nord, 50 m upstream bridge on Hienghène–Tnèdo road, 3.9 km S summit of Mt Tnèda, 2.2 km E Tnèdo, 20°43.085'S, 164°49.928'E, 29 m, 7.xii.2003, light trap, loc#071, leg. K.A. Johanson (NHRS); males, females, Province Nord, 1 m upstream road, below waterfall on Hienghène–Tnèdo road, 2.2 km SSW summit of Mt Unpac,4.9 km ESE Tnèdo, 20.73879°S, 164.85508°E, 7.xii.2003, light trap, loc#072, leg. K.A. Johanson (NHRS); males, females, Province Nord, Wan Pwé on stream, draining NNE side of Mt Panié, 3.9 km NW Cascade de Tao, 20°31.820'S, 164°47.016'E, 18.xii.2003, light trap, loc#085, leg. K.A. Johanson (NHRS); numerous males, females, Province Nord, Bouérabate Stream, S Mont Ninndo, along road Barabache–Boulagoma, 20°17.409'S, 164°11.242'E, 60 m, 19.xii.2003-7.i.2004, Malaise trap, loc#089, leg. K.A. Johanson (RHNS); males, females, Province Nord, Rivière Néhoué, camp Amenage de Néhoué, 20°25.037'S, 164° 13.222'E, 12 m, 19.xii.2003, light trap, loc#090, leg. K.A. Johanson (NHRS); males, females, Province Nord, Rivière Néhoué, camp Amenage de Néhoué, 20°25.015'S, 164° 13.245'E, 12 m, 19.xii.2003, light trap, loc#091, leg. K.A. Johanson (NHRS); male, females, Province Nord, Héémwâ Pwei River, 50 m upstream bridge on Touho– Hienghene road, 1.0 km N Paola, 20.76512°S, 165.10979°E, 22.xii.2003, light trap, loc#095, leg. K.A. Johanson (NHRS); numerous males, females, Province Nord, Ponandou Tiôgé River at Kögi, 3.9 km SSW Touho, 20°49.043'S, 165°13.551'E, 25 m, 26.xii.2003, light trap, loc#100, leg. K.A. Johanson (NHRS); males, females, Province Nord, Plaine des Gaïacs, Rivière Rouge, 14.2 km NW summit of Mt Rouge, 50 m upstream road RT1 Noumea–Koné, 21°31.573'S, 164°46.690'E, 23 m, 2.i.2004, light trap, loc#104, leg. K.A. Johanson (NHRS); 1 male, KAJ sp. F, Province Sud, Creek Froid, 10 m upstream bridge on La Foa–Koindé road, 200 m W crossroad to Ouipouin, 21°38.581'S, 165°56.672'E, 180 m, 4.i.2004, light trap, loc#105, leg. K.A. Johanson (NHRS); 1 male KAJ sp. F, Province Sud, Fö Néchédeva stream, 2 m upstream bridge on La Foa–Koindé road, 21°38.812'S, 165°56.076'E, 124 m, 4.i.2004, light trap, loc#106, leg. K.A. Johanson (NHRS); numerous males, females, Province Nord, Etnbl. thermal de la Crouen, along Riv. la Crouen, 30 m upstream road RM3, 21°32.105'S, 165°53.319'E, 15 m, 5.i.2004, Malaise trap, loc#110, leg. K.A. Johanson 1 male KAJ “F”, Province Nord, Forêt Plate, Ouendé River, at 2.5 km WNW summit of Katépouenda, 23.3 km E Pouembout, 21°07.474'S, 165°06.781'E, 470 m, 8-15.i.2004, Malaise trap, loc#113, leg. K.A. Johanson (NHRS); numerous males, females, Province Nord, 2.8 km ENE Bopope, Rivière Oua Mendiou, 100 m S RPN2 Koné–Poindimié, 20°54.455'S, 165°06.300'E, 78 m, 14.i.2003, light trap, loc#119, leg. K.A. Johanson (NHRS).

Remarks.

Hellyethira malleoforma is the only representative in New Caledonia of this diverse Australian genus that also occurs more broadly but less commonly in SE Asia and New Guinea. This species is widespread and often abundant in New Caledonia (Fig. 113). It was described from south-eastern Australia where it is one of the most common species in lower altitude streams.