Ewart, 1988 : 182
A revision of the Ewartia oldfieldi (Distant) species complex (Hemiptera: Cicadidae: Cicadettinae) with five new species from eastern and northern Australia
Popple, Lindsay W.
Zootaxa
2017
4263
3
401
449
8JSHT
[151,336,1077,1103]
Insecta
Cicadidae
Ewartia
Animalia
Hemiptera
11
412
Arthropoda
species
roberti
sp. nov.
Cicadetta oldfieldi(Distant): Ewart, 1988: 182–183, 191, 193, 196–197, Fig. 9J–M, Plate 2C. Types. Holotype: ♂[typed] ‘ 28°59’S 151°28’E’/ ’ AUSTRALIAQLD’ / ‘ Glenlyon Dam, via’ / ‘ Texas, SEQ.’ / ’ 11–12.Mar.2004. ’ / ‘ L. & W. Popple.’ / ‘231-0023’ / ‘QM reg. no. T207359’ ( QM); Paratypes: QUEENSLAND: 1♂ Indooroopilly, Brisbane Q., 28.ii.1990; 2♀ Stanthorpe, 11.ii.1990, A. I. Knight; 1 M Mackellar Ra. Rd, Bungabbeee SF, 15kmSE. Kyogle, NSW, 28.xii.1997, mv light, Watkinselev. 180m; 1♀QLD: Mt Woowoongaca 50mi. W. of Bundabergvia Dallarnil, 29–30.i.1972, H. Frauca; 1♂NW slope of Bluff Rg. nr BiggendenQLD, 10.i.1972, H. Frauca; 2♂QLD Biggenden, Bluff Range, 11.i.1971, H. Frauca, c. 1700ft; 1♂same data as previous, 12.xii.1972; 1♂same data as previous, 13.i.1977; 1♂QLD: Biggenden, Bluff Range, Southfhills, 9.xii.1972, H. Frauca; 2♂same data as previous, 19–20.xii.1972, H. Frauca; 2♂ 1♀QLD: Mt Walshsummit, 2200ft, Biggenden, 3.i.1972, H. Frauca; 1♂ CoastRanges, 11kmS. of Biggenden Q., 4.xii.1976, H. Frauca; 1♀QLD: Spring Ckca 30kmS. of Bundaberg, 23.xii.1976, H. Frauca; 1♂ 27.33S 151.59E, Prince Henry Heights, 620m, Toowoomba Q., 19.xii.1985, I. F. B. Common( all ANIC); 2♂ Qld: 25.552° Sx 151.469°E, Binjour Plateau, Redvale Rd, 21.xii.1997, C. J. Burwell, open forest, 340m; 1♂ 1♀SEQ: 25°42’S 151°26’ENipping Gully, Site6, 18– 19.xii.1998, G. Monteith, C. Gough& G. Maywald, 7532, Cicadetta oldfieldidet. A. Ewart, 1997; 1♂ Toowoomba, 26.x.1908, Hamlyn-Harris, Melampsalta oldfieldi; 1♀ Toowoomba, 1908, Hamlyn-Harris, Melampsalta oldfieldi; 3♂ 2♀QLD: 26°26’ Sx 150°31’E, Barakula, 330m, 17–18.xii.2001, G.Monteith& S. Wright, M.V. light, 10311; 1♀QLD: 28°03’ Sx 153°07’E‘ Glen Witheren’ General, 26–29.xii.2001, G. Montieth, M. V. light, 10352; 1♂SEQ: 24°55’ 150°59’E, Hurdle Gully, 14.8kmWSW Monto, open forest, 460m, 20.xii.1997, C. J. Burwell, S. Evans; 1♂ 1F Gooroolba, S.E.Q., 1.xii.1952, W.W.Abell, Melampsalta oldfieldi; 1♂N. of Samford, xi.1990, C. J. Burwell, Cicadetta oldfieldidet. A. Ewart1997 ( all QM); 1♂ Australia, Queensland, 4kmW. of Binjour, SEQ, 24.i.2003, L. & W. Popple, mv lamp, 231-0001; 2♂ 1♀ Australia Queensland, 6 kmwest of Thane, 11.xii.2001, Handcollected, L. W. Popple, 28°09’41”S 151°57’59”E, 231-0002 to 231-0004; 1♂ Australia Queensland, Willowvale, 5 km N. Warwick, 11.xii.2001, L. Popple, J. Moss, 231-0005; 1♂ Australia Queensland, 28.162S° 151.6287°E, Thanes Creekvia Warwick, L. Popple, J.Moss, 231-0004, Genitaliaprep. 