Dryopteris Adans. sect. Lophodium (Newman) C.Chr. ex H.Itô

Roux, Jacobus P., 2011, The fern genera Dryopteris and Nothoperanema (Dryopteridaceae) in Madagascar and neighbouring Indian Ocean islands, including Saint Paul, Adansonia (3) 33 (1), pp. 7-67 : 20

Dryopteris Adans. sect. Lophodium (Newman) C.Chr. ex H.Itô

Nova flora japonica 4: 65 (1939). — Lophodium Newman in New Phytologist 4: 371, app. XVI (1851). — Dryopteris Adans. group Lophodium (Newman) C.Chr. , Index filicum: XXI (1905). — Lectotype: Lophodium spinulosum (O.F.Muell.) Newman ( Polypodium spinulosum O.F.Muell. ), designated by Christensen (1905: XXI), now Dryopteris carthusiana (Vill.) H.P.Fuchs.

Aspidium Sw. group Spinulosa H.Christ, Farnkräuter der Erde: 261 (1897), nom. nud. (McNeill et al. 2006: Art. 32.1[d]). — Type: none indicated. Two species – A. spinulosum Sw. and A. aemula (Soland.) Sw. are listed. Aspidium spinulosum may be considered as the type.

REMARKS

A group of about 10 species chiefly occurring throughout the northern temperate regions of the world, Macaronesia, and northern Africa, with few species extending to the southern hemisphere.

Diagnostic features of the section are the short, erect or decumbent rhizome that is closely and irregularly branched and which is set with roots and closely spaced trophopods. The lamina is up to 3-pinnate and the lobes are strongly dentate. The fertile vein branches are frequently shortened and end in the sori, or they extend for a distance beyond the sorus. Indumentum is composed of glands, moniliform hairs, and scales, the scale apex and marginal outgrowths generally end in a long moniliform series of cells. Fraser-Jenkins (1986: 195) noted that the minutely spinulose perispores may be unique for the section.