Ewart, 1988 : 182 A revision of the Ewartia oldfieldi (Distant) species complex (Hemiptera: Cicadidae: Cicadettinae) with five new species from eastern and northern Australia Popple, Lindsay W. Zootaxa 2017 4263 3 401 449 8JSHT [151,336,1077,1103] Insecta Cicadidae Ewartia Animalia Hemiptera 11 412 Arthropoda species roberti sp. nov.     Cicadetta oldfieldi(Distant):  Ewart, 1988: 182–183, 191, 193, 196–197, Fig. 9J–M, Plate 2C.    Types.  Holotype: ♂[typed] ‘ 28°59’S 151°28’E’/ ’ AUSTRALIAQLD’ / ‘ Glenlyon Dam, via’ / ‘ Texas, SEQ.’ / ’  11–12.Mar.2004. ’ / ‘ L. & W. Popple.’ / ‘231-0023’ / ‘QM reg. no. T207359’ ( QM);  Paratypes: QUEENSLAND: 1♂ Indooroopilly, Brisbane Q.,  28.ii.1990;  2♀ Stanthorpe,  11.ii.1990, A. I. Knight; 1 M Mackellar Ra. Rd, Bungabbeee SF, 15kmSE. Kyogle, NSW,  28.xii.1997, mv light, Watkinselev.  180m;  1♀QLD: Mt Woowoongaca 50mi. W. of Bundabergvia Dallarnil,  29–30.i.1972, H. Frauca;  1♂NW slope of Bluff Rg. nr BiggendenQLD,  10.i.1972, H. Frauca;  2♂QLD Biggenden, Bluff Range,  11.i.1971, H. Frauca, c.  1700ft;  1♂same data as previous,  12.xii.1972;  1♂same data as previous,  13.i.1977;  1♂QLD: Biggenden, Bluff Range, Southfhills,  9.xii.1972, H. Frauca;  2♂same data as previous,  19–20.xii.1972, H. Frauca;  2♂ 1♀QLD: Mt Walshsummit,  2200ft, Biggenden,  3.i.1972, H. Frauca;  1♂ CoastRanges, 11kmS. of Biggenden Q.,  4.xii.1976, H. Frauca;  1♀QLD: Spring Ckca 30kmS. of Bundaberg,  23.xii.1976, H. Frauca;  1♂ 27.33S 151.59E, Prince Henry Heights,  620m, Toowoomba Q.,  19.xii.1985, I. F. B. Common( all ANIC);  2♂ Qld: 25.552° Sx 151.469°E, Binjour Plateau, Redvale Rd,  21.xii.1997, C. J. Burwell, open forest,  340m;  1♂ 1♀SEQ: 25°42’S 151°26’ENipping Gully, Site6,  18– 19.xii.1998, G. Monteith, C. Gough& G. Maywald, 7532,  Cicadetta oldfieldidet. A. Ewart, 1997;  1♂ Toowoomba,  26.x.1908, Hamlyn-Harris,  Melampsalta oldfieldi; 1♀ Toowoomba, 1908, Hamlyn-Harris,  Melampsalta oldfieldi;  3♂ 2♀QLD: 26°26’ Sx 150°31’E, Barakula,  330m,  17–18.xii.2001, G.Monteith& S. Wright, M.V. light, 10311;  1♀QLD: 28°03’ Sx 153°07’E‘ Glen Witheren’ General,  26–29.xii.2001, G. Montieth, M. V. light, 10352;  1♂SEQ: 24°55’ 150°59’E, Hurdle Gully,  14.8kmWSW Monto, open forest,  460m,  20.xii.1997, C. J. Burwell, S. Evans; 1♂ 1F Gooroolba, S.E.Q., 1.xii.1952, W.W.Abell,  Melampsalta oldfieldi;  1♂N. of Samford,  xi.1990, C. J. Burwell,  Cicadetta oldfieldidet. A. Ewart1997 ( all QM);  1♂ Australia, Queensland, 4kmW. of Binjour, SEQ,  24.i.2003, L. & W. Popple, mv lamp, 231-0001;  2♂ 1♀ Australia Queensland, 6 kmwest of Thane,  11.xii.2001, Handcollected, L. W. Popple, 28°09’41”S 151°57’59”E, 231-0002 to 231-0004;  1♂ Australia Queensland, Willowvale, 5 km N. Warwick,  11.xii.2001, L. Popple, J. Moss, 231-0005;  1♂ Australia Queensland, 28.162S° 151.6287°E, Thanes Creekvia Warwick, L. Popple, J.Moss, 231-0004, Genitaliaprep. 231-01;  1♂ Dinmore, Ipswich, SEQ,  6.iii.1998, L. Popple,  Acaciasp., 231-0006;  1♂ Camp Hill, Brisbane, SEQ,  3.iii.1998, L. Popple,  Acaciasp., 231-0007;  1♂ Myall Park, 8km N. Glenmorgan, SEQ,  27–28.xii.2001, L. Popple, A. Strange, 231-0008;  1♂ Austin Rd, Crows Nest, SEQ, mv lamp,  26.xii.2001, L. Popple, A. Strange, 231-0009;  1♂ Highfields, via Toowoomba, SEQ,  26.xii.2001, L. Popple, A. Strange, 231-0010;  1♂ Lytton, SEQ,  10.xii.2000, L. Popple, S. Billington, 231-0011;  3♂ Carina Heights, SEQ,  7.xi.1998, L. Popple, 231-0014-231-0016;  1♀ Carina Heights, SEQ,  3.xii.1999, W. Popple, 231-0017;  1♂ Australia, Queensland, Carina Heights, Brisbane, SEQ,  7–8.i.2003, L. W. Popple, Recorded L. W.Popple, 231-0018;  1♂ Australia, Queensland, Carina Heights, Brisbane, SEQ,  i.2003, L. W. Popple, 231-0019;  1♂ Australia, Queensland, Carina Heights, Brisbane, SEQ,  21.xii.2002, R. J. Popple, 231-0020;  1♀ 27°18’S 152°28’E, Australia, Qld, Mount