Novalena Chamberlin & Ivie
publication ID |
https://doi.org/ 10.5281/zenodo.571683 |
publication LSID |
lsid:zoobank.org:pub:894795FB-ED38-4132-92B9-6AC8C3E41566 |
DOI |
https://doi.org/10.5281/zenodo.6031526 |
persistent identifier |
https://treatment.plazi.org/id/E45787C5-FF86-FFF2-E1A1-62E5FA1AFA73 |
treatment provided by |
Plazi |
scientific name |
Novalena Chamberlin & Ivie |
status |
|
Key to species of Novalena Chamberlin & Ivie View in CoL View at ENA
The males of N. attenuata , N. bipartita , N. bipunctata , N. costata , N. laticava , N. ajusco , N. alvarezi , N. atzimbo , N. cieneguilla , N. comaltepec , N. divisadero , N. durango , N. irazu , N. jiquilpan , N. orizaba , N. paricutin , N. plata , N. popoca , N. prieta , N. saltoensis , N. sinaloa , N. triunfo , N. victoria , and N. volcanes are unknown. The females of N. alamo , N. bosencheve , N. ivei , and N. ixtlan are unknown. Epigyna of N. bipartita and N. laticava were examined by photos but these two species are not included in the key. Novalena orizaba was not examined and also is not included in the key.
1 Males.............................................................................................. 2
- Females........................................................................................... 30
2(1) Metatarsus and tarsus I and II with four rows of long, curved hairs ( Fig. 29).......................... N. ixtlan View in CoL sp. nov.
- Metatarsus and tarsus I and II without four rows of long, curved hairs........................................... 3
3(2) RTA with ventral projection ( Figs. 3 View FIGURE 3 d, 12a, 16e, 21d, 28g, 39b)................................................ 4
- RTA without ventral projection ( Figs. 13 View FIGURE 13 b, 18b, 32b, 42b)................................................... 24
4(3) Dorsal projection of RTA with two sharp points directed ventrally ( Figs. 7 View FIGURE 7 b, 12b).................................. 5
- Dorsal projection of RTA without two sharp points directed ventrally ( Figs. 20 View FIGURE 20 e, 26b).............................. 6
5(4) Ventral projection of RTA prominent in ventral view ( Fig. 12 View FIGURE 12 a).................................... N. lutzi (Gertsch)
- Ventral projection of RTA small in ventral view ( Fig. 7 View FIGURE 7 a)............................. N. calavera Chamberlin & Ivie View in CoL
6(4) Median apophysis as long as wide ( Fig. 16 View FIGURE 16 d).................................................. N. alamo View in CoL sp. nov.
- Median apophysis longer than wide ( Figs. 28 View FIGURE 28 d, 39a)......................................................... 7
7(6) Dorsal projection of conductor larger than ventral projection ( Figs. 28 View FIGURE 28 d, 39a)..................................... 8
- Dorsal projection of conductor smaller than ventral projection or projections of similar size ( Figs. 3 View FIGURE 3 c, 31a).............. 9
8(7) RTA with ventral and dorsal projections large and with deep excavation ( Fig. 28 View FIGURE 28 e)...................... N. iviei View in CoL sp. nov.
- RTA with ventral and dorsal projections small and without deep excavation ( Fig. 39 View FIGURE 39 b).................. N. rothi View in CoL sp. nov.
9(7) Fulcrum present ( Fig. 2 View FIGURE 2 a)...................................................... N. annamae (Gertsch & Davis)
- Fulcrum absent ( Figs. 26 View FIGURE 26 a, 30d)........................................................................ 10
10(9) Tegular median process with two projections ( Figs. 4 View FIGURE 4 a, 22a, 34d, 38a)......................................... 11
- Tegular median process with one projection ( Figs. 19 View FIGURE 19 a, 26a, 33d, 37a)......................................... 17
11(10) Mesal and ectal projections of tegular median process adjacent ( Figs. 30 View FIGURE 30 d, 31a, 34d).............................. 12
- Mesal and ectal projections of tegular median process separated ( Figs. 4 View FIGURE 4 a, 16l, 22a, 38a)........................... 14
12(10) Mesal and ectal projections of tegular median process of similar length ( Fig. 31 View FIGURE 31 a)............... N. mexiquensis View in CoL sp. nov.
