Conostegia lasiopoda Conostegia trianaei Conostegia sororia A Monograph of Conostegia (Melastomataceae, Miconieae) Kriebel, Ricardo PhytoKeys 2016 2016-07-20 67 1 326 F21D4AE0-4FE7-9BEF-DDAA-5EA2137307C0 Benth. Benth. Magnoliopsida Melastomataceae Conostegia CoL Plantae Conostegia lasiopoda Myrtales 129 130 Tracheophyta species lasiopoda   Conostegia lasiopodaBenth., Bot. Voy. Sulphur 96. 1844. Type: Costa Rica. Puntarenas: Cocos Island, no date, Barclay s.n.(K!).  Conostegia trianaeiCogn., DC. Monog. Phan. 7: 702. 1891. Type: Colombia. Choco: 1851-1857, J. Triana 3943 (holotype: BR!, isotypes: BM!, NY!, K!, US!, W).  Conostegia sororiaStandl., Field Mus. Nat. Hist., Bot. sere 22: 161. 1940. Type: Panama. Darien: Cana-Cuasi trail, Rio Cuasi (camp I), Chepigna, 250 m, 7 March 1940, M. and R. Terry 1414 (holotype: F!, isotypes: A!, MO!).  Description. Shrubs to small trees 1.5-6 m tall with stems that are first tetragonal then terete and setose with appressed single bristles 1-2 mm long and many minute sessile stellate hairs; nodal line present yet slight. Leaves at a node equal to subequal in size. Petioles 0.7-3.6 cm long, densely hirsute adaxially. Leaves 8.3-30 x3.6-11.7 cm, 3-5 nerved, ovate to obovate, cuneate to obtuse at the base, the apex acute to rounded and attenuate to abruptly acuminate,the margins entire to undulate-dentate, the adaxial surface glabrous, slightly concave and with deeply impressed tertiary venation, abaxially covered only with a layer of these tiny sessile stellae. Inflorescence terminal, 5-19.1 cm long, accessory branches absent or present, the branches subtended by deciduous, setose linear bracteoles 2-20 mm long, the clusters of flowers subtended by puberulent foliose, persistent to deciduous bracteoles 2-7 mm long. Pedicels of 0.5-2.0 mm. Flowers (5-)6(-8) merous, calyptrate. Floral buds 4-9 x2-5 mm, ovate to oblong-pyriform, the base rounded, the apex acuminate, constricted below the middle, the upper and lower portions undifferentiated, the hypanthium 3.5-4 x3.25-3.5 mm, pubescent. Petals 7-8 x7-8 mm, translucent white, obtriangular, spreading, glabrous, and slightly asymmetrical apically. Stamens 17-25, 4-6 mm, slightly zygomorphic apparently from the movement of the style, the filament 2.5-3 mm, white, anthers 1.5-2 x0.5-1 mm, linear-oblong, straight or recurved, light yellow., the pore terminal, less than 0.1 mm. Ovary 6-9 locular, inferior, apically glabrous and forming a low collar around the style. Style 5-8 mm, straight for most of its length and bending gently near the apex, vertical distance from the anther to the stigma 2-3 mm, the stigma capitate, 1-1.25 mm wide. Berry 8-10 x8-10 mm, light purple to purple-black. Seeds 0.3-0.5 mm long, ovoid, the testa smooth to roughened.   Figure 123.  Conostegia lasiopoda. ALeaf abaxial surface BInternode CLateral view of the flower DFruit EInflorescence branch showing bracteoles covering flower buds FLongitudinal section of a flower bud GFlower HLongitudinal section of a flower with petals removed IPetal JStamen Photographs A, Bby Reinaldo Aguilar and vouchered R. Aguilar 11265 C-Jvouchered from specimen R. Kriebel 5651.  Distribution (Fig. 124). Distributed in southeastern Nicaragua, wet forests of Costa Rica, somewhat restricted in Panama to mostly western Colombia, with a few specimens collected in northwestern Ecuador, also in Cocos Island, from sea level to 2100 m in elevation.   Figure 124.Distribution of  Conostegia lasiopoda.   