231-01; 1♂ Dinmore, Ipswich, SEQ, 6.iii.1998, L. Popple, Acaciasp., 231-0006; 1♂ Camp Hill, Brisbane, SEQ, 3.iii.1998, L. Popple, Acaciasp., 231-0007; 1♂ Myall Park, 8km N. Glenmorgan, SEQ, 27–28.xii.2001, L. Popple, A. Strange, 231-0008; 1♂ Austin Rd, Crows Nest, SEQ, mv lamp, 26.xii.2001, L. Popple, A. Strange, 231-0009; 1♂ Highfields, via Toowoomba, SEQ, 26.xii.2001, L. Popple, A. Strange, 231-0010; 1♂ Lytton, SEQ, 10.xii.2000, L. Popple, S. Billington, 231-0011; 3♂ Carina Heights, SEQ, 7.xi.1998, L. Popple, 231-0014-231-0016; 1♀ Carina Heights, SEQ, 3.xii.1999, W. Popple, 231-0017; 1♂ Australia, Queensland, Carina Heights, Brisbane, SEQ, 7–8.i.2003, L. W. Popple, Recorded L. W.Popple, 231-0018; 1♂ Australia, Queensland, Carina Heights, Brisbane, SEQ, i.2003, L. W. Popple, 231-0019; 1♂ Australia, Queensland, Carina Heights, Brisbane, SEQ, 21.xii.2002, R. J. Popple, 231-0020; 1♀ 27°18’S 152°28’E, Australia, Qld, Mount Glorious, 11.i.2004, M. Larsen, mv lamp, 231-0022; 1♂ Australia Queensland, 28°34’S 152°49’E, Afterlee Public School, Kyogle district, rural, L. Popple, R. MacSloy, 231–0024; 1♂ Australia Queensland, 25°22’42”S 151°07’46”E, Eidsvold, Burnett River, 28–29.xii.2004, L. W. Popple, 231-0025; 1♂ Australia Queensland, 28.162S° 151.6287°E, Thanes Creekvia Warwick, 28.i.2005, L.Poppleand R. MacSloy, 231-0027; 1♂ Australia Queensland, 27°24’47”S152°11’60”E, Helidon Hills, Seventeen Mile Road, 19.ii.2005, L. W. Popple, R. MacSloy, 231-0028; 1♂ Australia Queensland, 25°36’18”S 151°32’26”E, Mt Debatable(summit), 28.x.2005, L. W. Popple, Acaciasp., 231-0029; 1♂ Australia Queensland, Southwood NP, via Moonie, 5– 10.xii.2005, L. Popple, A. Ewart, 27°50’05”S 150°06’47”E, 231-0030; 1♂ 3♀ Australia Queensland, Lake Broadwater, via Dalby, 14–15.xii.2005, L. W. Popple, mv lamp, 27°21’02”S 151°05’34”E, 231-0031 to 231-0035; 1♂ Australia Queensland, Metroplex, Murrarie, 27°26’58”S 153°05’54”E, iii.2008, Ac. fimbriata, L. W. Popple, 231-0036; 1♂ Australia Queensland, Burrandoan Rd, SWof Kingaroy, 11.xii.2007, 26° 26°35’54”S 151°44’58”E, 231-0037; 1♂ AustraliaQLD, Roys Rd, Beerwah( F.S.C.), 26°51’19”S 153°00’24”E, 10.xii.2007, A. flavescens, L. Popple, R. MacSloy; 1♂ AustraliaQLD, Warwick— 5 kmS., parkland, Acaciasp., 22.i.2010, L. W. Popple, 28°13’31”S 152° 01’12”E, 231-0039 (all LWP); 1♀ Iron Bark, Crows