Glaresis australis Gordon and Hanley, 2014

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 44-45

Glaresis australis Gordon and Hanley , new species

Description. Male. Length 3.4 mm, width 1.8 mm; body form slightly elongate, feebly widened from elytral base to apical 1/3 ( Fig. 29A ). Color dark yellowish brown. Head with clypeal surface, frons and vertex with dense, pronounced, reticulate sculpture, clypeus and anterior 1/2 of frons with large, dense tubercles separated by less than diameter of a tubercle, setae short, indistinct. Clypeal apex emarginate, with coarse, evenly spaced tubercles, appearing somewhat serrate, lateral angles oblique, angulate ( Fig. 29B ). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin slightly abruptly rounded. Pronotum with deep, long, transverse fovea in anterior 1/4 extended completely across pronotum, small, slightly transverse fovea on each side of middle in anterior 1/2, wide, deep fovea on each side medially near lateral margin, deep central furrow extended from base to transverse furrow; surface densely rugose, with dense, extremely irregular, setae-bearing carinae, carinae often coalesced forming a somewhat reticulate pattern, setae decumbent, about 1/2 length of carina; anterior and lateral pronotal margins not bordered, anterior and basal margins crenulate. Elytra with surface slightly shiny, densely microreticulate; all striae strongly carinate, carinae appearing continuous but narrowly interrupted adjacent to space between interval punctures, each carinal segment bearing an apical seta about 1/2 length of segment; intervals with deep, round punctures ( Fig. 29C ). Metasternum long, dull, finely, densely microreticulate, median surface smooth medially, laterally with short, irregular ridges often coalesced, forming slightly reticulate pattern, median area flat, with faint median carina extended anteriorly from apical keel 1/2 distance to base; metasternal groove deep, inner margin abruptly tapered, outer margin abruptly ridged ( Fig. 29D ). Lateral protibial teeth not evenly spaced, 2nd and 3rd teeth close together. Mesotibia with 4 short, widely spaced spines laterally, tibia slightly projecting at apex. Posterior metatrochanteral margin serrate with several teeth; posterosuperior surface of metatrochanter with single small tooth, tooth not visible directly in ventral view. Metafemoral surface with widely spaced, elongate, obliquely transverse setae-bearing tubercles, surface microreticulate; width to length ratio 1.0:1.5, with wide flange on anterior margin; posterosuperior margin without teeth ( Fig. 29G ). Metatibia broadly triangular, surface finely, irregularly sculptured, with large, apically blunt, posteromedian lateral projection, small tubercles on outer margin from near base to lateral projection, row of coarse, pronounced, obliquely angulate row of tubercles extended from base nearly to apex medially, inner margin smooth, pubescent ( Fig. 29F ). Apex of 5th abdominal ventrite broadly, weakly rounded. Genitalia long, basal piece shorter than parameres, proximal end curved; median lobe shorter than parameres, slightly wider than paramere, curved upward in apical 1/3, apex rounded in ventral view; parameres weakly curved in lateral view, slightly curved on inner margin, apex bluntly rounded ( Fig. 29E ).

Female. Apex of 5th abdominal ventrite strongly, broadly rounded.

Variation. Length 2.7 to 3.5 mm, width 1.5 to 2.0 mm. Head with tubercles confined to clypeus and anterior 1/2 of frons, or frons entirely tuberculate, size and spacing of tubercles varies slightly; lateral mesotibial margin may have either 3 or 4 teeth; posterosuperior margin of metafemur occasionally has a single, very small tooth; posteromedian metatibial projection may be bluntly truncate apically, or apically bifid.

Type material. Holotype male; Baja California Sur: MEX: Baja Cal. Sur, 9.4 mi. W. hwy 1 on Ramal a San Felipe, IX-10-11-88: E. Riley, blacklight ( TAMU) . Paratypes, 57: (3) same data as holotype; (1) MEXICO: Baja CA Sur, Naranjas Road , 7 km W. Hwy 1, 100 m. el., 5-X-1991, BL + MV, RA Cunningham coll.; (1) MEX. Baja CA. Sur, 22 km S Jct. Hwy 19 + dirt Rd. 1.5, km S. Todos Santos , Vic. Rancho El Cachania, 380 m. El.; (6) Mex. Baja Cal. Sur, Las Barracas , 14-X 1985, Coll. P. DeBach, blk. light (1) MEXICO, Baja Calif. Sur, 14.8 km SE Todos Santos, 300 m. el., 30-IX-1990, RA Cunningham Collector; (1) MEX: Baja Cal. Sur, 9.5 mi. W. hwy 1 on Ramal Sn. Antonio de la Sierra, IX-19-1988: E. G. Riley, at blacklight; (1) MEX: Baja CA Sur, 10-22 km S. La Paz , Rt. 1, 16/VIII/94, J. Huether; (7) Mexico, Baja California Sur, 10 mi. N Cabo San Lucas nr. El Salto de San Nicolas , 1500', IX-4-1990, Blacklight, F. Andrews, T. Eichlin & A. Gilbert collectors; (3) MEX: Baja Calif. Sur, 4.2 mi. W Miraflores , IX-30-1981, F. Andrews & D. Faulkner, coll. at blacklight; (1) MEX: Baja Calif. Sur, El Triunfo , X-3-1981, F. Andrews & D. Faulkner, coll. at blacklight; (1) MEX: Baja Cal Sur, 9.4 mi. W hwy 1 on Ramal a San Felipe, IX-10- 88, A.J. Gilbert; (9) MEX: Baja Calif. Sur, Playa Los Cerritos , 11.2 mi. S Todos Santos, IX-28-1981, BLACKLIGHT, F. Andrews & D. Faulkner; (14) MEX: Baja Calif. Sur, 2.3 mi. SW San Bartolo , X-1-1981, F. Andrews & D. Faulkner, coll. at blacklight; (5) MEX: Baja Calif. Sur, 17.1 mi. S La Paz , IX-27-1981, F. Andrews & D. Faulkner; (3) BAJA CALIFORNIA, Playa Pedrito , 3 mi. S. Todos Santos, 14-XI-1981, Robert Gordon. ( CSCA) ( RCCC) ( TAMU) ( UCRC) ( USNM) .

Remarks. Glaresis australis is a comparatively large species of the mendica group distinguished from all other species by the strongly reticulate head surface; lack of posterosuperior metafemoral tooth; and large metatibial posteromedian lateral projection. All specimens examined were from Baja California Sur, although G. australis may occur on adjacent mainland Mexico.

Etymology. This species name is the Latin australis , meaning south, and refers to the Baja California Sur type locality.

Figures

Figures 29A-29G. Glaresis australis, new species. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum with metasternal groove and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.