New and additional distribution data of the genus Acupalpus LATREILLE 1829 from the Middle East and adjacent areas (Coleoptera, Carabidae, Harpalini, Stenolophina) Jaeger, B. Linzer biologische Beiträge 2011 2011-07-25 43 1 763 775 9VGQ (SCHAUM 1960) SCHAUM 1960 [178,707,1458,1480] Insecta Carabidae Acupalpus Animalia Coleoptera 4 767 Arthropoda species maculatus Acupalpus  D i s t r i b u t i o n i n M i d d l e E a s t: So far recorded from Cyprus( JAEGER 1999: 960, AUSTIN et al. 2008: 68, 69), Turkey( JAEGER 1992: 225, 1999: 960), Syria( JAEGER 1992: 225), Jordan( JAEGER 1992: 225), Israel( BODENHEIMER 1932: 53, JAEGER 1992: 225, 1999: 960), Iran( JAEGER 1992: 225, 1999: 960) and Afghanistan( JEDLIČKA 1955: 191, JAEGER 1999: 960). The following new material, including first records for Lebanon, Pakistan, India, and for the Iranian province Kerman, and for the Turkish provinces Çanakkale, Elâziğ, Erzincan, Kars, Samsun and Uşak, has been examined. The finds from northwest India( Jammu and Kashmir) represent the most eastern records of  A. maculatusin the southern West Palaearctic. E x a m i n e d m a t e r i a l: C y p r u s: Ezousa - Fluss Ufer, bei Kourdaka, IV. 2004, Apfel (1 – cAPF).  T u r k e y: Adana: Adana-Balcali,  VIII. 1980, Kasap(1 – MNHUB);  Karatas,  50 kmS Adana, 36°33’27"N, 35°20’15"E,  V. 2002, Skale(1 – cSKA). – Antalya: Beldibip. Kemer,  VII. 1998, Bulirsch(1 – cJAE);  Göynükp. Kemer, Bey Dagl.,  50-500 m,  VII. 1998, Bulirsch(1 – cJAE);  Sucati-Riverbei Asagigökdere,  380 m, 37°32+30°46,  V. 2006, Marggi(4 – cMAR, cJAE). – Aydin: Milet, Menderes Delta,  V. 1991, Schönmann& Schillhammer(1 – NHMW). – Çanakkale: N Ezine, Sarmisaki Fluss,  V. 1991, Schönmann& Schillhammer(1 – NHMW). – Elâziğ: Elazig– Ergani,  V. 1966, Türkei-Expedition Naturhistorisches Museum Wien(6 – NHMW). – Erzincan: Refahiye E, Sakaltutan Geçidi W, Stausee Schlammufer  1936 m, N 39°52.682' E 39°01.705',  VIII. 2002, Schnitter (4 – cSCHN). – Erzurum: N Ikizdere,  V. 1989, Schönmann& Schillhammer(1 – NHMW). – Kars: Asagi Fluss, ETuzluca,  VI. 1989, Schönmann& Schillhammer(1 – NHMW). – Konya: Bozkir, Yalihüyük,  V. 1998, Nodmar (1 – cJEA). – Muğla: Köyceğiz Gölü,  10 m, 36°55’19"N, 28°42’14"E, flood debris sifted,  III. 2002, Wunderle(5 – MNHUB). – Samsun:  15 kmNW Bafra, Karaboğaz Gölü, beach debris, 41°40’49"N 35°48’16"E,  III. 2009, Assing (21 – MNHUB);  Carsamba,  V. 1989, Jäch(1 – NHMW). – Sivas: Tödürge Gölü, ZaraW, Salzpfannen, Uferzone,  1337 m, N 39°51.924' E 37°37.380', Schnitter& Staven(5 – cSCHN). – Uşak: 30 kmW Uşak,  V. 2002, Skoupý (1 – cSKO). L e b a n o n Bekaa: Ammiq, Reserved Area, Feuchtgebiet, 870 m, 33°43'42"N, 35°47'09"E, V. 2006, Frenzel & Weipert (15 – cWEIP, cFRE, cJAE, NME).  I s r a e l: Central District: Breikhat Ya'ar, N Tel Aviv,  11 m(wetland),  III. 2008, Wrase(1 – cWRA);  Ga'ash, S Netanya(winter pool, Eleocharis),  III. 2008, Wrase(4 – cWRA);  Netanya, Breikhat Dora Res.