Nest, 6.xi.1975, [ A. Ewart]; 2♂ 1♀ Iron Bark, Crows Nest, 22.xi.1975, [ A. Ewart]; 2♂ Perseverance Dam, 19.x.1975, [ A. Ewart]; 1♀ Ironbark Qld, Crows Nest, 1.xi.1977[ A. Ewart]; 1♂ L. Broadwater, Dalby, Qld, 12.ii.1984, [ A. Ewart]; 3♂ 1♀ Wilkie Ck, L. Broadwater area, 21.ii.1987, A. E[wart]; 1 M Mt Nebo, SEQ, H. Horton, x.1987; 1♂‘ Rockwood’ Chinchilla, 26°55’S 150°25’E, Iron Bark Ridge, 29.i.1983; 1♂ L. Broadwater, SW. area, on A. spectabilis, 30kmSW. Dalby, Q, 28.xi.1987, D. M. Reeves; 1♂ L. Broadwater, Acacia, A. E[wart], ~ 30kmSW Dalby, 3.xi.1990; 1♂ Hellhole CreekQuarry, Auburn Rd, Baracula S.F., Q, 15.xii.1997; 1♂ Doolandella, S. E. Qld, 7.iii.1994, [A. Ewart]; 1♂ Doolandella, S. E. Qld, 18–19.xii.1995, [A. Ewart]; 3♂ L. Broadwater, E.P., SEQ., Bush Camp Entrance, A. E[wart], I. R[attray]; 3.i.2000, 27°19.89’S 151°05.76’E; 1♂ L. Broadwater, E.P., SEQ., Bush Camp Entrance, A. E[wart], I. R[attray]; 3.i.2000, 27°19,89’S 151°05.76’E, Recorded; 1♂ Binjour Plateau, 17 kmNW. Gayndah, S.E.Q. 22.xii.2000, A. E[wart], 25°31.99’S 151°30.00’E, Fieldrecording; 1♂ Leslie Dam, Warwick, SEQ, 30.xi.2001, A. E[wart], 28°14.32’S 151°55.54’E, Recorded; 1♂ Leslie Dam, Warwick, SEQ, 1.xii.2001, A. E[wart], Openwoodland, 28°15.13’S 151°55.67’E; 1♂‘ Allinga’pty, N. ChinchillaSEQ. Brigalow, A. E[wart], 9.i.2002, 26°39.79’S 150°38.06’E( all AE); 1♀ Anstead Qld, 16.xii.1999, M. Coombs; 1♀ Glastonbury Qld, 31.x.1999, G. Maywald, 26°13’S 152°32’E; 1♂ Eumundi Qld, 11.xii.2001, M. Coombs; 1♂ Gunalda Qld, 14.xii.2001, M. Coombs(all MC); 1♂ Benarkin, Blackbutt, 7.xi.1993, R. Eastwood; 1♀AU.QLD.YLC, Yarrill Ck, nr Boggabilla, 223m, 28°18.0’S 150°17.7’E, 2.i.2004, Cooley, Hill, Marshall, Moulds; 1♂ 16 kmS. of Miles, 19.xii.1987, T.A. Moulds; 2♀ Kinbombi Falls, near Goomeri, 19.xii.1976, M.S. & B.J. Moulds; 2♂ 2♀ Jimna, N.of Kilcoy, 17.xii.1979, A. Hiller; 1♂ IslaGorge, 22.x.1989, R. Eastwood; 1♂ 2♀ Coominglah Range, 24 kmN. of Monto, 6.i.1975, M.S. & B.J. Moulds, ♂ genitalia prep C66; 1♂ St Lucia, Brisbane, 10.viii.1980, I. Gynther; 1♂ 1♀ Wyberba, S. of Stanthorpe, ex. Coll. Vernon Fanning, purchased 1984, no other data; 1♂ Kenmore, 26.x.1984, J. North; 1♂ St Lucia, 24.ix.1983, J. North; 1♂ St Lucia, Brisbane, 10.x. 1987, R. Van Barneveld; 1♂ Ashgrove, Brisbane, 5.x.1983, J.T. North; 4♂ Leslie Dam, Warwick, 1.i.1993, R. Eastwood; 1♂[damaged], same data as previous, 7.i.1990; 1♂ Lake Broadwater, nr Dalby, site A, 27°21’S 151°06’E, 25.x.1986, G. and A. Daniels; 1♂same data as previous, 26.xii.1986; 1♂same data as previous, site B, 30.i.1987; 1♂same