Centris (Wagenknechtia) escomeli Cockerell, 1926

Vivallo, Felipe, 2013, Revision of the bee subgenus Centris (Wagenknechtia) Moure, 1950 (Hymenoptera: Apidae: Centridini), Zootaxa 3683 (5), pp. 501-537 : 509

Centris (Wagenknechtia) escomeli Cockerell, 1926

( Figures 5−8 and 40 )

Centris escomeli Cockerell, 1926 . 59: 28, 29 (Original description). Simpson, 1989. 14: 409, 410 (Cited). Toro et al., 1996. 23: 68, 69, 74 (Distribution, floral record). Chiappa, 1998. 22: 91 (Distribution). Montalva & Ruz, 2010. 35: 45 (List, distribution).

Centris (Wagenknechtia) escomeli ; Toro, 1986b. 13: 127 (Distribution). Cocucci et al., 1996. 34: 191 (Probable floral record). Vivallo et al., 2003: 78 (Distribution).

Centris (Penthemisia) garleppi ; Wagenknecht, 1971. 4: 279, 280 (Morphology, comparison with Centris nigerrima , distribution, floral record (misidentification)). Toro, 1986b. 13: 127 (List, distribution (misidentification)).

Comments. A complete redescription of the male and the description of the female of this species can be found in Vivallo et al. (2002), so here are indicated only diagnostic characters for the identification of both sexes of this species.

Diagnosis. Both sexes with blackish integument and brown pilosity ( Figs. 5–8 ). Female: fore elaiospathe located in the distal half of forebasitarsus. Hairs of hind elaiospathe almost overlapping. All claws with preapical tooth. Male: mandible long and slender, with the posterior margin of second and anterior edge of the third tooth broadly curved ( Fig. 7 ).

This is the only species of Centris (Wagenknechtia) with dark brown pilosity and without metallic reflections on the metasoma ( Figs. 6 and 8 ). The only two species with dark pilosity on the entire body are Centris rhodophthalma and C. vardyorum , but both species have intense bluish violet metallic reflections on the metasoma ( Figs. 22, 24 , 26 and 28 ).

Type material. Holotype male ( USNM, examined).

Type locality. Peru: Arequipa.

Floral records. See table 1.

Distribution. This species ranges from southern Peru (Arequipa) to the III Atacama Region (Quebrada de Paipote) in northern Chile. CHILE: ( Montalva & Ruz, 2010). II ANTOFAGASTA: Antofagasta ( Wagenknecht, 1971; Toro, 1986b; Chiappa, 1998; Vivallo et al., 2002). Quebrada de Cifuncho, Quebrada de Paposo, Quebrada de Taltal ( Toro et al., 1996; Vivallo et al., 2002). *Aguas Verdes, *Calama, *Chiuchiu, *Encanche, *La Negra, *Pampa Blanca, *Tumbre (Laskar). XV ARICA Y PARINACOTA: Chusmisa, Codpaquilla, Guatacondo, Parca, ( Wagenknecht, 1971; Vivallo et al., 2002). *Ariquilda, *Mamiña, *Quebrada de Guatacondo, *Socoroma, *Termas de Mamiña. * ATACAMA: Chañaral, Copiapó, Potrerillos, Quebrada de Paipote. III TARAPACÁ: ( Wagenknecht, 1971). *Iquique. PERÚ: ( Soukup, 1943; Vivallo et al., 2002, 2003; Montalva & Ruz, 2010). AREQUIPA: Arequipa ( Cockerell, 1926).