Glorious,  11.i.2004, M. Larsen, mv lamp, 231-0022;  1♂ Australia Queensland, 28°34’S 152°49’E, Afterlee Public School, Kyogle district, rural, L. Popple, R. MacSloy, 231–0024;  1♂ Australia Queensland, 25°22’42”S 151°07’46”E, Eidsvold, Burnett River,  28–29.xii.2004, L. W. Popple, 231-0025;  1♂ Australia Queensland, 28.162S° 151.6287°E, Thanes Creekvia Warwick,  28.i.2005, L.Poppleand R. MacSloy, 231-0027;  1♂ Australia Queensland, 27°24’47”S152°11’60”E, Helidon Hills, Seventeen Mile Road,  19.ii.2005, L. W. Popple, R. MacSloy, 231-0028;  1♂ Australia Queensland, 25°36’18”S 151°32’26”E, Mt Debatable(summit),  28.x.2005, L. W. Popple,  Acaciasp., 231-0029;  1♂ Australia Queensland, Southwood NP, via Moonie,  5– 10.xii.2005, L. Popple, A. Ewart, 27°50’05”S 150°06’47”E, 231-0030;  1♂ 3♀ Australia Queensland, Lake Broadwater, via Dalby,  14–15.xii.2005, L. W. Popple, mv lamp, 27°21’02”S 151°05’34”E, 231-0031 to 231-0035;  1♂ Australia Queensland, Metroplex, Murrarie, 27°26’58”S 153°05’54”E,  iii.2008,  Ac. fimbriata, L. W. Popple, 231-0036;  1♂ Australia Queensland, Burrandoan Rd, SWof Kingaroy,  11.xii.2007, 26° 26°35’54”S 151°44’58”E, 231-0037;  1♂ AustraliaQLD, Roys Rd, Beerwah( F.S.C.), 26°51’19”S 153°00’24”E,  10.xii.2007,  A. flavescens, L. Popple, R. MacSloy;  1♂ AustraliaQLD, Warwick— 5 kmS., parkland,  Acaciasp.,  22.i.2010, L. W. Popple, 28°13’31”S 152° 01’12”E, 231-0039 (all LWP);  1♀ Iron Bark, Crows Nest,  6.xi.1975, [ A. Ewart];  2♂ 1♀ Iron Bark, Crows Nest,  22.xi.1975, [ A. Ewart];  2♂ Perseverance Dam,  19.x.1975, [ A. Ewart];  1♀ Ironbark Qld, Crows Nest,  1.xi.1977[ A. Ewart];  1♂ L. Broadwater, Dalby, Qld,  12.ii.1984, [ A. Ewart];  3♂ 1♀ Wilkie Ck, L. Broadwater area,  21.ii.1987, A. E[wart]; 1 M Mt Nebo, SEQ, H. Horton,  x.1987;  1♂‘ Rockwood’ Chinchilla, 26°55’S 150°25’E, Iron Bark Ridge,  29.i.1983; 1♂ L. Broadwater, SW. area, on  A. spectabilis, 30kmSW. Dalby, Q, 28.xi.1987, D. M. Reeves;  1♂ L. Broadwater,  Acacia, A. E[wart], ~ 30kmSW Dalby,  3.xi.1990;  1♂  Hellhole CreekQuarry, Auburn Rd, Baracula S.F., Q,  15.xii.1997; 1♂ Doolandella, S. E. Qld, 7.iii.1994, [A. Ewart]; 1♂ Doolandella, S. E. Qld, 18–19.xii.1995, [A. Ewart]; 3♂ L. Broadwater, E.P., SEQ., Bush Camp Entrance, A. E[wart], I. R[attray]; 3.i.2000, 27°19.89’S 151°05.76’E; 1♂ L. Broadwater, E.P., SEQ., Bush Camp Entrance, A. E[wart], I. R[attray]; 3.i.2000, 27°19,89’S 151°05.76’E, Recorded;  1♂ Binjour Plateau, 17 kmNW. Gayndah, S.E.Q.  22.xii.2000, A. E[wart], 25°31.99’S 151°30.00’E, Fieldrecording;  1♂ Leslie Dam, Warwick, SEQ,  30.xi.2001, A. E[wart], 28°14.32’S 151°55.54’E, Recorded;  1♂ Leslie Dam, Warwick, SEQ,  1.xii.2001, A. E[wart], Openwoodland, 28°15.13’S 151°55.67’E;  1♂‘ Allinga’pty, N. ChinchillaSEQ. Brigalow, A. E[wart],  9.i.2002, 26°39.79’S 150°38.06’E( all AE);  1♀ Anstead Qld,  16.xii.1999, M. Coombs;  1♀ Glastonbury Qld,  31.x.1999, G. Maywald, 26°13’S 152°32’E;  1♂ Eumundi Qld,  11.xii.2001, M. Coombs;  1♂ Gunalda Qld,  14.xii.2001, M. Coombs(all MC);  1♂ Benarkin, Blackbutt,  7.xi.1993, R. Eastwood;  1♀AU.QLD.YLC, Yarrill Ck, nr Boggabilla,  223m, 28°18.0’S 150°17.7’E,  2.i.2004, Cooley, Hill, Marshall, Moulds;  1♂ 16 kmS. of Miles,  19.xii.1987, T.A. Moulds;  2♀ Kinbombi Falls, near Goomeri,  19.xii.1976, M.S. & B.J. Moulds;  2♂ 2♀ Jimna, N.of Kilcoy,  17.xii.1979, A. Hiller;  1♂ IslaGorge,  22.x.1989, R. Eastwood;  1♂ 2♀ Coominglah Range, 24 kmN. of Monto,  6.i.1975, M.S. & B.J. Moulds, ♂ genitalia prep C66;  1♂ St Lucia, Brisbane,  10.viii.1980, I. Gynther; 1♂ 1♀ Wyberba, S. of Stanthorpe, ex. Coll. Vernon Fanning, purchased 1984, no other data;  1♂ Kenmore,  26.x.1984, J. North;  1♂ St Lucia,  24.ix.1983, J. North;  1♂ St Lucia, Brisbane,  10.x. 1987, R. Van Barneveld;  1♂ Ashgrove, Brisbane,  5.x.1983, J.T. North;  4♂ Leslie Dam, Warwick,  