(wetland),  III. 2008, Wrase(4 – cWRA). - Haifa District: Carmel Ridge, Dalya/ Gared,  III. 1995, Sama(33 – SMNS, cJAE); coastal area  3 kmW Zikhron Ya'akov(edges of fish pond brackish water),  V. 2006, Wrase(22 – cWRA, cJAE);  Ein Afek Reserve, W Kiriat Motskin( Haifa) (loamy foreshore of brackish ponds),  III. 2008, Wrase(1 – cWRA). – North District: Upper Galilee, Ha Khula Valley, Ma'agar Einan lake,  73 m, 33°05.137'N 035°34.730'E(shore gravel, sandy/loamy soil),  V. 2006, Wrase(1 – cWRA);   Upper Galilee, N.shore of sea of Galilee-  200 m, Tamariscfloodplain forest, betw. Kfar Nakhum( Capernaum) and mouth of Jordanriver, III/  IV. 2008, Wrase (3 – cWRA). – Golan Hights: Bental Reservoir, WMerom Golan, ca.  1000 m, (stony wetland near shore),  VI. 2006, Wrase (5 – cWRA).  I r a n: Azabayian-e Gharbi:  18 kmSE Quarah Ziya'oddin( Aq River),  960 m, N 38°51'03" E 045°13'31",  VIII. 2008, Frisch& Serri(1 – MNHUB). – Gilan: 20 kmW Bandar Anzali,  VI. 2009, Skoupý(21 – cSKO, cJAE);  Asalem,  VI. 2009, Skoupý(7 – cSKO, cJAE);  Espeyli,  2000 m,  VI. 2009, Skoupy(1 – cSKO);  Rudbar,  VI. 2009, Skoupý(19 – cSKO, cJAE). – Golestan: S Gorgan,  12 kmS Nahar Khoran, Ziarat,  1200 m, N 36°40'52" E 054°27'54",  VI. 2006, Frisch& Serri(1 – MNHUB). – Kerman: Kerman, rivier 5 km Z Khabr, N 28°50'35.6"/ E 56°14'54.8",  III. 2007, Felix (1 – cFEL). – Mazandaran:  40 kmS Tavir,  660 m, meadow beside road, 35.25.12N 51.16.53E,  V. 2010, Frenzel(1 – cFRE);  Kalardasht: Rudbarak,  1790-1950 m, N 36°26'43" E 051°03'48",  VII. 2005, Frisch& Serri(1 – MNHUB);  Now Shahr, 36°36'17"N, 51°38'23"E,  VI. 2008, Frenzel(1 – cFRE);  Tonekabon: Dohezar,  400 m, N 36°40'29" E 050°49'40",  VII. 2005, Frisch& Serri(1 – MNHUB);  Alamdeh,  VI. 1974, Bernhauer(1 – cHEI);  Sari,  I. 1875, Kerim(1 – MCSNG). - Ostan- e Ardabil:  20 kmS Astara, 3816N, 4830E,  2 m,  VI. 2000, Hajdaj(1 – cWRA). A f g h a n i s t a n: Bala Murghab, 470 m, (23), VI. 1964, Jakeš (1 atypical female – MMB).  P a k i s t a n: Khyber Pakhtunkhwa Province( North-West Frontier Province): Kâgân-Tal, Naran,  2400-2700 m,  VII. 1981, Heinz(5 – MNHUB, cJAE);  Hazara, Kaghan,  2150 m,  VI. 1983, Besuchet& Löbl(2 – MHNG, cJAE);  Swat, Madyan,  1400 m,  V. 1983, Besuchet& Löbl(1 – MHNG);:  Swat, s/ Utrot,  2500-2600 m,  V. 1983, Besuchet& Löbl(1 – MHNG).  I n d i a: Jammu and Kashmir: Srinigar,  VI. 1984, Levey (2 – NMW);  Srinagar,  1600 m,  VI. 1976, Wittmer (1 – NHMB). It should be noted that the specimens from the Swat and Khagan Valley in northern Pakistan(see above) differ from typical members of  A. maculatus. The Pakistani specimens at a first glance remind one of  Acupalpus brunnipes, because five of them are uniformely blackish to dark blackish brown, having only the mouthparts, pronotal and elytral margins narrowly, and first elytral interval dark reddish brown. The remaining specimens have at least the outer elytral intervals vaguely lighter, and in one specimenindications of a small dark brown spot in front of the base of each elytron. In typical maculatusspecimens the pronotal margins are more widedly reddish yellow and the elytra are reddish yellow with an ±expanded dark central macula, which typically extends along intervals 5-6 to elytral base, so that a distinct oval reddish or yellowish patch remain at both sides of suture. Rarely the pronotum is almost black, and very rarely the elytra are completely dark and immaculate in single specimens occuring within