- Mesal projection of tegular median process longer than ectal projection ( Figs. 30 View FIGURE 30 d, 34d)........................... 13
13(12) Dorsal and ventral projections of RTA clearly differentiated ( Fig. 30 View FIGURE 30 e)........................... N. leonensis View in CoL sp. nov.
- Dorsal and ventral projections of RTA not clearly differentiated, represented by four small projections ( Fig. 34 View FIGURE 34 e)................................................................................................ N. poncei View in CoL sp. nov.
14(11) Tegular median process exceeding the length of conductor ( Fig. 38 View FIGURE 38 a)............................... N. punta View in CoL sp. nov.
- Tegular median process not exceeding the length of conductor ( Figs. 3 View FIGURE 3 c, 16l, 21b)................................ 15
15(14) Mesal projection of tegular median process with several teeth ( Fig. 21 View FIGURE 21 b, c)......................... N. dentata View in CoL sp. nov.
- Mesal projection of tegular median process without teeth.................................................... 16
16(15) RTA with dorsal projection larger than ventral projection ( Fig. 3 View FIGURE 3 d).................... N. approximata (Gertsch & Ivie)
- RTA with dorsal and ventral projections of similar size ( Fig. 16 View FIGURE 16 m)............................ N. bosencheve View in CoL sp. nov.
17(10) Embolus straight ( Figs. 19 View FIGURE 19 a, 20d)...................................................................... 18
- Embolus curved ( Figs. 10 View FIGURE 10 a, 26a, 33d)................................................................... 19
18(17) Ventral projection of RTA with a small denticle ( Fig. 19 View FIGURE 19 b)........................................ N. clara View in CoL sp. nov.
- Ventral projection of RTA without a small denticle ( Fig. 20 View FIGURE 20 e)..................................... N. creel View in CoL sp. nov.
19(17) Ventral projection of RTA with a spine-shaped subprocess ( Fig. 43 View FIGURE 43 e)............................. N. valdezi View in CoL sp. nov.
- Ventral projection of RTA without a spine-shaped subprocess ( Figs. 11 View FIGURE 11 b, 33e)................................... 20
20(19) Ventral projection of conductor notched ( Figs. 11 View FIGURE 11 b, 37a)..................................................... 21
- Ventral projection of conductor not notched ( Fig. 33 View FIGURE 33 d)...................................................... 22
21(20) Dorsal projection of RTA cupped ( Fig. 11 View FIGURE 11 b) and much larger than ventral projection ( Fig. 10 View FIGURE 10 c).............................................................................................. N. intermedia (Chamberlin & Gertsch) View in CoL
- Dorsal projection of RTA pointed and slightly larger than ventral projection ( Fig. 37 View FIGURE 37 b)................. N. puebla View in CoL sp. nov.
22(20) Dorsal and ventral projections of RTA of similar size ( Fig. 26 View FIGURE 26 b).................................. N. garnica View in CoL sp. nov.
- Dorsal projection of RTA larger than ventral projection ( Figs. 25 View FIGURE 25 b, 33e)........................................ 23
23(22) Ectal border of tegular median process undulate ( Fig. 25 View FIGURE 25 a)...................................... N. franckei View in CoL sp. nov.
- Ectal border of tegular median process slightly rounded ( Fig. 33 View FIGURE 33 d)................................. N. perote View in CoL sp. nov.
24(3) Conductor with dorsal projection larger than ventral projection; tegular median process with two short and pointed tips ( Fig. 17 View FIGURE 17 a).............................................................................. N. chamberlini View in CoL sp. nov.