1.i.1993, R. Eastwood;  1♂[damaged], same data as previous,  7.i.1990;  1♂ Lake Broadwater, nr Dalby, site A, 27°21’S 151°06’E,  25.x.1986, G. and A. Daniels;  1♂same data as previous,  26.xii.1986;  1♂same data as previous, site B,  30.i.1987;  1♂same data as previous,  30.ii.1987;  1♀ St Lucia,  9.xii.1983, C. E. Hagan;  1♂ Canningvalevia Warwick,  13.i.1993, R. Eastwood;  1♂same data as previous,  6.ii.1993;  1♀AU.QL.LBW, nr Lake Broadwater, 27°18.846’S 151°5.962’E,  357 m,  10.i.2002, Cooley, Hill, Marshall, Cowan& Moulds;  2♂ 1♀foot of Blackbutt Rg., 13 kmE. of Blackbutt, 26°53.34’S 152°12.88’E,  9.i.2002, Cooley, Cowan, Hill, Marshall; 1♀ S. Gympie, 23.xi.1986, R. Eastwood;  1♂AU.QL. SUN, Sundown N.P., E. of Texas, 28°55.184’S 151°34.9’E,  385 m,  4.i.2009, Hill, Marshall, Moulds, Owen;  1♂AU.QL.TXN, ~ 28 kmS. of Inglewood, 28°40.369’S 151°8.958’E,  347 m,  3.i.2009, Hill, Marshall, Moulds, Owen;  3♂ Wildash, Warwick,  14.i.1993, R. Eastwood;  1♂same data as previous,  20.ii.1995;  1♂same data as previous,  22.ii.1995;  1♂same data as previous,  9.xii.1994;  1♂ Blackbutt,  21.i.1989, R. Eastwood;  1♂ Connolly Dam, Warwick,  10.i.1990, R. Eastwood;  1♂ QueenslandUniversity, St Lucia, Brisbane, 27°30’S 153°06’E,  29.ii.1998, G. Daniels;  1♂ Capalaba, Brisbane,  8 Jan 2002, Cooley, Cowan, Hill, Marshall, Moss, Moulds; (  MSM); NEW SOUTH WALES:  1♂ Australia, N.S.W., Oaky Hillvia Legume,  1.xi.2003, L. Popple, R. MacSloy, 231-0021;  1♂ Australia NSW, 4 kmSSE. of Stannum,  4.i.2016, SL 100704, L. W. Popple, Recorded, 29.35249°S 151.803142°E, 231-0040 (both LWP);  3♂ Tamworth NSW,  16.i.1992, M. Coombs, 1 M Inverell NSW,  4.ii.1995, M. Coombs(all MC);  4♂ 1♀ WarrumbungleNPk, 31°29.22’S, 149°00.76’E,  420m,  16.ii.15, N. & D. Emery;  1♂ Whitegum Lookout, WarrumbungleNPk, 31°18.14’S, 149°02.05’E,  680 m,  12.i.14, C. & D. Emery;  1♂ Spring Ridge, 31°39.11’S; 150°25.48’E, 15.ii.15, N. & D. Emery;  1♂ 64 kmE Ebor, 30°04.08’S, 152°38.08’E,  654m,  4.i.13, N., C. & D. Emery;  2♂ 1♀ Killarney Gap, nr Bingara, 30°08.12’S, 150°04.33’E,  600m,  02.xii.13, N., C. & D. Emery(all DE); 5♂ 1♀ NSW.MRE, 50kmS. of Moree, N.S.W., 29°53.0’S 149°47.3’E, 1.i.2004, Cooley, Hill, Marshall, Moulds;  1♂same data as previous, VOUCHER 04. NSW.MRE.14, T. oldfieldi; 1♂ AU.NSW.INX ~ 15kmE. of Delungra, N.S.W., 29°43.837’S 150°56.804’E, 700m, 1.i.2005, Hill, Marshall, Moulds; 1♀ AU.NSW.INW, C. 19kmE. of Delungra, N.S.W., 29°44.097’S 150°59.147’E, 707m, 1.i.2005, Hill, Marshall, Moulds;  1♂same data as previous, VOUCHER: SIMON  LAB05.AU.NSW.INW.02,  Cicadetta oldfieldi; 1♂ AU.  NSW.ROP, c. 3kmW of  Warialda, N.S.W, 29°33.076’S 150°35.263’E,  591m,  1.i.2005, Hill, Marshall, Moulds;  6♂ 11 kmN. of Baradine,  1.i.1974, M.S. & B.J. Moulds;  2♂ 3♀ Afterleenear Kyogle,  10.i.1975, M.S. & B.J. Moulds;  1♂ Inverell,  2.i.1985, M.S. & B.J. Moulds;  1 ♂ Kyogle,  7.i.1985, W. Rixon(  MSM).   Etymology. Named after Mr Robert MacSloy of Alexandra Hills, Queensland. Mr MacSloy is one of only a small number of people who have the ability to recognise cicada species from their calls, and he has used this skill to contribute extensively to the current knowledge of the distributions and seasonality of cicadas over many years, particularly in south-east Queensland.   