Comibaena birectilinea, Han & Galsworthy & Xue, 2012
publication ID |
https://doi.org/ 10.1111/j.1096-3642.2012.00826.x |
persistent identifier |
https://treatment.plazi.org/id/0389DF41-FF89-A922-FC09-AA96FBDBFC7B |
treatment provided by |
Marcus |
scientific name |
Comibaena birectilinea |
status |
sp. nov. |
COMIBAENA BIRECTILINEA SP. NOV.
( FIGS 36 View Figures 27–54 , 90 View Figures 79–90 , 140 View Figures 132–155 , 197 View Figures 188–199 )
Description: Antenna in male with basal threequarters bipectinate, outer rami longer than inner rami. Frons white, a small green patch present in the holotype. Labial palpus with upper and lateral surfaces brown, ventral surface white. Vertex green. Dorsal side of thorax and abdomen pale green, intersegment white on abdomen. Hind tibia with two pairs of spurs in both sexes, not dilated, without hair-pencil, terminal extension absent.
Forewing length: ♀ ♂ 16–17 mm. Forewing green, with white antemedial and postmedial lines, both straight, the antemedial inclined outwards, the postmedial inwards. Hindwing pale green, anterior part whitish, with white submarginal line parallel to outer margin; discal spot dark green. Both wings with terminal line and fringes concolorous with wing surface. Underside: forewing same as upperside; hindwing with outwardly curved postmedial line. Frenulum well developed.
Male genitalia. Uncus totally bifid, tapering, bent outwards. Socii longer than uncus, tapering but blunt. Valva simple, apex blunt; costa protruding at middle. Vinculum short, concave at middle, with branches short and broad. Aedeagus slender.
Female genitalia. Sterigma irregularly shaped. Antrum rounded, with posterior margin deeply concave, depth about half length of antrum. Ductus bursae quite short. Corpus bursae weak (incomplete in specimen examined).
Diagnosis: Comibaena birectilinea is similar to Comibaena cenocraspis and Comibaena hypolampes in the male genitalia and wing pattern. However, the antemedial line is distinctly inclined outwards, and close to the postmedial line on the inner margin. In the male genitalia, the costa is protruding and angled, and the lateral protrusions of the vinculum are shorter and broader. The female genitalia are similar to those of Comibaena cenocraspis , but the concavity of the antrum is shallower. The wing pattern of Comibaena birectilinea resembles the members of Thetidia , but the frenulum is present in this species, whereas it is usually absent in Thetidia .
Holotype, ♂, China: Tibet: Jomda , 3400 m, 31.vii.1976, coll. Zhang Xuezhong, IZCAS slide no. Geom-00124 ( IZCAS) . Paratypes ( IZCAS): China: 2♀, same locality as holotype, 26–27.vii.1976, coll. Han Yinheng, Zhang Xuezhong, IZCAS slide no. Geom-00125 ; 1♀, Sichuan: Batang, Yidun , 3350 m, 16.viii.1982, coll. Zhang Xuezhong ; 1♂, Batang (Tibet), Untere Urwaldzone , c. 3800 m, 5.viii.1936, H. Höne ( ZFMK) ; same locality and collector , 3♂, 7.viii.1936, 1♂, 8.viii.1936; 1♂, 14.viii.1936, 1♂, 15.viii.1936 ( ZFMK) ; 1♂, Batang (Tibet), Im Tal des Yangtse , c. 2800 m, 18.ix.1936, H. Höne ( ZFMK) ; 1♂, Tibet, Yargong Zambala, P. Soulié, 1903, ex. coll. Oberthür ( ZFMK) .
Distribution: China (Sichuan, Tibet). Etymology: A combination of the Latin prefix bi - and rectilinea, referring to the linear antemedial and postmedial lines on the forewing.
Species not occurring in China
Square brackets indicate nature of material examined.
Comibaena albimarginata ( Warren, 1893) View in CoL [specimen(s); male and female genitalia photographs, Holloway (1996: figs 248, 265)]
Uliocnemis albimarginata Warren, 1893: 355 View in CoL . ( India: Sikkim)
Comibaena amoenaria ( Oberthür, 1880) View in CoL [specimen(s)]
Phorodesma amoenaria Oberthür, 1880: 48 View in CoL , pl. 9, figure 4. ( Russia: Askold Island)
Comibaena bajularia bajularia ( Denis & Schiffermüller, 1775) View in CoL [male and female genitalia photographs, Hausmann, (2001: fig. 19)]
Geometra bajularia Denis & Schiffermüller, 1775: 97 View in CoL . ( Austria)
Phalaena ditaria Fabricius, 1776: 286 View in CoL . ( Germany)
Phalaena glauca Fourcroy, 1785: 267 View in CoL . ( France)
Phalaena pustulata Hufnagel, 1767: 520 View in CoL . ( Germany) [Junior homonym of pustulata Müller, 1764 View in CoL , Arctiidae .]