- Conductor with dorsal and ventral projections of similar size or ventral projection larger than dorsal projection ( Figs. 18 View FIGURE 18 a); teg- ular median process otherwise ( Figs. 15 View FIGURE 15 a, 42a)........................................................... 25
25(24) Tegular median process hook-shaped ( Fig. 27 View FIGURE 27 a).............................................. N. gibarrai View in CoL sp. nov.
- Tegular median process otherwise ( Figs. 13 View FIGURE 13 a, 32a)......................................................... 26
26(25) Tegular median process with tip strongly sclerotized ( Figs. 15 View FIGURE 15 a, 32a)........................................... 27
- Tegular median process with tip as sclerotized as tegulum ( Figs. 13 View FIGURE 13 a, 18a, 42a)................................... 28
27(26) Tegular median process separated from embolus by its length ( Fig. 15 View FIGURE 15 a)............ N. simplex (F.O. Pickard-Cambridge) View in CoL
- Tegular median process separated from embolus by less than half its length ( Fig. 32 View FIGURE 32 a)................ N. oaxaca View in CoL sp. nov.
28(26) RTA with denticles ( Fig. 13 View FIGURE 13 b).................................................. N. shlomitae (García-Villafuerte) View in CoL
- RTA without denticles ( Figs. 18 View FIGURE 18 b, 42b).................................................................. 29
29(28) Tegulum rounded in lateral view; RTA with inconspicuous excavation ( Fig. 18 View FIGURE 18 b)................... N. cintalapa View in CoL sp. nov.
- Tegulum flat in lateral view; RTA with conspicuous excavation ( Fig. 42 View FIGURE 42 b)........................... N. tacana View in CoL sp. nov.
30(1) Copulatory openings adjacent, visible in posterior view ( Figs. 7 View FIGURE 7 e, 12e)......................................... 31
- Copulatory openings separated, usually visible in ventral view ( Figs. 2 View FIGURE 2 c, 5a, 16i, 40d, 44a)......................... 32
31(30) Copulatory openings as long as wide ( Fig. 7 View FIGURE 7 e)...................................... N. calavera Chamberlin & Ivie View in CoL
- Copulatory openings wider than long ( Fig. 12 View FIGURE 12 e)................................................. N. lutzi (Gertsch)
32(30) Lateral margins of atrium forming a spiral that surrounds the copulatory openings ( Figs. 17 View FIGURE 17 c, 34a, 35d)............... 33
- Lateral margins of atrium not forming a spiral ( Figs. 16 View FIGURE 16 a, 19c, 27c, 28a, 32c).................................... 35
33(32) Primary spermathecae strongly curved with anterior and posterior parts almost touching ( Fig. 35 View FIGURE 35 e)....... N. prieta View in CoL sp. nov.
- Primary spermathecae not strongly curved ( Fig. 34 View FIGURE 34 b)....................................................... 34
34(33) Primary spermathecae three times longer than wide and separated by their width ( Fig. 17 View FIGURE 17 d)........ N. chamberlini View in CoL sp. nov.
- Primary spermathecae two times longer than wide and separated by less than their width ( Fig. 34 View FIGURE 34 b)....... N. plata View in CoL sp. nov.
35(32) Copulatory openings in median position of atrium ( Figs. 26 View FIGURE 26 c, 39c)............................................. 36
- Copulatory openings in lateral position of atrium ( Figs. 16 View FIGURE 16 a, 24a, 30f).......................................... 37
36(35) Atrium with spurs ( Fig. 39 View FIGURE 39 c)................................................................ N. rothi View in CoL sp. nov.
- Atrium without spurs ( Fig. 26 View FIGURE 26 c)........................................................... N. garnica View in CoL sp. nov.
37(35) Primary spermathecae strongly curved and four times longer than wide ( Figs. 36 View FIGURE 36 a, 37d)................ N. puebla View in CoL sp. nov.
