Fissidentoblatta polyphagella ( Bey-Bienko 1963 ) Bohn, 2024

Bohn, Horst, 2024, The spine armament of the legs as an important means for the characterisation of the genera of Corydiinae and their relationships (Blattodea, Corydiidae), Zootaxa 5482 (1), pp. 1-79 : 39-40

No. 45. Fissidentoblatta polyphagella ( Bey-Bienko 1963) , comb. nov.

Figs. 3G , 14G,H

Arenivaga (Psammoblatta) polyphagella Bey-Bienko, 1963: 252 , fig. 1; Princis 1971: 1136.

Male characters. Tibia armament: [2.7.0][7.7.2][11.7.4/5], dorsal spines of hindtibia fairly evenly distributed. The right fronttibia has three dorsal spines, two in dorso-posterior and one in dorso-anterior position, resulting in 10 spines as total number, which is unusual for Palaearctic genera; on the left fronttibia there is only one spine on each side. The short distance and the smaller size of the two spines suggest that wounding might have caused a splitting and doubling of the spine.—Tarsal arolia: present.—Subgenital plate: with styli, Fig. 14G .—Right phallomere: Fig. 3G , 14H .

Female: unknown.

Material studied: Holotype, 1♂, Кух-Шер ДерваЗе, 1860 m, 20.IX.1957, К. ЛИндберг (prep.: Bo 1399). (M. St.Petersburg).

Species of unknown position

No. 46. “ Polyphaga” kruegeri Salfi, 1927

Polyphaga kruegeri Salfi, 1927: 231 , fig. 3.

Hetrogamodes kruegeri : Chopard 1929: 334, Figs. 10–11 , 68–70, 78–80, 88.

Arenivaga (Heterogamisca) kruegeri : Princis 1962: 69.

Hemelytroblatta kruegeri : Grandcolas 1994a: 157.

Male characters (according to Salfi). Wings: fully developed, tegmina without subcosta lobe.—Femur armament: all femora without apical spine.—Tibia armament: [1.8.0][7.7.0][10.7.4], dorsal spines of hindtibia evenly distributed.—Tarsal arolia: absent.

Remarks. Type material not traceable; already Chopard (1929) did not see the type specimen and could only refer to Salfi’s description, which does not contain informations about the subgenital plate concerning the distribution of bristles or the presence or absence of styles. According to the tibia armament it could either be a Mollidentoblatta or a Psammoblatta . In the Paris Museum there are two male specimens from Djanet (SE Algeria near the border to Libya) determined by Chopard as Heterogamodes kruegeri ; but the armament of the midtibia [6.8.1] excludes its belonging to this species. The specimens are described herein as a new species, Heterogamodes (Heterogamisca) sulcata (No. 7).

Female: unknown

Material, presumably lost. Type, 1♂, Libya, Porto Bardia , 20.VIII.1925 .

No. 47. “ Heterogamodes” incerta Chopard, 1929

Heterogamodes incerta Chopard, 1929: 306 , fig. 57.

Arenivaga (Psammoblatta) incerta : Bey-Bienko 1950: 319; Princis 1962: 68.

Characters of female. Femur armament: all femora without apical spine.—Tibia armament: [1.8.0][7.7.1][10.7.4], dorsal spines distributed in two groups (7+3). The armament of mid- and hindtibia slightly deviates from the pattern of P. africana , in contrast to Chopard’s statement.

Other characters apart from leg armament not known, since the male gender of the species is still unknown. According to femur and tibia armament either belonging to Mollidentoblatta or Psammoblatta .

Material studied: Type, 1♀, Turkestan , Sikora. (M. Wien) .

No. 48. “ Heterogamia” hirsuta Saussure, 1899

Heterogamia hirsuta Saussure, 1899: 583 .

Heterogamodes hirsuta : Chopard 1929: 301.

Arenivaga (Psammoblatta) hirsuta : Princis 1962: 67.

Remarks. Species described by Saussure after a single female from southern Africa. Chopard (1929) had already— without effect—tried to get the type specimen for study; it is obviously lost. Since the original description does not contain any information concerning the femur and tibia armament, it is not possible to place the species in one of the known genera. Chopard (1929) included it in his genus Heterogamodes on account of the dense coverage with long hairs.