Diagnosis(Plate 1C, 1D; Figs 2B, 8A, 9A). Male and female specimens match the description given for  E. oldfieldi, though with some differences, as follows. Ventral surface of head mostly green to pale yellow-brown; postclypeus green, often with some dark brown coloration along midline; anteclypeus green to pale brown; rostrum pale brown anteriorly, becoming brown to dark brown towards apex. Dorsal surface of head green to olive-brown with a black band extending between the eyes and surrounding the interior margins of ocelli, though not extending to anterior margin; ocelli pink. Eyes deep red in living specimens, faded to brown in preserved specimens. Antennae dark brown, paler towards apex. Thorax green, sometimes fading to olive in dried specimens. Pronotum green with a thin, brown midline extending from the back of the head. Mesonotum green to olive-brown; midline broad, brown to pale brown, with lateral edges distinctly yellow; submedian sigilla darker than midline, brown to dark brown and typically darkest along parapsidal sutures; lateral sigilla green to olive-brown, sometimes indistinct. Legs with coxae and femora green or olive brown; tibia olive to pale brown; tarsi pale brown, becoming darker brown towards apex of claws. Wings with fore wing costal veins pale green to pale brown; pterostigma light brown. Hind wing veins pale brown or light green, becoming darker on posterior side of apical cells. Abdomen with tergites green or faded to olive-green; midline approximately 1 mmwide, brown, bordered by pale brown or yellow. Sternites pale green to yellow brown. Female abdominal segment 9 green, with a brown midline; ovipositor sheath extends approximately 1 mmbeyond the termination of abdominal segment 9.   Measurements. N= 26♂ 8♀. Means and ranges (in parentheses), mm; BL: ♂17.1 (15.3–18.6), ♀21.4 (19.9– 23.1); FWL: ♂22.2 (19.3–25.9), ♀25.7 (23.7–26.5); FWW: ♂7.5 (6.8–8.0), ♀8.6 (8.2–9.0); HW: ♂6.1 (5.3– 7.0), ♀6.8 (6.4–7.0); PW: ♂5.2 (4.5–6.2), ♀5.9 (5.6–6.3); AW: ♂5.2 (4.8–5.8), ♀5.7 (5.3–6.0).  Distinguishing features.  Ewartia roberti  n. sp.has a conspicuous, brown stripe medially along the dorsum, which extends from the proximal edge of the pronotum posteriorly to the apex of the abdomen. The stripe is distinctly narrow on the pronotum and to some extent along the abdomen, but is contrastingly broad on the mesonotum. This feature distinguishes it readily from  E. brevis,  E. carina  n. sp.and  E. cuensis.A similar stripe is also present in  E. etesia  n. sp.,  E. lapidosa  n. sp.,  E. oldfieldiand  E. thamna  n. sp.; however, its presentation in  E. roberti  n. sp.is subtly unique. In  E. roberti  n. sp., the stripe is narrower than the distance between the inner margins of the lateral ocelli. This distinguishes it from both  E. oldfieldiand  E. lapidosa  n. sp., in which the stripe is conspicuously broader than the distance between the lateral ocelli, and most conspicuously broader at the anterior margin of the pronotum (compare Figs 2A, 2B, 2C). It also distinguishes it from  E. etesia  n. sp., in which the stripe is equal to or broader than the distance between the inner margins of the lateral ocelli ( Figure 2D). In  E. roberti  n. sp., the black colouration on the dorsal side of the head is extensive, and, in particular, it surrounds the lateral ocelli. This distinguishes it from  E. thamna  n. sp., which has no black colouration on the dorsal side of the head. In addition, females of  E. roberti  n. sp.can be distinguished from  E. oldfieldiand  E. etesia  n. sp.by their short ovipositor length, which extends < 1 mmbeyond the apex of abdominal segment 9. In  E. oldfieldi, this extends 2.0– 2.5 mm, and in  E. etesia  n. sp., it extends approximately 3 mm.   Distribution and habitat. This species occurs from near Monto and IslaGorge in south-east Queenslandsouth to near Macksville, Spring Ridge and the Warrumbungle National Park in northern New South Wales( Fig. 10). Adults are present from late August until April. It occurs in association with various wattles (  Acaciaspp.) in dry sclerophyll forests, heathlands and also in urban environments, including road verges.  Acacia fimbriatais a major adult food plant in Brisbane. In Brisbane Forest Park,  A. glaucocarpais inhabited. In the inland, specimens have been collected from  A. spectabilis. Other wattle species with small phyllodes tend to be favoured. Acacias with large, fleshy phyllodes (e.g.  