- Primary spermathecae less curved or straight and three times or less longer than wide ( Figs. 23 View FIGURE 23 a, 33g, 44e)............ 38
38(37) Atrium hexagonal-shaped ( Fig. 40 View FIGURE 40 d)....................................................... N. sinaloa View in CoL sp. nov.
- Atrium not hexagonal-shaped ( Figs. 5 View FIGURE 5 d, 16f, 19c, 31c)...................................................... 39
39(38) Distance between anterior and posterior margins of atrium as long or longer than half the plate length in posterior view ( Figs. 5 View FIGURE 5 f, 11e, 16c, 30h)....................................................................................40
- Distance between anterior and posterior margins of atrium shorter than half the plate length in posterior view ( Figs. 4 View FIGURE 4 e, 16h, 20h, 32e)........................................................................................... 43
40(39) Atrium with anterior margin strongly sclerotized; spurs adjacent ( Figs. 16 View FIGURE 16 a, 30f).................................. 41
- Atrium with anterior margin not sclerotized; spurs separated ( Figs. 4 View FIGURE 4 c, 13c, 27c, 34f, 40a).......................... 42
41(40) Spurs long ( Fig. 16 View FIGURE 16 a); primary spermathecae with a dorsal transverse mark ( Fig. 16 View FIGURE 16 b)................. N. ajusco View in CoL sp. nov.
- Spurs short ( Fig. 30 View FIGURE 30 f); primary spermathecae without a dorsal transverse mark ( Fig. 30 View FIGURE 30 g)............. N. leonensis View in CoL sp. nov.
42(40) Secondary spermathecae bifurcated; primary spermathecae adjacent ( Fig. 5 View FIGURE 5 e)............... N. bipunctata Roth View in CoL stat. rev.
- Secondary spermathecae not bifurcated ( Fig. 10 View FIGURE 10 f); primary spermathecae separated ( Fig. 11 View FIGURE 11 d)................................................................................................ N. intermedia (Chamberlin & Gertsch) View in CoL
43(39) Atrium with posterior margin thick ( Figs. 13 View FIGURE 13 c, 15c, 28a, 42c, 43a)............................................. 44
- Atrium with posterior margin thin ( Figs. 7 View FIGURE 7 f, 16i, 24d, 27c, 35a)............................................... 49
44(43) Atrium with posterior margin divided in two or three large and rounded parts ( Figs. 18 View FIGURE 18 c, 42c, 43a)................... 45
- Atrium with posterior margin more or less straight ( Figs. 13 View FIGURE 13 c, 15c, 28a)........................................ 47
45(44) Median plate wider towards ventral margin in posterior view ( Fig. 43 View FIGURE 43 c)............................. N. triunfo View in CoL sp. nov.
- Median plate uniformly wide in posterior view ( Figs. 18 View FIGURE 18 e, 42e)............................................... 46
46(45) Secondary spermathecae directed anteriorly ( Fig. 18 View FIGURE 18 d); spurs near to lateral margin of atrium ( Fig. 18 View FIGURE 18 c). N. cintalapa View in CoL sp. nov.
- Secondary spermathecae directed dorsally ( Fig. 42 View FIGURE 42 d); spurs in median position of atrium ( Fig. 42 View FIGURE 42 c)....... N. tacana View in CoL sp. nov.
47(44) Anterior margin of atrium with lateral lobes (ectal to spurs) ( Fig. 15 View FIGURE 15 c).............. N. simplex (F.O. Pickard-Cambridge) View in CoL
- Anterior margin of atrium without lateral lobes ( Fig. 13 View FIGURE 13 c).................................................... 48
48(47) Secondary spermathecae thick ( Fig. 28 View FIGURE 28 b); spurs separated by more than three times their basal width ( Fig. 28 View FIGURE 28 a)................................................................................................... N. irazu View in CoL sp. nov.
- Secondary spermathecae thin ( Fig. 13 View FIGURE 13 d); spurs separated by twice their basal width ( Fig. 13 View FIGURE 13 c).................................................................................................... N. shlomitae (García-Villafuerte) View in CoL
49(43) Copulatory ducts curved in longitudinal direction ( Fig. 27 View FIGURE 27 d).................................... N. gibarrai View in CoL sp. nov.