Material examined. CHILE: II ANTOFAGASTA: 1 male: Atacama Aguas Verdes 22/10/1968 P. Millas, col ( PUCV). 1 male and 1 female: Encanche 10/11/1974 Sielfeld, col\ Coll: Sielfeld/ MNHN Chile ( MNHN). 1 male: La Negra 12/92 H. Toro, col ( PUCV). 2 females: Antofagasta Chiuchiu 24/01/1972 H. Toro, col\ en Medicago sativa ( PUCV). 2 females: Antofagasta Chiuchiu 24/01/1972 M. Rojas, col ( PUCV). 2 females: Antofagasta Chiuchiu 24/01/1972 Montenegro, col\ en Medicago sativa ( PUCV). 4 males and 8 females: II R. 30 Km. E. Calama 01/1992 S. Rodríguez, col ( PUCV). 1 female: Calama Antofagasta Recarrhen 02/01/0972 R. Mendoza, col\ en Medicago sativa ( IDEA). 1 female: Tumbre N. W. Laskar Vol. Antofagasta Prov. 3500 mts. 11/1965\ Centris (W.) cf escomeli Cockerell, 1920 F. Zanella det., 1998 ( AMNH). XV ARICA Y PARINACOTA: 1 male: Quebrada Guatacondo, 2200 msnm. 15/03/1967 Loayza, col\ Coll. Cerda/ MNHN Chile ( MNHN). 1 male: Ariquilda N. E. Iquique 28-29/04/1969, L. Peña, col ( MNHN). 1 male: Tarapacá Guatacondo 09/1971 H. Toro, col\ en Medicago sativa ( CTMI). 1 female: Tarapacá Guatacondo 09/1971 L. Ruz, col\ en Medicago sativa ( CTMI). 1 male: Socoroma 16/04/1997 H. Vargas C., col ( IDEA). 1 male: Mamiña 19/03/1968 ( DZUP). 1 male: Guatacondo Tarapacá 19/09/1971 ( PUCV). 10 males and 2 females: Tarapacá Guatacondo 19/09/1971 L. Ruz, col\ en Medicago sativa ( PUCV). 6 males and 2 females: Tarapacá Guatacondo 09/1971 H. Toro, col\ en Medicago sativa ( PUCV). 4 males: Tarapacá Guatacondo 18/04/1971 Montenegro, col\ en Medicago sativa ( PUCV). 1 male: Tarapacá Guatacondo 19/04/1971 Montenegro, col\ en Medicago sativa ( PUCV). 4 males: Tarapacá Chusmisa 19/ 04/1971 H. Toro, col\ en Medicago sativa ( PUCV). 1 male: Tarapacá Chusmisa 19/04/1971 Montenegro col\ en Medicago sativa ( PUCV). 2 males: Tarapacá Codpaquilla 22/04/1971 W. Sielfeld, col ( PUCV). 1 male: Chile Region I 20 km W Chuzmisa 11.iv.00 L. Packer\ Centris (Wagenknechtia) cf. escomeli det. J. S. Ascher 2004 ( PCYU). 1 male: Tarapacá Codpaquilla 22/04/1971 Zuñiga, col ( PUCV). 1 male: Tarapacá Codpaquilla 22/04/ 1971 L. Ruz, col\ en Medicago sativa ( PUCV). III ATACAMA: 1 female: Chañaral Pampa Blanca 04/12/1990 R. Mendoza, col ( IDEA). 1 male: On Loasa \ Chile, Region II R 5 km 1006, Chañaral S26°14.97’ W70°28.83’ 19.xi.2002 J. Grixti & A. Zayed\ Centris (Wagenknechtia) cf. escomeli det. J. Ascher 2004 ( PCYU). 1 female: Potrerillos 15/01/1970 Sielfeld, col\ Coll: Sielfeld/ MNHN Chile ( MNHN). 1 male: Copiapó Quebrada de Paipote 17/10/1988 H. Vargas C. y D. Bobadilla G., col\ en Prosopis chilensis ( IDEA). 8 males and 2 females: Copiapó 15/ 01/1970 Jorquera, col\ Coll: Sielfeld/ MNHN Chile ( MNHN). I TARAPACÁ: 2 males: Iquique Termas de Mamiña 14/07/1977 H. Vargas, col ( IDEA). PERÚ: AREQUIPA: 1 male: Arequipa Peru (Escomel)\ Type No. 40493 U.S. N.M.\ Centris escomeli Ckll. Type \ USNM ENT 0 0 534193 ( USNM). 1 male and 3 females: Arequipa 12/1926 Dr. Escomel, col. Hemisia clypeata Fr. ( MACN). 1 male: Perú Arequipa 2200 m. 19/02/1955 A. Meza, coll ( CAS). 1 male: Arequipa 2200 m. 05/1955 A. Meza, col ( DZUP).

Figures

FIGURE 40. Distribution records for Centris escomeli Cockerell, 1926 and C. orellanai Ruiz, 1940.

FIGURES 5 – 8. Centris escomeli Cockerell, 1926. Fig. 5. Female (Guatacondo, Chile): Head, frontal view. Fig. 6. Habitus, lateral view. Fig. 7. Male (Guatacondo, Chile): Head, frontal view. Fig. 8. Habitus, lateral view. Scale bars 1 mm.

FIGURES 21 – 24. Centris rhodophthalma Pérez, 1911. Fig. 21. Female (El Pangue, Chile): Head, frontal view. Fig. 22. Habitus, lateral view. Fig. 23. Male (El Pangue, Chile): Head, frontal view. Fig. 24. Habitus, lateral view. Scale bars 1 mm.

FIGURES 25 – 28. Centris vardyorum Roig-Alsina, 2000. Fig. 25. Female (El Challao, Argentina): Head, frontal view. Fig. 26. Habitus, lateral view. Fig. 27. Male (El Challao, Argentina): Head, frontal view. Fig. 28. Habitus, lateral view. Scale bars 1 mm.