A. concurrens), are only utilised occasionally. In most seasons,  E. roberti  n. sp.is a common species throughout its range. Local congregations of singing males are often prominently scattered throughout areas of suitable habitat. Adults are active from Late August or September to May, being most common from October to February. This species overlaps in geographical distribution with  E. oldfieldiand  E. lapidosa  n. sp., and sometimes occurs in sympatry with each of these taxa.  Calling song. The complex calling song mode of  E. roberti  n. sp.( Figs 11–13) contains repeated, exceptionally long subphrases (25.3– 36.6 sduration; all statistics n= 70 recordings). Each subphrase is composed of a long series of discrete macrosyllables (each 2–5 syllables, 0.029– 0.080 sduration), each separated by a (typically increasing) set of 1–5 discrete syllables (each 11–13 ms duration, gaps 0.053– 0.092 sduration), which is then followed by series of 3–4 echemes of increasing duration (0.107– 0.441 s), each punctuated by syllable sequences of decreasing duration (each between 0.337 and 1.234 s, containing intersyllable gaps of 0.009– 0.045 sduration), a climatic echeme (0.296– 2.351 sduration) and, finally, a long syllable sequence (4.416– 7.407 sduration, with intersyllable gaps of 0.031– 0.084 sduration). An accentuation then follows the syllable sequence at the end of most subphrases. This comprises typically 3–4 macrosyllables (each containing 2–5 syllables), each separated either by a syllable sequence (1–5 syllables, with intersyllable gaps of 0.031– 0.086 sduration) or, more rarely, an uninterrupted gap (0.142– 0.201 s), and followed by a final gap (0.123– 0.155 sduration). This calling song mode is typically produced at intervals throughout the day. The simple calling song mode of this species ( Fig. 14) contains monotonously repeated phrases (0.3– 1.1 sduration). Each phrase typically contains a single echeme (0.25– 1.07 sduration) followed by a gap (0.053– 0.118 sduration); however, as the call progresses, the long echemes may be preceded by 1–2 discrete syllables or macrosyllables (each 0.011– 0.062 sduration), each followed by gaps (0.026– 0.064 sduration). This calling song mode predominates at dusk and is also produced occasionally at intervals during the day. The presentation of the frequency spectra does not change between song modes. In each mode the calling song typically has a highest amplitude frequency plateau between 10.0 and 14.5 kHz, with a dominant frequency between 11.3 and 13.5 kHz (mean=12.3, n=22; e.g. Fig. 7B). The structure of the calling song of  E. roberti  n. sp.is remarkably consistent across its wide distribution. The echeme, syllable sequence and subphrase durations do exhibit some variation, but this variation appears to be present within and among individuals in the broader population, without exhibiting any clear spatial pattern. Much of this variation considered likely to be at least partly temperature dependent. 1502352774 2004-03-11 2004-03-12 2004-03-11 Australia -28.983334 Glenlyon Dam 1229 151.46666 11 412 1 1 Texas holotype 1502352772 1990-02-28 L. & W. Popple. & Brisbane Q. Australia Indooroopilly 11 412 1 1 QUEENSLAND paratype 1502352768 1990-02-11 1997-12-28 1990-02-11 NSW A. I. Knight & M Mackellar Ra. Rd & Bungabbeee SF & Watkins Australia 180 Kyogle Stanthorpe 11 412 2 2 QUEENSLAND paratype 1502352785 1972-01-29 1972-01-30 1972-01-29 H. Frauca Australia Mt Woowoonga Dallarnil Bundaberg 11 412 1 1 QUEENSLAND paratype 1502352784 1972-01-10 H. Frauca Australia Biggenden 11 412 1 1 QUEENSLAND paratype 1502352765 [395,1133,1430,1455] 1971-01-11 H. Frauca Australia 518 Bluff Range Biggenden 11 412 2 2 QUEENSLAND paratype 1502352799 1972-12-12 H. Frauca Australia 518 Bluff Range