Comibaena bajularia tikhonovi Hausmann, 2000
Comibaena bajularia tikhonovi Hausmann, 2000: 302 , fig. d. ( Russia)
Comibaena barnsi barnsi Prout, 1930 View in CoL [specimen(s)]
Comibaena barnsi Prout, 1930: 14 View in CoL . ( Zaïre)
Comibaena barnsi equatorialis Debauche, 1941 View in CoL
Comibaena barnsi equatorialis Debauche, 1941: 302 View in CoL , pl. 10, figure 3; pl. 11, figure 6. ( Zaïre)
Comibaena biviaria Hampson, 1909 View in CoL [specimen(s)]
Comibaena biviaria Hampson, 1909: 126 View in CoL , pl. 4, figure 60. (Ruwenzori [south-west Uganda /north-east Zaïre])
Comibaena castaneata ( Warren, 1906) View in CoL [specimen(s)]
Comostolodes castaneata Warren, 1906: 87 View in CoL . ( Papua New Guinea)
Probolosceles agathia Warren, 1912: 80 View in CoL . ( Papua New Guinea) [specimen(s)]
Comibaena chalybeata Moore, 1868 View in CoL [specimen(s); male genitalia]
Comibaena chalybeata Moore, 1868: 639 View in CoL . ( India)
Comibaena cheramota ( Meyrick, 1886) View in CoL [specimen(s)]
Iodis cheramota Meyrick, 1886: 203 . ( Fiji)
Comibaena connata ( Warren, 1898) View in CoL [specimen(s)]
Probolosceles connata Warren, 1898: 15 View in CoL . ( Australia)
Comibaena coryphata ( Felder & Rogenhofer, 1875) View in CoL [specimen(s)]
Racheospila coryphata Felder & Rogenhofer, 1875 View in CoL : pl. 127, figure 10. ( South Africa: Knysna)
Comibaena diluta ( Warren, 1895) View in CoL [specimen(s)]
Colutoceras diluta Warren, 1895: 88 View in CoL . ( Japan)
Euchloris obsoletaria Leech, 1897: 238 View in CoL . ( Japan) [specimen(s); female genitalia]
Comibaena erythrospila Yazaki, 1998 View in CoL
Comibaena erythrospila Yazaki, 1998: 4 View in CoL , pl. 129, figure 7; text-figure 840. ( Nepal)
Comibaena esmeralda ( Warren, 1898) View in CoL [specimen(s)]
Probolosceles View in CoL ? esmeralda Warren, 1898: 15 View in CoL . ( Nigeria)
Comibaena flavitaenia ( Warren, 1898) View in CoL [specimen(s)]
Oenospila flavitaenia Warren, 1898: 13 View in CoL . ( Nigeria)
Comibaena hemictenes Prout, 1917 View in CoL [specimen(s)]
Comibaena hemictenes Prout, 1917: 114 View in CoL , pl. 7, figure 21. ( Ivory Coast)
Comibaena ingrata ( Wileman, 1911) View in CoL [specimen(s)]
Thalera ingrata Wileman, 1911: 343 View in CoL . ( Japan)
Comibaena insulana Inoue, 1986 View in CoL [specimen(s)]
Comibaena insulana Inoue, 1986: 53 View in CoL , figures 8a, b, 9a, b. ( Japan)
Comibaena integranota Hampson, 1893 View in CoL [specimen(s); male genitalia]
Comiboena integranota Hampson, 1893: 37 , 146, pl. 170, figure 13. ( Sri Lanka)
Comibaena leucochloraria ( Mabille, 1880) View in CoL [specimen(s)]
Phorodesma leucochloraria Mabille, 1880 View in CoL : xxii. ( Madagascar)
Comibaena leucospilata ( Walker, 1863) View in CoL [specimen(s)]
Geometra leucospilata Walker, 1863: 1554 View in CoL . ( South Africa)
Comibaena levequei Leraut, 2009 [specimens(s)]
Chlorochaeta longipennis Leraut, 2009: 46 View in CoL . ( Algeria)
Comibaena longipennis ( Warren, 1904) View in CoL [specimen(s)]
Chlorochaeta longipennis Warren, 1904: 464 View in CoL . ( Niger)
Comibaena mariae mariae ( Lucas, 1888) View in CoL [male genitalia]
Iodis mariae mariae Lucas, 1888: 1266 . ( Australia: Queensland)
Probolosceles albipunctata Warren, 1898: 15 View in CoL . ( Australia: Queensland) [specimen(s)]
Comibaena mariae syndyas Prout, 1916b View in CoL [specimen(s)]
Comibaena mariae syndyas Prout, 1916b: 205 View in CoL . (New Guinea)
Comibaena meyricki Prout, 1933 View in CoL [specimen(s); male genitalia]
Comibaena quadrinotata meyricki Prout, 1933: 92 View in CoL . (Talaut)
Comibaena multigruma Prout, 1920 View in CoL [specimen(s)]
Comibaena multigruma Prout, 1920: 299 View in CoL . ( Papua New Guinea)
Comibaena nepalensis Yazaki, 2000 [specimen and male genitalia photograph, Yazaki, (2000: pl. 162: 1; fig. 1248)]
Comibaena nepalensis Yazaki, 2000: 9 .
