Cribellopsis moulladei ( Saint-Marc, 1974 ) Schlagintweit, 2020

Schlagintweit, Felix, 2020, Cribellopsis Moulladei (Saint-Marc, 1974) Nov. Comb. (Foraminiferida, Orbitolinidae): An Albian Marker Taxon Of The Southern Neotethysian Margin, Acta Palaeontologica Romaniae 16 (1), pp. 37-41 : 37-38

Cribellopsis moulladei ( Saint-Marc, 1974) comb. nov.

Fig. 1a–s

1974 Simplorbitolina moulladei n. sp. – Saint-Marc, p. 225, pl. 2, figs. 3-8.

1977 Coskinolina sunnilandensis Maync – Prestat, pl. 12, figs. 1–2.

1977 Coskinolina sp. – Prestat, pl. 12, figs. 3–7, 14–17.

1977 Iraqia sp. – Prestat, pl. 12, figs. 8–10 (B-forms).

1981 Paracoskinolina prestati n. sp. – Peybernès and Delmas, 81, figs. 1–7.

1982 Paracoskinolina prestati nom. nud. – Peybernès, pl. 2, figs. 9–12.

1989 Cribellopsis arnaudae n. sp. – Chiocchini. p. 42, pls. 1–2.

2012 Cribellopsis arnaudae Chiocchini – Chiocchini et al., pl. 69, fig. 1 pars, figs. 2–10.

2018 Cribellopsis aff. arnaudae Chiocchini – Cruz Abad, p. 111, figs. 92–93.

2019 in press Cribellopsis arnaudae Chiocchini – Schmitt et al., fig. 5D.

Descriptions: Test finely agglutinating, medium- to high conical. The cone base is almost flat in juvenile specimens, becoming convex in adult forms. Large sized, medium-conical specimens, which may show a slight depression in the central part of the base are interpreted as microspheric forms ( Fig. 1j, r ). The megalospheric embryo is situated at the rounded apex or positioned slightly eccentrically ( Fig. 1a–f ,?h). Sometimes a twochambered simple (= no partitions) embryo consisting of a subspherical protoconch and a smaller hemispherical deuteroconch is observable ( Fig. 1b–c ,?h). There are up to ~25 uniserial chambers in adult specimens. The welldeveloped marginal zone is subdivided by two orders of beams, primary beams and up to 3 short intercalary beams ( Fig. 1 o-p, s). The primary beams are either straight or slightly undulating. Inwards they fuse laterally thereby forming a loose central network ( Fig. 1n, p, s ). In axial sections, they display an alternating position between subsequent chambers and a tendency to thicken in direction to the apex resulting in a triangular shape. In shallow tangential sections the beams exhibit an alternating rectangular pattern ( Fig. 1c, h, l, q ). The foramina distributed in the central zone are predominantly arranged oblique but may be also vertical.

Dimensions: see Table 1 for Cribellopsis moulladei (Saint-Marc) nov. comb., compared to proposed synonyms. Further comments are provided in the subchapter "Comparisons".

Comparisons: The following comparisons and remarks essentially refer to the proposed synonymy with the species Paracoskinolina prestati , Cribellopsis arnaudae , and a form recently described by Cruz-Abad (2018) as Cribellopsis aff. arnaudae . It is worth mentioning that all the mentioned taxa do not show any structural differences and cannot satisfactorily be separated biometrically ( Table 1 ; and further comments in the text). For " Simplorbitolina " moulladei , no section displaying the embryo was provided. Saint-Marc (1974) just noted an initial spire of 3 to 4 chambers, but a simple (= non-septulate) bilocular embryo (protoconch + deuteroconch) is barely recognizable in the specimen illustrated in his plate 2, figure 6 (re-illustrated in Fig. 1h ). A slightly eccentric biloculine embryo is reported from both " Paracoskinolina " prestati (see Fig. 1b ), and Cribellopsis arnaudae ( Fig. 1c ). The differences of the mentioned taxa refer to the dimensional data as indicated in the original works.

Figures

Fig. 1 Cribellopsis moulladei (Saint-Marc) nov. comb., Albian of Somalia (a–b, j–l, r), Italy (c–g, m, p–q, s), and Lebanon (h–i, n–o; type material). a–f Axial sections of megalospheric specimens. g, r Subaxial sections of possibly microspheric specimens. h–i, k Subaxial sections of megalospheric specimens. j, m Tangential-oblique sections. l, q Tangential sections. n–p, s Transverse sections. a–b, k extracted from Peybernès & Delmas (1981, pl. 1, fig. 1–2, h) as Paracoskinolina prestati; j, l extracted from Peybernès (1982, pl. 2, fig. 12, fig. 11) illustrated as Paracoskinolina prestati; c, f–g, m, p, q, s extracted from Chiocchini (1989, pl. 1, fig. 14, fig. 5, pl. 2, fig. 8, pl. 1, fig. 11, fig. 10, fig. 12, fig. 4) illustrated as Cribellopsis arnaudae n. sp.; d extracted from Schmitt et al. (2019, in press, fig. 5d) illustrated as Cribellopsis arnaudae Chiocchini. e extracted from Cruz-Abad (2018, fig. 93J) illustrated as Cribellopsis aff. arnaudae Chiocchini; h–i, n–o extracted from Saint-Marc (1974, pl. 2, figs. 6, 3–4, 7) illustrated as Simplorbitolina moulladei n. sp. r extracted from Prestat (1977, pl. 12, fig. 10) illustrated as Iraqia sp. Abbreviations: b = beam (or main partition), deu = deuteroconch, ib = intercalary beam, pr = proloculus