- Copulatory ducts straight or curved in lateral direction ( Figs. 2 View FIGURE 2 d, 5b, 33b, 44e)................................... 50
50(49) Atrium with a pair of deep pits ( Fig. 24 View FIGURE 24 d)................................................... N. durango View in CoL sp. nov.
- Atrium without a pair of deep pits ( Figs. 16 View FIGURE 16 f, 20a, 24a, 40a).................................................. 51
51(50) Primary spermathecae separated by more than their width ( Fig. 30 View FIGURE 30 b)............................. N. jiquilpan View in CoL sp. nov.
- Primary spermathecae adjacent or separated by less than their width ( Figs. 16 View FIGURE 16 j, 23a, 25d, 33g, 43g)................... 52
52(51) Atrium strongly procurved (copulatory openings anterior to spurs) ( Figs. 20 View FIGURE 20 a, 22c, 34f, 38c, 43f).................... 53
- Atrium straight or slightly procurved (copulatory openings on the same level or posterior to spurs) ( Figs. 2 View FIGURE 2 c, 24a, 32c, 44d)..................................................................................................... 58
53(52) Atrium with anterior margin strongly sclerotized ( Fig. 22 View FIGURE 22 c)..................................... N. dentata View in CoL sp. nov.
- Atrium with anterior margin not sclerotized ( Figs. 16 View FIGURE 16 f, 20a, 34f, 38c, 43f)....................................... 54
54(53) Secondary spermathecae longer than copulatory ducts width ( Fig. 43 View FIGURE 43 g)............................ N. valdezi View in CoL sp. nov.
- Secondary spermathecae otherwise ( Figs. 34 View FIGURE 34 g, 38d)........................................................ 55
55(54) Secondary spermathecae thick (almost as wide as copulatory ducts) ( Fig. 20 View FIGURE 20 b)................... N. comaltepec View in CoL sp. nov.
- Secondary spermathecae short and thin ( Figs. 16 View FIGURE 16 g, 34g, 38d)................................................. 56
56(55) Primary spermathecae strongly curved and without a transverse dorsal mark ( Fig. 38 View FIGURE 38 d)................. N. punta View in CoL sp. nov.
- Primary spermathecae less curved and with a transverse dorsal mark ( Figs. 16 View FIGURE 16 g, 34g).............................. 57
57(56) Spurs large, their basal width occupying almost half the atrium width ( Fig. 34 View FIGURE 34 f)...................... N. poncei View in CoL sp. nov.
- Spurs small, their basal width occupying less than half the atrium width ( Fig. 16 View FIGURE 16 f)................... N. alvarezi View in CoL sp. nov.
58(52) Atrium with lateral hoods ( Fig. 35 View FIGURE 35 a)........................................................ N. popoca View in CoL sp. nov.
- Atrium without lateral hoods ( Figs. 7 View FIGURE 7 f, 20f, 31c, 44a)....................................................... 59
59(58) Atrium with anterior or lateral sclerotized tips ( Figs. 2 View FIGURE 2 c, 5a, 31c, 33f).......................................... 60
- Atrium without anterior or lateral sclerotized tips ( Figs. 4 View FIGURE 4 c, 17f, 25c, 33a)....................................... 64
60(59) Atrium with anterior sclerotized tips ( Figs. 5 View FIGURE 5 a, 32c, 33f)..................................................... 61
- Atrium with lateral sclerotized tips ( Figs. 2 View FIGURE 2 c, 31c).......................................................... 63
61(60) Primary spermathecae curved ( Fig. 33 View FIGURE 33 g)..................................................... N. perote View in CoL sp. nov.