Biggenden 11 412 1 1 QUEENSLAND paratype 1502352790 [295,712,1466,1491] 1977-01-13 H. Frauca Australia 518 Bluff Range Biggenden 11 412 1 1 QUEENSLAND paratype 1502352797 1972-12-09 H. Frauca Australia Biggenden South Bluff Range 11 412 1 1 QUEENSLAND paratype 1502352802 [242,840,1502,1527] 1972-12-19 1972-12-20 1972-12-19 H. Frauca Australia Biggenden South Bluff Range 11 412 2 2 QUEENSLAND paratype 1502352810 1972-01-03 H. Frauca Australia 671 Biggenden Mt Walsh 11 412 3 1 2 QUEENSLAND paratype 1502352806 [387,1155,1538,1563] 1976-12-04 Biggenden Q. & H. Frauca Australia Coast 11 412 1 1 Coast paratype 1502352808 1976-12-23 H. Frauca Australia Bundaberg Spring Ck 11 412 1 1 Coast paratype 1502352796 1985-12-19 ANIC Toowoomba Q. & I. F. B. Common Australia 620 -27.33 Prince Henry Heights 742 151.59 11 412 1 1 Coast paratype 1502352807 1997-12-21 C. J. Burwell Australia Qld 340 Binjour Plateau 151.469 Sx 11 412 2 2 Coast paratype 1502352803 1998-12-18 1998-12-19 1998-12-18 G. Monteith & C. Gough & G. Maywald Australia A. Ewart -25.7 Site 1245 151.43333 11 412 2 1 1 Coast paratype 1502352789 1908-10-26 Hamlyn-Harris Australia Toowoomba 11 412 1 1 Coast paratype 1502352791 [151,1390,1754,1779] 2001-12-17 2001-12-18 2001-12-17 G. Monteith & S. Wright Australia 330 Barakula 150.51666 Sx 11 412 5 2 3 Coast paratype 1502352798 2001-12-26 2001-12-29 2001-12-26 G. Montieth Australia Glen Witheren' General 153.11667 Sx 11 412 1 1 Coast paratype 1502352811 1997-12-20 C. J. Burwell & S. Evans Australia 460 Hurdle Gully 150.98334 11 412 1 1 Coast paratype 1502352793 1990-11 C. J. Burwell Australia A. Ewart Samford 11 412 1 1 Coast paratype 1502352809 2003-01-24 L. & W. Popple Australia Binjour 11 412 1 1 Queensland paratype 1502352787 2001-12-11 Hand & L. W. Popple Australia -28.161388 Thane 20 151.96638 11 412 3 1 2 Queensland paratype 1502352801 2001-12-11 N. Warwick & L. Popple & J. Moss Australia Willowvale 11 412 1 1 Queensland paratype 1502352792 Warwick, L. & Popple, J. & Genitalia Australia 12 413 -28.162 Thanes Creek 55 151.6287 11 412 1 1 Queensland paratype 1502352795 1998-03-06 L. Popple Australia Ipswich Dinmore 12 413 1 1 Queensland paratype 1502352800 [534,1387,188,213] 1998-03-03 L. Popple Australia Brisbane Camp Hill 12 413 1 1 Queensland paratype 1502352794 2001-12-27 2001-12-28 2001-12-27 N. Glenmorgan & L. Popple & A. Strange Australia Myall Park 12 413 1 1 Queensland paratype 1502352804 2001-12-26 L. Popple & A. Strange Australia Crows Nest Austin Rd 12 413 1 1 Queensland paratype 1502352805 2001-12-26 L. Popple & A. Strange Australia Toowoomba Highfields 12 413 1 1 Queensland paratype 1502352788 [667,1390,295,320] 2000-12-10 L. Popple & S. Billington Australia Lytton 12 413 1 1 Queensland paratype 1502352822 1998-11-07 L. Popple Australia Heights 12 413 3 3 Queensland paratype 1502352814 1999-12-03 W. Popple Australia Heights 12 413 1 1 Queensland paratype 1502352823 2003-01-07 2003-01-08 2003-01-07 L. W. Popple & Recorded L. W. Australia Brisbane Heights 12 413 1 1 Queensland paratype 1502352836 [365,1390,404,429] 2003-01 L. W. Popple Australia Brisbane Heights 12 413 1 1 Queensland paratype 1502352826 2002-12-21 R. J. Popple Australia Brisbane Heights 12 413 1 1 Queensland paratype 1502352835 2004-01-11 M. Larsen Australia -27.3 Mount Glorious 1237 152.46666 Qld 12 413 1 1 paratype 1502352828 Afterlee Public School & L. Popple & R. MacSloy Australia -28.566668 Kyogle district 1231 152.81667 12 413 1 1 Queensland paratype 1502352820 2004-12-28 2004-12-29 2004-12-28 L. W. Popple Australia -25.378334 Burnett River 21 151.12944 Eidsvold 12 413 1 1 Queensland paratype 1502352833 2005-01-28 L. Popple & R. MacSloy Australia -28.162 Warwick 55 151.6287 Thanes Creek 12 413 1 1 Queensland paratype 1502352832 2005-02-19 L. W. Popple & R. MacSloy Australia -27.413055 