Araliaephyllum luciferum (KRYSHT.) GOLOVN.

Golovneva, Lina, Bugdaeva, Eugenia, Volynets, Elena, Sun, Yuewu & Zolina, Anastasia, 2021, Angiosperm Diversification In The Early Cretaceous Of Primorye, Far East Of Russia, Fossil Imprint 77 (2), pp. 231-255 : 247

Araliaephyllum luciferum (KRYSHT.) GOLOVN.

Text-fig. 10a–d

1929 Aralia lucifera KRYSHT. , p. 1360, pl. LVIII, figs 1, 2.

2018 Araliaephyllum luciferum (KRYSHT.) GOLOVN. , p. 23, fig. 6C, E, F, H.

L e c t o t y p e. No. 3013/1, coll. TSNIGR Museum (designated by Golovneva 2018), Southern Primorye, Suchan (Partizansk) coal basin, left bank of the Malaya Sitza (Postyshevka) River near Novoveselaya village, Severosuchan Formation, lower-middle Albian ( Kryshtofovich 1929: pl. LVIII, fig. 2).

D e s c r i p t i o n. Leaves of Araliaephyllum luciferum (KRYSHT.) GOLOVN. are simple, trilobate, fan-shaped, ovate and rounded in outline, 1.5–3.5 cm long, 1.3–2.6 cm wide, cuneate base, decurrent near the petiole, and serrate margin. Teeth are small, adpressed. Apices of lobes are acute or rounded. The central lobe is the largest; its length is equal to 1/4–1/2 of the blade length. Lateral lobes are shorter and narrower, usually directed to the leaf apex. Sinuses between lobes are shallow and narrow, with rounded apices. Venation is palmate, 3-nerved, brochidodromous. Midvein is straight. Lateral basal veins are slightly curved, diverging from the midvein at an angle of 20–35° in 3–5 mm above leaf margin, ending in the lobe apices. Basal veins bear several curved basiscopic branches, which form a series of loops near the margin. Three to five pairs of secondary veins diverge from the midvein at an angle of 35–55°, also forming loops. Tertiary venation is very thin and poorly preserved.

C o m p a r i s o n a n d r e m a r k s. Araliaephyllum luciferum differs from other species of this genus from eastern Russia in having a serrate margin. This character was reported for several species of Araliaephyllum or “ Aralia ” from Cenomanian Peruc flora of the Czech Republic, but all of them have deeper dissected leaf blades and longer lobes ( Velenovský 1882, 1884, Greguš and Kvaček 2015). Besides that, lobes of Araliaephyllum formosum (HEER) GREGUŠ et J.KVAČEK from the Maletín are serrate only in the apical parts. “ Aralia ” decurrens VELEN. from Vyšehořovice ( Velenovský 1884) has very narrow lobes; lateral lobes of “ Aralia ” minor VELEN. are significantly smaller than the medial ones, and the leaf base is not cuneate ( Velenovský 1884).

“ Aralia ” calomorpha SAPORTA from the Albian of Buarcos in Portugal ( Saporta 1894) also has a serrate margin, but differs from Araliaephyllum luciferum in having narrow lobes separated by deep incisions. This species was considered as a presumable report of lauraceous leaves ( Friis et al. 2011). Toothed leaves of “ Aralia ” wellingtoniana LESQ. from the Raritan Formation ( USA) differ in larger size (up to 20 cm), and in deeply trilobate blades, with long, lanceolate lobes ( Lesquereux 1892).

Figures

Text-fig. 10. a–d: Araliaephyllum luciferum (KRYSHT.) GOLOVN., Partizansk, Severosuchan Formation, Aptian, a – spec. TSNIGRM 3013/1, lectotype, b – spec TSNIGRM 3013/2, c – spec. TSNIGRM 3013/4, d – spec. TSNIGRM 3013/3; e, j: Pandanites ahnertii (KRYSHT.) GOLOVN., Konstantinovka, Lipovtsy Formation, Aptian, TSNIGRM 3013/5, e – fragment of leaf margin, j – general view, j1 – lectotype, j2 – leaf, figured in (e); f: Araliaephyllum ussuriense (KRASSILOV) GOLOVN., Andreev Inlet, Frentsevka Formation, early-middle Albian, spec. IBBS 28/103; g: Araliaephyllum vittenburgii GOLOVN. et VOLYNETS, Podgorodenka coalfield, Galenki Formation, early-middle Albian, spec. IBBS 41/1, g1 – spec. IBBS 41/1a, holotype, g2 – spec. IBBS 41/1b; h: Dicotylophyllum sp. 1, Podgorodenka coalfield, Lipovtsy Formation, Aptian, spec. IBBS 28L-4; i: Sapindopsis sp., 3rd Kamenka, Frentsevka Formation, early-middle Albian, spec. IBSS 325/8. Scale bar 5 mm in a–h and 1 cm in i, j.