Bornmüller (1907: 784) The comparative study of Alkanna and Huynhia (Boraginaceae, Lithospermeae), with description of a new species in Iran Khalvati, Sina Ranjbar, Massoud Phytotaxa 2021 2021-11-09 524 4 216 234 BSQQ Boissier (1846: 32) Boissier 32 1846 [164,575,158,185] Magnoliopsida Boraginaceae Alkanna Plantae Boraginales 10 226 Tracheophyta species frigida   Type:—  IRAN. Kohgiluyehand Boyer-Ahmad: Dena Mt.[In argillosis humidis alpis Kuh-Daëna post nivem deliquescentem],  8 July 1842,  Kotschy573( holotype G! [G-00176782]; isotype BM! [ BM013839850], C! [ C10008670], E! [ E00284793], FI! [ FI009503], GOET! [ GOET005793], H! [ H1613716], K! [K-001085737], MO! [MO-503924], P! [ P00606828], US! [ US01050205], W! [W-0051322], WAG! [ WAG0058072]). Fig. 9.   =  Alkanna frigidavariety pallida  Bornmüller (1907: 784).   Type:—  IRAN. Tehran: Tochal Mt.[ M.  Elburs, in jugo Lädd alpium Totschal],  3000 m,  4 June 1902,  Bornmüller7703( holotype JE! [JE-00012012]).   Description:—Perennial, root rather thick, stem 3–10 (–15) cm long, multi-branched, branched from base, densely patent-pilose. Basal leaves oblong to ovate, 30–100 × 5–12 mm, acute to sub-acute, tapering into the petiole, with entire, flat, or sometimes slightly undulate margins, white densely tuberculate setose. Cauline leaves oblong to linear, 10–20 × 3–5 mm, acute or subacute, sessile, attenuate at base, with an evident midrib, veinless, densely white setose. Inflorescence a compact terminal head; pedicels short, leafy, peduncle 1–2 cmlong; bracts oblong to ovate, 5–10 × 2–5 mm, densely to sparsely tuberculate setiform and pilose. Pedicels erect, 1–2 mmlong at flowering, recurved 2–3 mmlong in fruit and pilose. Calyx 3–5 mmlong in flowering time, densely hirsute, and eglandular hairy. Corolla sulphur yellow, occasionally with purple spots, 10–12 (–15) mm long, glabrous outside, limb 3–5 mmin diam., tube 2–3 mmwide, and 6–10 mmlong. Nutlets 5–6 × 3–4 mm, densely tuberculate; beak horizontal to strongly curved.  Additional specimens examined:—  IRAN.  Alborz: Kandavan, 36º 5’ N, 51º 11’ E, s.d.,  Wendelbo& Shirdelpour11650 ( TARI);   Chaharmahaland   Bakhtiari: Lordegan, between Monj& Char-Tagh, Tang-e Zendan, 31º 30’ N, 50º 49’ E, s.d.,  Mozaffarian59924( TARI);   Isfahan: Semirom, Khafr, Dena Mt., 30º 58’ N, 51º 30’ E, s.d., Riazi5982 ( TARI);   Ghazvin: Alamut, 36º 31’ N, 50º 28’ E, s.d.,  Mirfakhrayi1382( TARI);   Semnan: 35 kmnorth of Damghan, Touyeh, 36º 2’ N, 53º 51’ E, s.d.,  Wendelbo& Assadi29488( TARI);   Tehran: Damavand, Valiran village, 35º 40’ N, 52º 5’ E, s.d.,  Mozaffarian32345( TARI). 3400134310 FI009503 1842-07-08 G, BM, C, E, FI, GOET, H, K, MO, P, W, WAG Iran Kotschy Kohgiluyeh Dena Mt. 10 226 BM013839850, C10008670, E00284793, GOET005793, H1613716, P00606828, WAG0058072 1 holotype 3400134330 [219,513,378,400] M Iran Tochal Mt. 10 226 1 Tehran 3400134335 1902-06-04 JE Iran 3000 Bornmuller 10 226 1 Alborz holotype 3400134344 TARI Wendelbo & Shirdelpour Iran 36.083332 Kandavan 1190 51.183334 11 227 1 Alborz 3400134317 [308,526,512,539] Iran Chaharmahal 11 227 1 Alborz 3400134362 TARI Mozaffarian Iran Lordegan 31.5 Char-Tagh 1216 50.816666 Monj 11 227 1 Bakhtiari 3400134323 TARI Riazi Iran Semirom 30.966667 Dena Mt. 1218 51.5 Khafr 11 227 1 Esfahan 3400134324 [233,1034,583,610] TARI Mirfakhrayi Iran 36.516666 Alamut 1186 50.466667 11 227 1 Ghazvin 3400134336 TARI Wendelbo & Assadi Iran 36.033333 Touyeh 1190 53.85 Damghan 11 227 1 Semnan 3400134329 TARI Mozaffarian Iran 35.666668 Valiran village 1191 52.083332 Damavand 11 227 1 Tehran