data as previous, 30.ii.1987; 1♀ St Lucia, 9.xii.1983, C. E. Hagan; 1♂ Canningvalevia Warwick, 13.i.1993, R. Eastwood; 1♂same data as previous, 6.ii.1993; 1♀AU.QL.LBW, nr Lake Broadwater, 27°18.846’S 151°5.962’E, 357 m, 10.i.2002, Cooley, Hill, Marshall, Cowan& Moulds; 2♂ 1♀foot of Blackbutt Rg., 13 kmE. of Blackbutt, 26°53.34’S 152°12.88’E, 9.i.2002, Cooley, Cowan, Hill, Marshall; 1♀ S. Gympie, 23.xi.1986, R. Eastwood; 1♂AU.QL. SUN, Sundown N.P., E. of Texas, 28°55.184’S 151°34.9’E, 385 m, 4.i.2009, Hill, Marshall, Moulds, Owen; 1♂AU.QL.TXN, ~ 28 kmS. of Inglewood, 28°40.369’S 151°8.958’E, 347 m, 3.i.2009, Hill, Marshall, Moulds, Owen; 3♂ Wildash, Warwick, 14.i.1993, R. Eastwood; 1♂same data as previous, 20.ii.1995; 1♂same data as previous, 22.ii.1995; 1♂same data as previous, 9.xii.1994; 1♂ Blackbutt, 21.i.1989, R. Eastwood; 1♂ Connolly Dam, Warwick, 10.i.1990, R. Eastwood; 1♂ QueenslandUniversity, St Lucia, Brisbane, 27°30’S 153°06’E, 29.ii.1998, G. Daniels; 1♂ Capalaba, Brisbane, 8 Jan 2002, Cooley, Cowan, Hill, Marshall, Moss, Moulds; ( MSM); NEW SOUTH WALES: 1♂ Australia, N.S.W., Oaky Hillvia Legume, 1.xi.2003, L. Popple, R. MacSloy, 231-0021; 1♂ Australia NSW, 4 kmSSE. of Stannum, 4.i.2016, SL 100704, L. W. Popple, Recorded, 29.35249°S 151.803142°E, 231-0040 (both LWP); 3♂ Tamworth NSW, 16.i.1992, M. Coombs, 1 M Inverell NSW, 4.ii.1995, M. Coombs(all MC); 4♂ 1♀ WarrumbungleNPk, 31°29.22’S, 149°00.76’E, 420m, 16.ii.15, N. & D. Emery; 1♂ Whitegum Lookout, WarrumbungleNPk, 31°18.14’S, 149°02.05’E, 680 m, 12.i.14, C. & D. Emery; 1♂ Spring Ridge, 31°39.11’S; 150°25.48’E, 15.ii.15, N. & D. Emery; 1♂ 64 kmE Ebor, 30°04.08’S, 152°38.08’E, 654m, 4.i.13, N., C. & D. Emery; 2♂ 1♀ Killarney Gap, nr Bingara, 30°08.12’S, 150°04.33’E, 600m, 02.xii.13, N., C. & D. Emery(all DE); 5♂ 1♀ NSW.MRE, 50kmS. of Moree, N.S.W., 29°53.0’S 149°47.3’E, 1.i.2004, Cooley, Hill, Marshall, Moulds; 1♂same data as previous, VOUCHER 04. NSW.MRE.14, T. oldfieldi; 1♂ AU.NSW.INX ~ 15kmE. of Delungra, N.S.W., 29°43.837’S 150°56.804’E, 700m, 1.i.2005, Hill, Marshall, Moulds; 1♀ AU.NSW.INW, C. 19kmE. of Delungra, N.S.W., 29°44.097’S 150°59.147’E, 707m, 1.i.2005, Hill, Marshall, Moulds; 1♂same data as previous, VOUCHER: SIMON LAB05.AU.NSW.INW.02, Cicadetta oldfieldi; 1♂ AU. NSW.ROP, c. 3kmW of Warialda, N.S.W, 29°33.076’S 150°35.263’E, 591m, 1.i.2005, Hill, Marshall, Moulds; 6♂ 11 kmN. of Baradine, 1.i.1974, M.S. & B.J. Moulds; 2♂ 3♀ Afterleenear Kyogle, 10.i.1975, M.S. & B.J. Moulds; 1♂ Inverell, 2.i.1985, M.S. & B.J. Moulds; 1 ♂ Kyogle, 7.i.1985, W. Rixon( MSM).