- Primary spermathecae straight ( Figs. 5 View FIGURE 5 b, 32d)............................................................. 62
62(61) Atrium with posterior margin protruded ( Fig. 5 View FIGURE 5 a)............................. N. attenuata (F.O. Pickard-Cambridge)
- Atrium with posterior margin not protruded ( Fig. 32 View FIGURE 32 c)......................................... N. oaxaca View in CoL sp. nov.
63(60) Median plate wider towards ventral margin in posterior view ( Fig. 31 View FIGURE 31 e); secondary spermathecae conspicuous ( Fig. 31 View FIGURE 31 d)..................................................................................... N. mexiquensis View in CoL sp. nov.
- Median plate uniformly wide in posterior view ( Fig. 2 View FIGURE 2 e); secondary spermathecae small ( Fig. 2 View FIGURE 2 d)................................................................................................... N. annamae (Gertsch & Davis)
64(59) Spurs directed mesally ( Figs. 4 View FIGURE 4 c, 19c, 25c, 44d)............................................................ 65
- Spurs directed posteriorly ( Figs. 20 View FIGURE 20 f, 24a, 33a, 44a)........................................................ 68
65(64) Atrium occupying more than half the plate width ( Figs. 19 View FIGURE 19 c, 44d)............................................. 66
- Atrium occupying half the plate width ( Figs. 4 View FIGURE 4 c, 25c)....................................................... 67
66(65) Spurs anterior to copulatory openings ( Fig. 44 View FIGURE 44 d)............................................. N. volcanes View in CoL sp. nov.
- Spurs covering copulatory openings ( Fig. 19 View FIGURE 19 c)................................................. N. clara View in CoL sp. nov.
67(65) Atrium length half the plate length ( Fig. 4 View FIGURE 4 c)....................................... N. approximata (Gertsch & Ivie)
- Atrium length less than half the plate length ( Fig. 25 View FIGURE 25 c)......................................... N. franckei View in CoL sp. nov.
68(64) Atrium with a central yellow protuberance ( Fig. 17 View FIGURE 17 h)....................................... N. cieneguilla View in CoL sp. nov.
- Atrium without a central yellow protuberance............................................................. 69
69(68) Spurs separated by their basal width ( Figs. 16 View FIGURE 16 i, 44a)........................................................ 70
- Spurs separated by more their basal width ( Figs. 7 View FIGURE 7 f, 20f, 24a, 33a, 40a)......................................... 71
70(69) Spurs mesal to copulatory openings ( Fig. 44 View FIGURE 44 a)................................................ N. victoria View in CoL sp. nov.
- Spurs anterior to copulatory openings ( Fig. 16 View FIGURE 16 i)............................................... N. atzimbo View in CoL sp. nov.
71(69) Spurs two times longer than wide ( Fig. 33 View FIGURE 33 a)................................................ N. paricutin View in CoL sp. nov.
- Spurs as long as wide ( Figs. 7 View FIGURE 7 f, 20f, 24a, 40a)............................................................. 72
72(71) Posterior margin of atrium protruding ventrally ( Fig. 40 View FIGURE 40 a)..................................... N. saltoensis View in CoL sp. nov.
- Posterior margin of atrium not protruding ( Fig. 7 View FIGURE 7 f)......................................................... 73
73(72) Lateral margins of atrium straight ( Fig. 20 View FIGURE 20 f); primary spermathecae oblique ( Fig. 20 View FIGURE 20 g).................. N. creel View in CoL sp. nov.
- Lateral margins otherwise; primary spermathecae straight ( Figs. 7 View FIGURE 7 g, 24b)....................................... 74
74(73) Lateral margins of atrium curved ( Fig. 7 View FIGURE 7 f); posterior part of primary spermathecae wider than anterior part ( Fig. 7 View FIGURE 7 g)............................................................................... N. costata (F.O. Pickard-Cambridge)
- Lateral margins oblique ( Fig. 24 View FIGURE 24 a); primary spermathecae uniformly wide ( Fig. 24 View FIGURE 24 b)............... N. divisadero View in CoL sp. nov.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.