populations of typically coloured specimens. In addition to these colour differences the eyes are somewhat less prominent; the pronotal basal impressions are more distinctly punctate; the microsculpture at apical half of the head is more isodiametric, instead of more tranversely stretched, and on pronotum and elytral intervals the meshes are generally somewhat coarser and more distinct. However, in single specimens of the Pakistani populations as well as in those from other places the mentioned differences are less clear. A comparison of the external shape of the median lobe of aedeagus and its internal sac revealed no difference between the Pakistani specimens and those from other areas. The population from northern Pakistanis here treated as an intraspecific geographical form without formal taxonomic rank. [178,836,559,579] 1980-08 MNHUB Kasap Adana-Balcali 5 768 1 Adana 1998-07 2002-05 1998-07 Skale & Beldibi & Kemer & Bulirsch 36.5575 Karatas 19 35.337498 5 768 1 Adana [418,1082,607,627] 1998-07 Bulirsch Goynuk 275 Bey Dagl. Kemer 5 768 1 Adana 1966-05 2009-03 1966-05 NHMW, MNHUB Marggi & Milet & Menderes Delta & Schonmann & Schillhammer & N Ezine & Refahiye E & Sakaltutan Gecidi W & N Ikizdere & Fluss, E & Wunderle Sucati-River 380 41.68028 Sarmisaki Fluss 19 35.804443 Asagigokdere 5 768 1 Aydin 1989-05 2002-05 1989-05 NHMW Jach & Schnitter & Staven Carsamba 1337 39.8654 Salzpfannen 1 37.623 Todurge Golu 5 768 1 Sivas [178,1082,1007,1027] 2008-03 Breikhat Ya'ar & N Tel Aviv 11 Central District Wrase 5 768 1 [223,870,1031,1051] 2008-03 Ga'ash & S Netanya & Eleocharis Wrase 5 768 1 1995-03 2008-03 1995-03 SMNS Netanya & Breikhat Dora Res. & Carmel Ridge & Dalya & Gared Sama Wrase Haifa District 5 768 1 2006-05 Wrase 5 768 1 2006-05 2008-03 2006-05 Ein Afek Reserve, W & Kiriat Motskin & Haifa Wrase 73 33.085617 Wrase 1 35.578835 Ma'agar Einan lake 5 768 1 2006-06 2008-04 2006-06 Upper Galilee, N. & Galilee & Tamarisc & Kfar Nakhum & Capernaum & Bental Reservoir, W Jordan 200 Upper Galilee 5 768 1 2008-08 2009-06 2008-08 MNHUB Frisch & Serri & Skoupy Azabayian-e Gharbi 960 38.85083 Bandar Anzali 19 45.225277 Aq River 5 768 1 Gilan [482,890,1327,1347] 2009-06 Skoupy Asalem 5 768 1 Gilan 2009-06 Skoupy 2000 Espeyli 5 768 1 Gilan 2006-06 2010-05 2006-06 MNHUB S Gorgan & Frisch & Serri & Z Khabr & Frenzel 1200 28.843224 Skoupy 1 56.248554 Rudbar 5 768 1 Golestan 2005-07 MNHUB Frisch & Serri 1870 36.44528 Rudbarak 19 51.06333 Kalardasht 5 768 1 Golestan 2008-06 Frenzel 36.60472 Now Shahr 19 51.639725 5 768 1 Golestan 2005-07 MNHUB Frisch & Serri 400 36.674725 Dohezar 19 50.827778 Tonekabon 5 768 1 Golestan [324,693,1519,1539] 1974-06 Bernhauer Alamdeh 5 768 1 Golestan 1875-01 2000-06 1875-01 MCSNG Kerim & Hajdaj 2 Sari 5 768 1 Ardabil 1981-07 MNHUB Heinz 2550 Naran Kagan-Tal 6 769 1 Khyber Pakhtunkhwa 1983-06 MHNG Besuchet & Lobl 2150 Kaghan Hazara 6 769 1 Khyber Pakhtunkhwa 1983-05 MHNG Besuchet & Lobl 1400 Madyan Swat 6 769 1 Khyber Pakhtunkhwa [320,922,288,308] 1983-05 MHNG Besuchet & Lobl 2550 Utrot Swat 6 769 1 Khyber Pakhtunkhwa [178,823,320,340] 1984-06 NMW Srinigar 6 769 1 Jammu and Kashmir 1976-06 NHMB 1600 Srinagar 6 769 1 Jammu and Kashmir