Etymology. Named after Mr Robert MacSloy of Alexandra Hills, Queensland. Mr MacSloy is one of only a small number of people who have the ability to recognise cicada species from their calls, and he has used this skill to contribute extensively to the current knowledge of the distributions and seasonality of cicadas over many years, particularly in south-east Queensland.
Diagnosis(Plate 1C, 1D; Figs 2B, 8A, 9A). Male and female specimens match the description given for E. oldfieldi, though with some differences, as follows. Ventral surface of head mostly green to pale yellow-brown; postclypeus green, often with some dark brown coloration along midline; anteclypeus green to pale brown; rostrum pale brown anteriorly, becoming brown to dark brown towards apex. Dorsal surface of head green to olive-brown with a black band extending between the eyes and surrounding the interior margins of ocelli, though not extending to anterior margin; ocelli pink. Eyes deep red in living specimens, faded to brown in preserved specimens. Antennae dark brown, paler towards apex. Thorax green, sometimes fading to olive in dried specimens. Pronotum green with a thin, brown midline extending from the back of the head. Mesonotum green to olive-brown; midline broad, brown to pale brown, with lateral edges distinctly yellow; submedian sigilla darker than midline, brown to dark brown and typically darkest along parapsidal sutures; lateral sigilla green to olive-brown, sometimes indistinct. Legs with coxae and femora green or olive brown; tibia olive to pale brown; tarsi pale brown, becoming darker brown towards apex of claws. Wings with fore wing costal veins pale green to pale brown; pterostigma light brown. Hind wing veins pale brown or light green, becoming darker on posterior side of apical cells. Abdomen with tergites green or faded to olive-green; midline approximately 1 mmwide, brown, bordered by pale brown or yellow. Sternites pale green to yellow brown. Female abdominal segment 9 green, with a brown midline; ovipositor sheath extends approximately 1 mmbeyond the termination of abdominal segment 9.
Measurements. N= 26♂ 8♀. Means and ranges (in parentheses), mm; BL: ♂17.1 (15.3–18.6), ♀21.4 (19.9– 23.1); FWL: ♂22.2 (19.3–25.9), ♀25.7 (23.7–26.5); FWW: ♂7.5 (6.8–8.0), ♀8.6 (8.2–9.0); HW: ♂6.1 (5.3– 7.0), ♀6.8 (6.4–7.0); PW: ♂5.2 (4.5–6.2), ♀5.9 (5.6–6.3); AW: ♂5.2 (4.8–5.8), ♀5.7 (5.3–6.0). Distinguishing features. Ewartia roberti n. sp.has a conspicuous, brown stripe medially along the dorsum, which extends from the proximal edge of the pronotum posteriorly to the apex of the abdomen. The stripe is distinctly narrow on the pronotum and to some extent along the abdomen, but is contrastingly broad on the mesonotum. This feature distinguishes it readily from E. brevis, E. carina n. sp.and E. cuensis.A similar stripe is also present in E. etesia n. sp., E. lapidosa n. sp., E. oldfieldiand E. thamna n. sp.; however, its presentation in E. roberti n. sp.is subtly unique. In E. roberti n. sp., the stripe is narrower than the distance between the inner margins of the lateral ocelli. This distinguishes it from both E. oldfieldiand E. lapidosa n. sp., in which the stripe is conspicuously broader than the distance between the lateral ocelli, and most conspicuously broader at the anterior margin of the pronotum (compare Figs 2A, 2B, 2C). It also distinguishes it from E. etesia n. sp., in which the stripe is equal to or broader than the distance between the inner margins of the lateral ocelli ( Figure 2D). In E. roberti n. sp., the black colouration on the dorsal side of the head is extensive, and, in particular, it surrounds the lateral ocelli. This distinguishes it from E. thamna n. sp., which has no black colouration on the dorsal side of the head. In addition, females of E. roberti n. sp.can be distinguished from E. oldfieldiand E. etesia n. sp.by their short ovipositor length, which extends < 1 mmbeyond the apex of abdominal segment 9. In E. oldfieldi, this extends 2.0– 2.5 mm, and in E. etesia n. sp., it extends approximately 3 mm.