Comibaena pseudoneriaria Wehrli, 1926 View in CoL [male and female genitalia photograph, Hausmann (2001: fig. 20)]
Comibaena pseudoneriaria Wehrli, 1926: 163 View in CoL . ( Spain)
Comibaena punctaria ( Swinhoe, 1904) View in CoL [specimen(s)]
Probolosceles punctaria Swinhoe, 1904: 550 View in CoL . ( Madagascar)
Comibaena rhodolopha Prout, 1915 View in CoL [specimen(s)]
Comibaena rhodolopha Prout, 1915: 313 View in CoL . ( Zimbabwe)
Comibaena rhodonia Prout, 1913b View in CoL [specimen(s)]
Comibaena rhodonia Prout, 1913b: 416 View in CoL . (Irian Jaya)
Comibaena rufitornus Prout, 1916c View in CoL [specimen(s)]
Comibaena rufitornus Prout, 1916c: 275 View in CoL . ( Kenya)
Comibaena serrulata Fletcher, 1963 View in CoL [specimen(s); male genitalia]
Comibaena serrulata Fletcher, 1963: 116 View in CoL , pl. 8, figures 1–5 ( Iran)
Comibaena viridifimbria ( Warren, 1906) View in CoL [specimen(s); male genitalia]
Comostolodes viridifimbria Warren, 1906: 87 View in CoL . ( Papua New Guinea)
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
Kingdom |
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Phylum |
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Class |
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Order |
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Family |
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Genus |
Comibaena birectilinea
Han, Hongxiang, Galsworthy, Anthony C. & Xue, Dayong 2012 |
Chlorochaeta longipennis
Leraut P 2009: 46 |
Comibaena bajularia tikhonovi
Hausmann A 2000: 302 |
Comibaena nepalensis
Yazaki K 2000: 9 |
Comibaena erythrospila
Yazaki K 1998: 4 |
Comibaena insulana
Inoue H 1986: 53 |
Comibaena serrulata
Fletcher DS 1963: 116 |
Comibaena quadrinotata meyricki
Prout LB 1933: 92 |
Comibaena barnsi
Prout LB 1930: 14 |
Comibaena multigruma
Prout LB 1920: 299 |
Comibaena hemictenes
Prout LB 1917: 114 |
Comibaena mariae syndyas
Prout LB 1916: 205 |
Comibaena rufitornus
Prout LB 1916: 275 |
Comibaena rhodolopha
Prout LB 1915: 313 |
Comibaena rhodonia
Prout LB 1913: 416 |
Probolosceles agathia
Warren W 1912: 80 |
Thalera ingrata
Wileman AE 1911: 343 |
Comibaena biviaria
Hampson GF 1909: 126 |
Comostolodes castaneata
Warren W 1906: 87 |
Comostolodes viridifimbria
Warren W 1906: 87 |
Chlorochaeta longipennis
Warren W 1904: 464 |
Probolosceles punctaria
Swinhoe C 1904: 550 |
Probolosceles connata
Warren W 1898: 15 |
Probolosceles
Warren W 1898: 15 |
Oenospila flavitaenia
Warren W 1898: 13 |
Probolosceles albipunctata
Warren W 1898: 15 |
Euchloris obsoletaria
Leech JH 1897: 238 |
Colutoceras diluta
Warren W 1895: 88 |
Uliocnemis albimarginata
Warren W 1893: 355 |
Comiboena integranota
Hampson GF 1893: 37 |
Iodis mariae
Lucas TP 1888: 1266 |
Iodis cheramota
Meyrick E 1886: 203 |
Phorodesma amoenaria Oberthür, 1880: 48
Oberthur C 1880: 48 |
Comibaena chalybeata
Moore F 1868: 639 |
Geometra leucospilata
Walker F 1863: 1554 |
Phalaena glauca
Fourcroy AFD 1785: 267 |
Phalaena ditaria
Fabricius JC 1776: 286 |
Geometra bajularia Denis & Schiffermüller, 1775: 97
Denis M & Schiffermuller I 1775: 97 |
Phalaena pustulata
Hufnagel JS 1767: 520 |