Seventeen Mile Road Helidon Hills 12 413 1 1 Queensland paratype 1502352816 2005-10-28 L. W. Popple Australia -25.605 Mt Debatable 21 151.54056 12 413 1 1 Queensland paratype 1502352813 2005-12-05 2005-12-10 2005-12-05 Southwood NP & L. Popple & A. Ewart Australia -27.834723 Moonie 20 150.11307 12 413 1 1 Queensland paratype 1502352821 2005-12-14 2005-12-15 2005-12-14 L. W. Popple Australia -27.350555 Dalby 20 151.09277 Lake Broadwater 12 413 4 3 1 Queensland paratype 1502352830 2008-03 Australia -27.449444 Murrarie 20 153.09833 Metroplex 12 413 1 1 Queensland paratype 1502352817 2007-12-11 Burrandoan Rd, SW Australia -26.598333 Kingaroy 20 151.74945 12 413 1 1 Queensland paratype 1502352827 2007-12-10 A.F. Sanborn Australia -26.855278 Beerwah 20 153.00667 Roys Rd 12 413 1 1 paratype 1502352812 2010-01-22 L. W. Popple Australia -28.225279 Australia 20 152.01999 12 413 1 1 paratype 1502352829 [718,1287,943,968] 1975-11-06 A. Ewart Australia Crows Nest Iron Bark 12 413 1 1 paratype 1502352815 1975-11-22 A. Ewart Australia Crows Nest Iron Bark 12 413 3 1 2 paratype 1502352825 [634,1150,980,1005] 1975-10-19 A. Ewart Australia Perseverance Dam 12 413 2 2 paratype 1502352824 1977-11-01 A. Ewart Australia Crows Nest Ironbark Qld 12 413 1 1 paratype 1502352818 [494,1151,1015,1040] 1984-02-12 L. Broadwater & A. Ewart Australia Qld Dalby 12 413 1 1 paratype 1502352831 1987-02-21 1987-10 1987-02-21 Wilkie Ck, L. & M Mt Nebo & H. Horton Australia Broadwater area 12 413 4 1 3 paratype 1502352834 1983-01-29 Australia -26.916666 Iron Bark Ridge 1239 150.41667 Rockwood' Chinchilla 12 413 1 1 paratype 1502352819 [498,1281,1124,1149] 1990-11-03 L. Broadwater Australia Dalby 12 413 1 1 paratype 1502352848 1997-12-15 Baracula S. F. Australia Hellhole Creek Quarry Auburn Rd Hellhole Creek 12 413 1 1 paratype 1502352843 2000-12-22 Gayndah, S. E. Q. & Field Australia -25.533167 Binjour Plateau 12 151.5 12 413 1 1 paratype 1502352846 2001-11-30 Recorded Australia -28.238667 Warwick 12 151.92566 Leslie Dam 12 413 1 1 paratype 1502352859 2001-12-01 Open Australia -28.252167 Warwick 12 151.92783 Leslie Dam 12 413 1 1 paratype 1502352851 2002-01-09 N. Chinchilla & Brigalow, A. E Australia -26.663166 Allinga' 12 150.63434 12 413 1 1 paratype 1502352852 [674,1174,1412,1437] 1999-12-16 M. Coombs Australia Anstead Qld 12 413 1 1 paratype 1502352856 1999-10-31 G. Maywald Australia -26.216667 Glastonbury Qld 1243 152.53334 12 413 1 1 paratype 1502352842 [687,1209,1447,1472] 2001-12-11 M. Coombs Australia Eumundi Qld 12 413 1 1 paratype 1502352861 2001-12-14 M. Coombs Australia Gunalda Qld 12 413 1 1 paratype 1502352853 [560,1121,1484,1509] 1993-11-07 R. Eastwood Australia Blackbutt Benarkin 12 413 1 1 paratype 1502352847 2004-01-02 Cooley & Hill & Marshall & Moulds Australia 223 -28.3 Boggabilla 123 150.295 Yarrill Ck 12 413 1 1 paratype 1502352839 1987-12-19 T. A. Moulds Australia Miles 12 413 1 1 paratype 1502352845 [451,1230,1556,1581] 1976-12-19 M. S. & B. J. Moulds Australia Goomeri Kinbombi Falls 12 413 2 2 paratype 1502352860 1979-12-17 Jimna, N. & A. Hiller Australia Kilcoy 12 413 4 2 2 paratype 1502352841 [527,973,1591,1616] 1989-10-22 R. Eastwood Iceland Isla 12 413 1 1 paratype 1502352855 1975-01-06 M. S. & B. J. Moulds Iceland Monto Coominglah Range 12 413 3 2 1 paratype 1502352844 [849,1388,1628,1653] 1980-08-10 I. Gynther Iceland Brisbane St Lucia 12 413 1 1 paratype 1502352854 1984-10-26 J. North Iceland Kenmore 12 413 1 1 paratype 1502352840 [259,643,1700,1725] 1983-09-24 J. North Iceland St Lucia 12 413 1 1 paratype 1502352850 [655,1265,1700,1725] 1987-10-10 R. Van Barneveld Iceland Brisbane St Lucia 12 413 1 1 paratype 1502352849 1983-10-05 J. T. North Iceland Brisbane Ashgrove 12 413 1 1 paratype 1502352837 [517,1090,1735,1760] 1993-01-01 R. Eastwood Iceland Warwick Leslie Dam 12 