Distribution and habitat. This species occurs from near Monto and IslaGorge in south-east Queenslandsouth to near Macksville, Spring Ridge and the Warrumbungle National Park in northern New South Wales( Fig. 10). Adults are present from late August until April. It occurs in association with various wattles ( Acaciaspp.) in dry sclerophyll forests, heathlands and also in urban environments, including road verges. Acacia fimbriatais a major adult food plant in Brisbane. In Brisbane Forest Park, A. glaucocarpais inhabited. In the inland, specimens have been collected from A. spectabilis. Other wattle species with small phyllodes tend to be favoured. Acacias with large, fleshy phyllodes (e.g. A. concurrens), are only utilised occasionally. In most seasons, E. roberti n. sp.is a common species throughout its range. Local congregations of singing males are often prominently scattered throughout areas of suitable habitat. Adults are active from Late August or September to May, being most common from October to February. This species overlaps in geographical distribution with E. oldfieldiand E. lapidosa n. sp., and sometimes occurs in sympatry with each of these taxa. Calling song. The complex calling song mode of E. roberti n. sp.( Figs 11–13) contains repeated, exceptionally long subphrases (25.3– 36.6 sduration; all statistics n= 70 recordings). Each subphrase is composed of a long series of discrete macrosyllables (each 2–5 syllables, 0.029– 0.080 sduration), each separated by a (typically increasing) set of 1–5 discrete syllables (each 11–13 ms duration, gaps 0.053– 0.092 sduration), which is then followed by series of 3–4 echemes of increasing duration (0.107– 0.441 s), each punctuated by syllable sequences of decreasing duration (each between 0.337 and 1.234 s, containing intersyllable gaps of 0.009– 0.045 sduration), a climatic echeme (0.296– 2.351 sduration) and, finally, a long syllable sequence (4.416– 7.407 sduration, with intersyllable gaps of 0.031– 0.084 sduration). An accentuation then follows the syllable sequence at the end of most subphrases. This comprises typically 3–4 macrosyllables (each containing 2–5 syllables), each separated either by a syllable sequence (1–5 syllables, with intersyllable gaps of 0.031– 0.086 sduration) or, more rarely, an uninterrupted gap (0.142– 0.201 s), and followed by a final gap (0.123– 0.155 sduration). This calling song mode is typically produced at intervals throughout the day. The simple calling song mode of this species ( Fig. 14) contains monotonously repeated phrases (0.3– 1.1 sduration). Each phrase typically contains a single echeme (0.25– 1.07 sduration) followed by a gap (0.053– 0.118 sduration); however, as the call progresses, the long echemes may be preceded by 1–2 discrete syllables or macrosyllables (each 0.011– 0.062 sduration), each followed by gaps (0.026– 0.064 sduration). This calling song mode predominates at dusk and is also produced occasionally at intervals during the day. The presentation of the frequency spectra does not change between song modes. In each mode the calling song typically has a highest amplitude frequency plateau between 10.0 and 14.5 kHz, with a dominant frequency between 11.3 and 13.5 kHz (mean=12.3, n=22; e.g. Fig. 7B). The structure of the calling song of E. roberti n. sp.is remarkably consistent across its wide distribution. The echeme, syllable sequence and subphrase durations do exhibit some variation, but this variation appears to be present within and among individuals in the broader population, without exhibiting any clear spatial pattern. Much of this variation considered likely to be at least partly temperature dependent.
