Anthomyzinae, Czerny, 1903

ROHÁýEK, JindĜich, 2014, Reliquantha eocena sp. nov., first tertiary representative of an extant genus of Anthomyzidae (Diptera), Acta Entomologica Musei Nationalis Pragae 54 (2), pp. 773-784 : 782-783

Key to fossil (all Tertiary) taxa of the subfamily Anthomyzinae

1 Orbit with only 2 ors long. Mesonotum with both dc macrosetae situated close to scutellum ( ROHÁýEK 1998: Fig. 8 ). f 1 with ctenidial spine short but distinct (ROHÁ- ýEK 1998: ¿g. 11). Wing narrow and terminal section of CuA 1 long ( ROHÁýEK 1998: Fig. 10 ). Female T4–T7 ochreous to yellow, each with transverse dark brown posterior band ( ROHÁýEK 1998: Fig. 11 ); female preabdominal sterna very broad ( ROHÁýEK 1998: Fig. 12 ). ............... Grimalantha vulnerata Roháþek, 1998 (Dominican amber)

– Orbit with 3 ors long ( Fig. 6 ). Mesonotum with both dc macrosetae postsutural, situated more anteriorly ( Figs 4, 5 ). f 1 without ctenidial spine ( Fig. 8 ). Wing wider and terminal section of CuA 1 short ( Fig. 10 ). Preabdominal terga uniformly dark brown and female preabdominal sterna narrower ( ROHÁýEK 2013a: Fig. 15C). .................... 2

2(1) Antenna with 1st flagellomere strongly bent ventrally ( ROHÁýEK 2013a: Fig. 14E; Fig. 5 ). Frons with oc setae erect and mesonotum with two medial rows of ac microsetae situated very close to each other; also stpl setae arising close to each other ( ROHÁýEK 2013a: Fig. 14A). Male cercus reduced, very small ( ROHÁýEK 2013a: Fig. 15B). [ Lacrimyza Roháþek, 2013 ]. ............................................................................................ 3

– Antenna with 1st flagellomere less bent ventrally (more porrect) ( Figs 4 , 6 ). Frons with oc setae normal ( Fig. 6 ) and mesonotum with 4 ac rows regularly arranged; distance between stpl setae normal. Male cercus large and brown pigmented ( Figs 11, 12 ). ..... ................................................................ Reliquantha eocena sp. nov. (Baltic amber)

3(2) 1st antennal flagellomere whitish yellow to white, with long marginal pilosity; frons wider and paler anteriorly ( ROHÁýEK 2013a: Fig. 14A). Parafacialia and gena whitish yellow and silvery white microtomentose. Wing narrower ( ROHÁýEK 2013a: Fig. 14C), more than 3 times as long as wide. Male preabdomen with S3 the largest sternum. Gonostylus without setae on apex ( ROHÁýEK 2013a: Fig. 15A). .................................... ................................................... Lacrimyza lacrimosa Roháþek, 2013 (Baltic amber)

– 1st antennal flagellomere orange-yellow, with very short marginal pilosity; frons narrower and darker anteriorly ( ROHÁýEK 2013a: Fig. 16A). Parafacialia and gena ochreous-yellow and golden microtomentose. Wing wider ( ROHÁýEK 2013a: ¿g. 16E), less than 2.9 times as long as wide. Male preabdomen with S4 the largest sternum. Gonostylus with a few longer setae on apex ( ROHÁýEK 2013a: Fig. 16B). .............................. ................................................... Lacrimyza christelae Roháþek, 2013 (Baltic amber)

Figures

Figs 6–12. Reliquantha eocena sp. nov., male holotype. 6 – head dorsolaterally; 7 – right hind femur, posteriorly; 8 – left fore femur, posteriorly; 9 – venation in basal part of wing (different view than in Fig. 10); 10 – left wing; 11 – abdomen laterally (left side); 12 – postabdomen with genitalia, ventrocaudally. Reconstructed parts in dashed lines. All scales = 0.3 mm. For abbreviations see text (p. 774).

Figs 4–5. Baltic amber fossil Anthomyzidae. 4 – Reliquantha eocena sp. nov., male holotype, left laterally (body length 2.30 mm). 5 – Lacrimyza lacrimosa Roháþek, 2013, male holotype, left laterally (body length 2.22 mm) (adopted from ROHÁýEK 2013a: Fig. 13A). Photo by J. Roháþek.