Euphaea pahyapi Hämäläinen, 1985

Phan, Quoc Toan, Kompier, Tom, Karube, Haruki & Hayashi, Fumio, 2018, A synopsis of the Euphaeidae (Odonata: Zygoptera) of Vietnam, with descriptions of two new species of Euphaea, Zootaxa 4375 (2), pp. 151-190 : 176-177

Euphaea pahyapi Hämäläinen, 1985

( Figs 15d , 18f , 19a–d )

Euphaea pahyapi: Hämäläinen (1985) , pp. 341–344, Figs 1–4 ; Kitagawa & Katatani (2003), p. 16, Fig. 4 ; Hämäläinen (2017), p. 11.

Euphaea khaochongensis: Asahina (1985) , pp. 27–29, Figs 25–28.

Euphaea ochracea: Bui (2007) , pp. 1, 6 (photos) [Specimens from Phu Quoc]; Kosterin et al. (2012), p. 153, Fig. 2 (Specimens from Kbal Chhay Cascades, Preah Sihanouk Province).

Euphaea sp.: Kosterin (2016), p. 19.

Materials examined. [ Vietnam] 1♂, Phu Quoc Isl. , Kien Giang Prov., 30.XII.2015, T. Kompier leg. (TKC) ; 2♂, same location and collector, 14.IV.2016 (TKC).

Other materials confirmed by field observations. [ Vietnam] 6 exs, 19–20.VIII.2016, Phu Quoc Isl. , Kien Giang Prov., T. Kompier.

Notes. This species is newly recorded for Vietnam, although Bui (2007) published the same taxon first as Euphaea ochracea . Hämäläinen (1985) described Euphaea pahyapi from the peninsular part of Thailand based on male specimens. Asahina (1985) also described Euphaea khaochongensis from another province of the same area, but this was treated as a junior synonym of E. pahyapi by Pinratana et al. (1988) and in Hämäläinen & Pinratana (1999). Euphaea pahyapi can be distinguished from all Vietnamese Euphaea species by the unique completely and evenly amber-colored wings of the male ( Fig. 18d ). Color pattern of the head is variable, entirely black as in the original description ( Hämäläinen 1985) or with dirty blue markings as in Asahina's (1985) description and our specimens from Vietnam (Phu Quoc Island) ( Fig. 19b ). Abdomen and anal appendages are as in Euphaea masoni , with hairs only on S3 ( Fig. 15d ). Vesicle of Euphaea pahyapi differs from that of E. masoni by being heart-shaped, pointed at its posterior margin ( Fig. 18f ) whereas it is ax-shaped with the posterior margin rounded in E. masoni ( Fig. 18e ). Female of this species is still undescribed.

The record of Euphaea ochracea from Kbal Chhay Cascades of Preah Sihanouk Province, Cambodia in Kosterin et al. (2012) is a misidentification and concerns in fact E. pahyapi ( Kosterin 2016, and pers. comm.)

Distribution. Vietnam (Phu Quoc Island), Thailand (Ranong, Krabi, and Trang Provinces) ( Hämäläinen 2017), Cambodia (Preah Sihanouk Province) ( Kosterin et al. 2012, and Kosterin pers. comm.).

Figures

FIGURE 2. Anisopleura qingyuanensis. [2a–d] ♂ and [2e–f] ♀, Tu Le. (2a) head and thorax; (2b) head, oblique-dorsal view; (2c) appendages, dorsal view; (2d) appendages, lateral view; (2e) head and thorax; (2f) head, oblique-dorsal view.

FIGURE 4. Bayadera bidentata. [3a–e] ♂, Tam Dao and [4f–h] ♀. (4a) head and thorax; (4b) appendages, dorsal view; (4c) appendages, ventral view; (4d) appendages, lateral view; (4e) wings; (4f) head and thorax, Guangdong, China; (4g) head and thorax, Guangxi, China, and Vietnam; (4h) abdominal tip.

FIGURE 18. Euphaea spp. [18a–f] vesicle of male and [18g–j] prothorax of female, dorsal view. (18a, b, g) E. saola sp. nov. from (18a) A Luoi and (18b) Chu Mom Ray; (18c, h) E.guerini, Bhalee; (18d, i) E. hirta, Nam Nung; (18e, j) E.masoni, Lang Van; (18f)E. pahyapi, Phu Quoc.

FIGURE 19. Euphaea pahyapi. [19a–d] ♂, Phu Quoc. (19a) head and thorax; (19b) head, antero-dorsal view; (19c) appendages, lateral view; (19d) wings.

FIGURE15.Schematic drawingsof abdominal hair tufts inlateral view ofmale Euphaeaspp.(15a) E.saola sp.nov.; (15b)E. guerini; (15c) E.hirta; (15d) E.masoni and E. pahyapi. I–X show abdominal segments S1–10.

FIGURE1. Anisopleurabipugio. [1a–c] ♂,A Luoi and [1d–h]♀, Hon Ba.(1a)head and thorax;(1b)appendages,dorsal view; (1c) appendages, lateral view; (1d) body; (1e) head, frontal view; (1f) prothorax,lateral view; (1g) prothorax, dorsal view; (1h) basal half of HW.

FIGURE3. Anisopleurayunnanensis. [3a–c] ♂,Yunnan and [3d]♀,Sapa.(3a) head and thorax; (3b) appendages, dorsalview; (3c) appendages, lateral view; (3d) head and thorax.