1502352774
2004-03-11
2004-03-12
2004-03-11
Australia
-28.983334
Glenlyon Dam
1229
151.46666
11
412
1
1
Texas
holotype
1502352772
1990-02-28
L. & W. Popple. & Brisbane Q.
Australia
Indooroopilly
11
412
1
1
QUEENSLAND
paratype
1502352768
1990-02-11
1997-12-28
1990-02-11
NSW
A. I. Knight & M Mackellar Ra. Rd & Bungabbeee SF & Watkins
Australia
180
Kyogle
Stanthorpe
11
412
2
2
QUEENSLAND
paratype
1502352785
1972-01-29
1972-01-30
1972-01-29
H. Frauca
Australia
Mt Woowoonga
Dallarnil
Bundaberg
11
412
1
1
QUEENSLAND
paratype
1502352784
1972-01-10
H. Frauca
Australia
Biggenden
11
412
1
1
QUEENSLAND
paratype
1502352765
[395,1133,1430,1455]
1971-01-11
H. Frauca
Australia
518
Bluff Range
Biggenden
11
412
2
2
QUEENSLAND
paratype
1502352799
1972-12-12
H. Frauca
Australia
518
Bluff Range
Biggenden
11
412
1
1
QUEENSLAND
paratype
1502352790
[295,712,1466,1491]
1977-01-13
H. Frauca
Australia
518
Bluff Range
Biggenden
11
412
1
1
QUEENSLAND
paratype
1502352797
1972-12-09
H. Frauca
Australia
Biggenden
South
Bluff Range
11
412
1
1
QUEENSLAND
paratype
1502352802
[242,840,1502,1527]
1972-12-19
1972-12-20
1972-12-19
H. Frauca
Australia
Biggenden
South
Bluff Range
11
412
2
2
QUEENSLAND
paratype
1502352810
1972-01-03
H. Frauca
Australia
671
Biggenden
Mt Walsh
11
412
3
1
2
QUEENSLAND
paratype
1502352806
[387,1155,1538,1563]
1976-12-04
Biggenden Q. & H. Frauca
Australia
Coast
11
412
1
1
Coast
paratype
1502352808
1976-12-23
H. Frauca
Australia
Bundaberg
Spring Ck
11
412
1
1
Coast
paratype
1502352796
1985-12-19
ANIC
Toowoomba Q. & I. F. B. Common
Australia
620
-27.33
Prince Henry Heights
742
151.59
11
412
1
1
Coast
paratype
1502352807
1997-12-21
C. J. Burwell
Australia
Qld
340
Binjour Plateau
151.469
Sx
11
412
2
2
Coast
paratype
1502352803
1998-12-18
1998-12-19
1998-12-18
G. Monteith & C. Gough & G. Maywald
Australia
A. Ewart
-25.7
Site
1245
151.43333
11
412
2
1
1
Coast
paratype
1502352789
1908-10-26
Hamlyn-Harris
Australia
Toowoomba
11
412
1
1
Coast
paratype
1502352791
[151,1390,1754,1779]
2001-12-17
2001-12-18
2001-12-17
G. Monteith & S. Wright
Australia
330
Barakula
150.51666
Sx
11
412
5
2
3
Coast
paratype
1502352798
2001-12-26
2001-12-29
2001-12-26
G. Montieth
Australia
Glen Witheren' General
153.11667
Sx
11
412
1
1
Coast
paratype
1502352811
1997-12-20
C. J. Burwell & S. Evans
Australia
460
Hurdle Gully
150.98334
11
412
1
1
Coast
paratype
1502352793
1990-11
C. J. Burwell
Australia
A. Ewart
Samford
11
412
1
1
Coast
paratype
1502352809
2003-01-24
L. & W. Popple
Australia
Binjour
11
412
1
1
Queensland
paratype
1502352787
2001-12-11
Hand & L. W. Popple
Australia
-28.161388
Thane
20
151.96638
11
412
3
1
2
Queensland
paratype
1502352801
2001-12-11
N. Warwick & L. Popple & J. Moss
Australia
Willowvale
11
412
1
1
Queensland
paratype
1502352792
Warwick, L. & Popple, J. & Genitalia
Australia
12
413
-28.162
Thanes Creek
55
151.6287
11
412
1
1
Queensland
paratype
1502352795
1998-03-06
L. Popple
Australia
Ipswich
Dinmore
12
413
1
1
Queensland
paratype
1502352800
[534,1387,188,213]
1998-03-03
L. Popple
Australia
Brisbane
Camp Hill
12
413
1
1
Queensland
paratype
1502352794
2001-12-27
2001-12-28
2001-12-27
N. Glenmorgan & L. Popple & A. Strange
Australia
Myall Park
12
413
1
1
Queensland
paratype
1502352804
2001-12-26
L. Popple & A. Strange
Australia
Crows Nest
Austin Rd
12
413
1
1
Queensland
paratype
1502352805
2001-12-26
L. Popple & A. Strange
Australia
Toowoomba
Highfields
12
413
1
1
Queensland
paratype
1502352788
[667,1390,295,320]