Figures

FIGURE 3. The right phallomere and its parts. A: Complete right phallomere of Type 1 in standard view, seen from the left (inner face) towards the right (outer face); the position of the other two axes of the body are indicated by the labels ant(erior)— post(erior) and dors(al)—ventr(al). The three sclerites of the phallomere are R1M, R2, and R3. The largest dent of R2 is called main dent (MD), in this case stepped in two etages, upper etage (ue). R1M with a tubercular ridge (tr), the two arrows point to an edge where R1M and R2 articulate. B: Most posterior part of Type 6 phallomere in ventral view, showing the frontside of R1M and the backside of the main dent, with its vertical arm (va) in focus, the horizontal arm (ha) out of focus. C,D: Isolated R1M sclerite, with the sclerotised part (sc) at the left, in D (Type 1) with a tubercular ridge (tr), the membraneous part at the right, in C (Type 2) with a gutterlike deepening (g), arrows indicate where R1M articulates with R2. E–G. Phallomeres in ventral view showing the backside of the main dent, R1M with a ridge (r). E: Phallomere Type 1/2, r of R1M as in Type 2 without tubercular elevations, ue in outline rounded as in Type 1, but with a small lobelike projection (l) recalling the pointed tip of Type 2. F,G: Type 3 phallomeres, main dent with a deep incision (arrow).—Species: A: Heterogamisca arenarum, Type; B: Psammoblatta minuta, Type; C: Mollidentoblatta persica, Holotype; D: Heterogamisca longipilosa, Holotype; E: Atuberculoblatta intermedia, B 1416; F: Fissidentoblatta afghana, Paratype; G: F. polyphagella, Holotype.—Enlargements: Scale in B 0.1 mm, in all others 0.25 mm.

FIGURE 10. Subgenital plate (at the left, ventral view) and posterior part of the right phallomere (at the right: B ventral view, D,F,H standard view) of male.—Species:A,B: Mollidentoblatta hebraica, Bo 1364; C,D: M. latifrons, Paratype, Bo 1411; E,F: M. marismortui, Bo 1362; G,H: M. marmorata, Syntype, Bo 1382.—Phallomere type: Type 2 in all corresponding figures.— Explanations: R1M, R2, R3 sclerites of the right phallomere, MD main dent of R2, ue its upper etage, in ventral view (B) showing the pointed upper end; s stylus.—Enlargements: Scale 0.25 mm.

FIGURE 11. Subgenital plate (at the left, ventral view) and posterior part of the right phallomere (at the right, standard view) of male.—Species: A,B: Mollidentoblatta oblonga, Type, Bo 1393, right stylus lost; C,D: M. persica, Holotype, Bo 1385, dents of R2 in D isolated; E,F: M. pilosella. (E) Holotype, Bo 1426; (F) Bo 1410; G,H: M. rugosa, Bo 1389.—Phallomere type: Type 2 in all corresponding figures.—Explanations: R1M, R2, R3 sclerites of the right phallomere, MD main dent of R2, ue its upper etage; s stylus—Enlargements: Scale 0.25 mm.

FIGURE 14. Subgenital plate (at the left, ventral view) and posterior part of the right phallomere (at the right, standard view) of male.—Species: A,B: Psammoblatta roseni. (A) Bo 1332; (B) Bo 1420; C,D: P. subhyalina. (C) Paratype, Bo 1383; (D) Cotype, R1M broken; E,F: Fissidentoblatta afghana, Paratype, Bo 1398; G,H: F. polyphagella, Holotype, Bo 1399.—Phallomere types: B,D:Type 6; F,H: Type 3.—Explanations: R1M, R2, R3 sclerites of the right phallomere, MD main dent of R2, in (B, D) not stepped, with a horizontal (ha) and vertical arm (va), in (F, H) with a deep incision or step (arrowheads); d denticles on the surface of R1M, s stylus.—Enlargements: Scale 0.25 mm.