413 4 4 paratype 1502352857 1990-01-07 R. Eastwood Iceland Warwick Leslie Dam 12 413 1 1 paratype 1502352858 [372,1388,1772,1797] 1986-10-25 G. and A. Daniels Iceland -27.35 Dalby 1237 151.1 Lake Broadwater 12 413 1 1 paratype 1502352838 1986-12-26 G. and A. Daniels Iceland -27.35 Dalby 1237 151.1 Lake Broadwater 12 413 1 1 paratype 1502352874 [579,1104,1807,1832] 1987-01-30 G. and A. Daniels Iceland -27.35 Dalby 1237 151.1 Lake Broadwater 12 413 1 1 paratype 1502352869 1987-02-30 G. and A. Daniels Iceland -27.35 Dalby 1237 151.1 Lake Broadwater 12 413 1 1 paratype 1502352870 [279,699,1844,1869] 1983-12-09 C. E. Hagan Iceland St Lucia 12 413 1 1 paratype 1502352884 [710,1325,1844,1869] 1993-01-13 R. Eastwood Iceland Warwick Canningvale 12 413 1 1 paratype 1502352879 1993-02-06 R. Eastwood Iceland Warwick Canningvale 12 413 1 1 paratype 1502352878 2002-01-10 Cooley & Hill & Marshall & Cowan & Moulds Iceland 357 -27.3141 Lake Broadwater 1 151.09937 12 413 1 1 paratype 1502352876 2002-01-09 Cooley & Cowan & Hill & Marshall Iceland -26.889 Blackbutt 12 152.21466 12 413 3 1 2 paratype 1502352866 2009-01-04 SUN Sundown N. P. & Hill & Marshall & Moulds & Owen Iceland 385 -28.919733 Texas 81 151.58167 12 413 1 1 Texas paratype 1502352897 2009-01-03 Hill & Marshall & Moulds & Owen Iceland 347 13 414 -28.672817 Inglewood 1 151.1493 12 413 1 1 Texas paratype 1502352880 [151,686,188,213] 1993-01-14 R. Eastwood Iceland Warwick Wildash 13 414 3 3 Texas paratype 1502352867 [699,1124,188,213] 1995-02-20 R. Eastwood Iceland Warwick Wildash 13 414 1 1 Texas paratype 1502352871 1995-02-22 R. Eastwood Iceland Warwick Wildash 13 414 1 1 Texas paratype 1502352865 [284,727,223,248] 1994-12-09 R. Eastwood Iceland Warwick Wildash 13 414 1 1 Texas paratype 1502352883 [743,1195,223,248] 1989-01-21 R. Eastwood Iceland Blackbutt 13 414 1 1 Texas paratype 1502352873 1990-01-10 R. Eastwood Iceland Warwick Connolly Dam 13 414 1 1 Texas paratype 1502352877 1998-02-29 G. Daniels Iceland -27.5 Brisbane 1236 153.1 St Lucia 13 414 1 1 Queensland paratype 1502352868 2002-01-08 MSM Cooley & Cowan & Hill & Marshall & Moss & Moulds Iceland Brisbane Capalaba 13 414 1 1 NEW SOUTH WALES paratype 1502352881 [361,1389,332,357] 2003-11-01 L. Popple & R. MacSloy Australia Legume Oaky Hill 13 414 1 1 paratype 1502352863 2016-01-04 NSW L. W. Popple & Recorded Australia -29.35249 Stannum 1 151.80315 13 414 1 1 paratype 1502352875 1992-01-16 1995-02-04 1992-01-16 NSW M. Coombs & M Inverell Australia Tamworth 13 414 3 3 paratype 1502352872 [227,1254,439,464] 2015-02-16 N. & D. Emery Australia 420 -31.487 Warrumbungle 12 149.01266 13 414 5 1 4 paratype 1502352862 2014-01-12 C. & D. Emery Australia 680 -31.302334 Warrumbungle 12 149.03416 Whitegum Lookout 13 414 1 1 paratype 1502352888 Australia -31.651833 Spring Ridge 13 414 1 1 paratype 1502352882 2013-01-04 N., C. & D. Emery Australia 654 -30.068 Ebor 12 152.63467 13 414 1 1 paratype 1502352864 2013-12-02 N., C. & D. Emery Australia 600 -30.135334 Bingara 12 150.07217 Killarney Gap 13 414 3 1 2 paratype 1502352896 [254,1053,620,645] 2013-12-02 NSW N., C. & D. Emery Australia 600 -30.135334 Bingara 12 150.07217 Killarney Gap 13 414 1 1 paratype 1502352893 2013-12-02 SIMON, LAB N., C. & D. Emery Australia 600 -30.135334 Bingara 12 150.07217 Killarney Gap 13 414 LAB 05 1 1 paratype 1502352891 2005-01-01 NSW Warialda, N. S. W & Hill & Marshall & Moulds Australia 591 -29.551266 Warialda 1 150.58772 13 414 1 paratype 1502352920 1974-01-01 M. S. & B. J. Moulds Australia Baradine 13 414 6 6 paratype 1502352901 [610,1290,799,824] 1975-01-10 M. S. & B. J. Moulds Australia Kyogle Afterlee 13 414 5 3 2 paratype 1502352900 1985-01-02 M. S. & B. J. Moulds Australia Inverell 13 414 1 1 paratype 1502352903 [498,956,836,861] 1985-01-07 MSM W. Rixon Australia Kyogle 13 414 1 1 paratype