Exocelina takime (Balke, 1998)

Shaverdo, Helena, Surbakti, Suriani, Warikar, Evie L., Sagata, Katayo & Balke, Michael, 2019, Nine new species groups, 15 new species, and one new subspecies of New Guinea diving beetles of the genus Exocelina Broun, 1886 (Coleoptera, Dytiscidae, Copelatinae), ZooKeys 878, pp. 73-143 : 73

18. Exocelina takime (Balke, 1998) Figs 39 , 44

Copelatus (Papuadytes) takime Balke, 1998: 336; Nilsson 2001: 77 (catalogue).

Papuadytes takime (Balke, 1998): Nilsson and Fery 2006: 56 (comb. nov.).

Exocelina takime (Balke, 1998): Nilsson 2007: 34 (comb. nov.).

Type locality.

Indonesia: Papua Province: Pegunungan Bintang Regency, Bime, ca. 04°20'S, 140°12'E, 1400 m a.s.l.

Type material studied.

Holotype: male "IRIAN JAYA: 11.9.1993 Bime - Calab Gebiet, Bime, 1400m", "leg. M. Balke (12) ca. 140°12'E 04°20'S ", “HOLOTYPUS” [red], " Copelatus takime Balke des. 1997" [red] (NHMW). Paratypes: 10 males, 14 females with the same label as the holotype, one female with an additional green label "DNA M.Balke 3291" (NHMW). 11 males, 2 females "IRIAN JAYA: 22.9.1993 Bime - Calab Gebiet, Bime, 1400m", "ca. 140°12'E 04°20'S, leg. Balke (16)", one of the males with an additional green label "DNA M.Balke 3290" (NHMW). 3 males, 1 female "IRIAN JAYA Zentralmassiv 140°25'E 04°24'S ", "Kali Takime, 900m 18.8.1992 leg. Balke (17)" (NHMW). 7 males, 6 females "IRIAN JAYA: 29.9.1993 Eme Gebiet Emdoman, 800m", "ca. 139°55'E 04°14'S, leg. M. Balke (24)" (NHMW). All these specimens are with red paratype labels "PARATYPUS Copelatus takime Balke des. 1997" or "Paratypus Copelatus takime Balke des. 1997" [red]. Note: in the original description ( Balke 1998), 8 males and 4 females are reported from the locality 16; there are 11 males, 2 females in the NHMW, therefore, some mistakes were made in presenting the type material in the original description or/and during labeling the material.

Additional material.

2 females "IRIAN JAYA: 11.9.1993 Bime - Calab Gebiet, Bime, 1400m", "leg. M. Balke (12) ca. 140°12'E 04°20'S " (NHMW).

Diagnosis.

For complete description, see Balke (1998: 336). Beetle medium-sized: TL-H 4.1-4.5 mm; oblong-oval; dark brown to piceous, with brownish pronotal sides and head anteriorly; shiny, but with distinct dorsal punctation and weakly impressed microreticulation; pronotum with narrow lateral bead ( Fig. 39 ); male protarsomere 4 with large, thick, strongly curved anterolateral hook-like seta; male protarsomere 5 with anterior band of ca. 40 and posterior row of eight relatively long setae ( Fig. 44D ); median lobe robust, in lateral view, evenly curved to narrowly rounded, not curved downwards apex; in ventral view, with more strongly thickened lateral sides, distinctly expanded in middle and narrowing to broadly pointed apex; paramere with distinct dorsal notch and subdistal part well developed, with long and dense subdistal and inconspicuous proximal setae ( Fig. 44 A–C ).

Affinities.

In the area of its distribution, E. takime co-occurs with E. aipomek , E. ascendens and species of the E. bacchusi , E. ekari , E. aipo , E. okbapensis , E. casuarina , and E. danae groups. From species of the E. ekari group, the species differs in larger size, evidently stronger dorsal punctation, and the shape of the median lobe. In the latter character, E. takime differs also from the species of the remaining groups. For separating it from some of these species, also presence of the pronotal bead and simple male antennae, and shape and setation of the paramere can be used. For affinities within the group, see the “Key”.

Distribution.

Indonesia: Papua Province: Pegunungan Bintang Regency. The species is known only from the type material, i.e., Borme - Bime - Emdoman area ( Fig. 45 ).

Figures

Figure 45. Map of the eastern part of New Guinea showing distribution of the species of the E. larsoni and takime groups.

Figures 35 - 39. Habitus and colouration 35 Exocelina larsoni (Balke, 1998) 36 E. nomax (J. Balfour-Browne, 1939) A holotype B specimen from Tapini 37 E. warahulenensis sp. nov. 38 E. mianminensis sp. nov. 39 E. takime (Balke, 1998).

Figures 43, 44. 43 Exocelina mianminensis sp. nov. 44 E. takime (Balke, 1998) A median lobe in ventral view B median lobe in lateral view C paramere in external view D male protarsomeres 4 - 5 in ventral view.