Conostegia lasiopodais usually a quite distinctive species. The leaf veins are elevated and arise exactly at the base ("perfectly nerved"), the petioles are setose adaxially and the flowers have foliaceous bracts covering them. When these bracts fall, distinguishing   Conostegialasiopodafrom  Conostegia centronioidescan be difficult. The latter tends to be at least somewhat plinerved and lack the adaxial setose petioles.  Conostegia rubiginosais also similar but lacks the foliaceous bracts covering the floral bracts and the indument in flower bud is more evident. Also,  Conostegia rubiginosatends to have smaller coriaceous leaves. The stamens of  Conostegia lasiopodahave a conspicuous anther shoulder (Figs 49, 123) which has not been confirmed in  Conostegia centronioidesor  Conostegia rubiginosaand the style is exserted (Fig. 123) like the latter two taxa and as usual in section  Australis. Lastly, the calyptra has been observed to rupture into pieces at anthesis (pers. obs. and photographs by Reinaldo Aguilar).  Specimens examined.   NICARAGUA(fide Schnell). Zelaya: Salto La Oropendula, Rio Rama, Stevens 8962 (MO).    COSTA RICA. Alajuela: San Ramon, R. B. Manuel Alberto Brenes, Estacion  RioSan Lorenzo, Kriebel 902 (INB);    Limon:  Pococi, P.N.Braulio Carrillo, EstacionQuebrada Gonzalez, Kriebel 203 (INB); Matina, P. N. Barbilla, Cuenca del Matina, SectorColonia Puriscalena, Sendero Cerro Azul hasta  RioSurubres, Mora 1162 (INB, MO, NY).   Puntarenas: R.F. Golfo Dulce,  Serraniasde Golfito, Estacion  RioBonitopor la Filaque va a Cerro La Gamba, Aguilar and Albertin5115 (NY); Distrito, Sierpe, Reserva Forestal Golfo Dulce, Mogos 1 kmantes de llegar a la entrada del Porvenir a 16 km. de Chacarita, Aguilar 11265 (NY); E.B. Las Cruces, downhill from the greenhouses along the trail to the  rioJava, Boyle et al. 6435 (NY); along stream leading into Chatham Bay, Cocos Island, Fournier 303 (CR, NY);  BahiaWaferaguas arriba del  RioGenio, Gonzalez1200 (INB, NY); Reserva Forestal innominada 2 K al norte de la entrada de Chacaritapor la carretera interamericana, Gonzalez3594 (INB, NY);  EstacionLa Gamba, Kriebel 1078 (INB).    PANAMA.  Panama: Sendero de Interpretacion, 1 kmal este del Campamento de los guardabosques de INRENARE, Correa and Montenegro 11122 (NY).   Veraguas: Parque Nacional Santa Fe, aproximadamente de 3 a 6 kmpasando la Escuela Agricola Alto de Piedra, Kriebel and Burke 5722 (NY, PMA).    COLOMBIA. Antioquia: P.N. Natural Las Orquideas, Vereda Venados Abajo, Pedraza et al. 2323 (NY).    Choco:  RioSan Juan, cercanias de Palestina, Cuatrecasas 16938 (NY); area of Baudo, on right bank of river Baudoabout 12.5 kmupstream of estuary, near estuary of Quebrada Carpio, 1 kmupstream of camp site, Quebrada Anguerado, Fuchs and Zanella 21770 (MO, NY); At RioIro on Hwy Sfrom Istmina, Juncosa 2484 (MO, NY).   El Valle:  RioCalima( regiondel Choco), entre Malaguita y Palestina, Cuatrecasas21331 (NY).   ECUADOR. Esmeraldas: San Lorenzo cantonRicuarte, Reserva Indigena Awa, Tipaz et al. 2107 (MO, NY). 2016-01-01 2016-12-31 2016-01-01 San Ramon, R. B COSTA RICA Rio San Lorenzo 1 Alajuela 2016-01-01 2016-12-31 2016-01-01 Pococi, P. N., Matina, P. N Costa Rica Rio Surubres Cuenca del Matina 1 Limon 2016-01-01 2016-12-31 2016-01-01 R. F. Golfo Dulce, la Fila, de Chacarita, E. B. Las Cruces Costa Rica Serranias de Golfito Cerro La Gamba Rio Bonito 1 Puntarenas 2016-01-01 2016-12-31 2016-01-01 de Interpretacion PANAMA Sendero de Interpretacion 1 Panama 2016-01-01 2016-12-31 2016-01-01 la Escuela Agricola Alto de Piedra Panama Escuela Agricola Alto de Piedra 1 Veraguas 2016-01-01 2016-12-31 2016-01-01 P. N. Natural Las Orquideas COLOMBIA Antioquia 1 Antioquia 2016-01-01 2016-12-31 2016-01-01 de Palestina, Hwy S Colombia Rio San Juan At Rio Iro on Hwy river Baudo 1 Choco 2016-01-01 2016-12-31 2016-01-01 Colombia Rio Calima 1 El Valle