Cranfillia bakeri (C.Chr.), 2020

Bauret, Lucie, Vázquez, Rubén, Molino, Sonia, Gaudeul, Myriam, Rakotondrainibe, France, Gasper, André Luís de & Rouhan, Germinal, 2020, New molecular and morphological evidences favor a combination of Blechnum bakeri C. Chr. in Cranfillia Gasper & V. A. O. Dittrich (Blechnaceae, Polypodiopsida), thus extending the distribution of Cranfillia to Madagascar and East Africa, Adansonia (3) 42 (18), pp. 279-289 : 286

Cranfillia bakeri (C.Chr.)

Vázquez Ferreira & S.Molino, comb. nov.

Blechnum bakeri C.Chr. , Index Filicum : 151 (1905), nom. nov. for Lomaria pubescens Baker ; Journal of the Linnean Society, Botany 15: 415 (1876), non Kunze (1843); nec Blechnum pubescens Desv. (1827) , nec Hooker (1837). — Austroblechnum bakeri (C.Chr.) Gasper & V.A.O.Dittrich, Phytotaxa 275 (3): 191-227 (2016). — Type: Madagascar, Antananarivo, W. Pool s.n. p.p. (lecto-,K[K000424012]!, first-step designated bySchelpe [1967: 348], second-step designated here; isolecto-, K[K000424011]! excl. specim.: fronds annotated as G and H).

Blechnum polypodioides (Sw.) Kuhn var. holstii Hieron., Pflanzenwelt Ost-Afrikas und der Nachbargebiete 1, C: 81 (1895). — Type: “usb., Mbaramu”, C . Holst 2479 (lecto-, B200031479 !, designated by Parris (2006: 4); isolecto-, P [ P00113566 ]!) .

Blechnum bakeri C.Chr. var. glabrum Bonap. , Notes ptéridologiques 7: 210 (1918) (as “glabra”). — Type: Madagascar. Mandraka, C. d’Alleizette 82 in Herb. Bonaparte (holo-, P [ P00483198 ]!) .

Blechnum ivohibense C.Chr., Archives de Botanique (Caen) , Bulletin mensuel 2: 211-212 (1928). — Type: Madagascar. Pic d’Ivohibe , Bara , H. Humbert 3300 p.p. in Herb. C. Christensen (holo-, BM[ BM001066246 photo]!; iso-, B[ B200031506 photo]! p.p., excl. specim: two small individuals on the right and left; excl. specim. P01625371 !). SeeRakotondrainibe et al. (2013) for details.

Blechnum umbrosum Peter, Repertorium specierum novarum regni vegetabilis, Beiheft 40 (1), Lieferung 1: 82; Descriptiones : 9, t. 3: 5-8 (1929). — Type: Tanzania. West Usambara , Kisimba above Mazumbai, IV.1916, A. Peter 16489 (lecto-, B[ B200034334 photo!], designated by Parris (2006: 4); isolecto-, BM[ BM000585222 photo]!, BR[ BR0000008054487 photo!], GOET007151 photo!, K[ K001092775 photo!], K[ K001092776 photo!], P[ P00483201 ]!, US00135471 photo!).

Blechnum ivohibense C.Chr. var. hirsutum C.Chr., Dansk Botanisk Arkiv 7: 106 (1932). — Type : Madagascar. Pic d’Ivohibe, H. Humbert 3299 (lecto-, BM[ BM001066247 photo!]; designated by Rakotondrainibe et al. (2013: 151-193); isolecto-, P[ P01625808 ]!); remaining syntypes: Analamazaotra, H. Perrier de la Bâthie 7550 (syn-, P[ P00835613 !, P01625804 !]); H. Perrier de la Bâthie 7625 (syn-, P[ P01625802 !, P01625803 !]).

NOMENCLATURAL COMMENTS

Schelpe (1967) cited two sheets at K as the lectotype of Blechnum bakeri , and only conducted the valid first-step lectotypification of the name (Art. 9.17 of ICN; Turland et al. 2018). Since those two sheets at K are not clearly labeled as being part of a single specimen (Art. 8.3 of ICN; Turland et al. 2018), we here conducted the second-step lectotypification, selecting the sheet with two isolated sterile fronds (annotated E, F). From the same sheet K000424012, the two fertile fronds (annotated G, H) are excluded from the lectotype, as they must be following the first-step lectotypification ( Schelpe 1967); we however ascribed the fertile frond H to Cranfillia bakeri Vázquez Ferreira & S.Molino , comb. nov., while frond G was ascribed to Lomaridium biforme (Baker) Gasper & V.A.O.Dittrich.

In the protologue of Blechnum bakeri var. glabrum, Bonaparte (1918) cited C. d’Alleizette 82 (in the Bonaparte’s Herbarium) as the type. This gathering is represented by a single sheet at P, which is the holotype; Rakotondrainibe et al. (2013) erroneously cited P00483199 as an isotype, as this latter sheet in fact belongs to the distinct gathering C. d’Alleizette 82m (i.e. including “m” as a suffix).