2000-12-10
L. Popple & S. Billington
Australia
Lytton
12
413
1
1
Queensland
paratype
1502352822
1998-11-07
L. Popple
Australia
Heights
12
413
3
3
Queensland
paratype
1502352814
1999-12-03
W. Popple
Australia
Heights
12
413
1
1
Queensland
paratype
1502352823
2003-01-07
2003-01-08
2003-01-07
L. W. Popple & Recorded L. W.
Australia
Brisbane
Heights
12
413
1
1
Queensland
paratype
1502352836
[365,1390,404,429]
2003-01
L. W. Popple
Australia
Brisbane
Heights
12
413
1
1
Queensland
paratype
1502352826
2002-12-21
R. J. Popple
Australia
Brisbane
Heights
12
413
1
1
Queensland
paratype
1502352835
2004-01-11
M. Larsen
Australia
-27.3
Mount Glorious
1237
152.46666
Qld
12
413
1
1
paratype
1502352828
Afterlee Public School & L. Popple & R. MacSloy
Australia
-28.566668
Kyogle district
1231
152.81667
12
413
1
1
Queensland
paratype
1502352820
2004-12-28
2004-12-29
2004-12-28
L. W. Popple
Australia
-25.378334
Burnett River
21
151.12944
Eidsvold
12
413
1
1
Queensland
paratype
1502352833
2005-01-28
L. Popple & R. MacSloy
Australia
-28.162
Warwick
55
151.6287
Thanes Creek
12
413
1
1
Queensland
paratype
1502352832
2005-02-19
L. W. Popple & R. MacSloy
Australia
-27.413055
Seventeen Mile Road
Helidon Hills
12
413
1
1
Queensland
paratype
1502352816
2005-10-28
L. W. Popple
Australia
-25.605
Mt Debatable
21
151.54056
12
413
1
1
Queensland
paratype
1502352813
2005-12-05
2005-12-10
2005-12-05
Southwood NP & L. Popple & A. Ewart
Australia
-27.834723
Moonie
20
150.11307
12
413
1
1
Queensland
paratype
1502352821
2005-12-14
2005-12-15
2005-12-14
L. W. Popple
Australia
-27.350555
Dalby
20
151.09277
Lake Broadwater
12
413
4
3
1
Queensland
paratype
1502352830
2008-03
Australia
-27.449444
Murrarie
20
153.09833
Metroplex
12
413
1
1
Queensland
paratype
1502352817
2007-12-11
Burrandoan Rd, SW
Australia
-26.598333
Kingaroy
20
151.74945
12
413
1
1
Queensland
paratype
1502352827
2007-12-10
A.F. Sanborn
Australia
-26.855278
Beerwah
20
153.00667
Roys Rd
12
413
1
1
paratype
1502352812
2010-01-22
L. W. Popple
Australia
-28.225279
Australia
20
152.01999
12
413
1
1
paratype
1502352829
[718,1287,943,968]
1975-11-06
A. Ewart
Australia
Crows Nest
Iron Bark
12
413
1
1
paratype
1502352815
1975-11-22
A. Ewart
Australia
Crows Nest
Iron Bark
12
413
3
1
2
paratype
1502352825
[634,1150,980,1005]
1975-10-19
A. Ewart
Australia
Perseverance Dam
12
413
2
2
paratype
1502352824
1977-11-01
A. Ewart
Australia
Crows Nest
Ironbark Qld
12
413
1
1
paratype
1502352818
[494,1151,1015,1040]
1984-02-12
L. Broadwater & A. Ewart
Australia
Qld
Dalby
12
413
1
1
paratype
1502352831
1987-02-21
1987-10
1987-02-21
Wilkie Ck, L. & M Mt Nebo & H. Horton
Australia
Broadwater area
12
413
4
1
3
paratype
1502352834
1983-01-29
Australia
-26.916666
Iron Bark Ridge
1239
150.41667
Rockwood' Chinchilla
12
413
1
1
paratype
1502352819
[498,1281,1124,1149]
1990-11-03
L. Broadwater
Australia
Dalby
12
413
1
1
paratype
1502352848
1997-12-15
Baracula S. F.
Australia
Hellhole Creek Quarry
Auburn Rd
Hellhole Creek
12
413
1
1
paratype
1502352843
2000-12-22
Gayndah, S. E. Q. & Field
Australia
-25.533167
Binjour Plateau
12
151.5
12
413
1
1
paratype
1502352846
2001-11-30
Recorded
Australia
-28.238667
Warwick
12
151.92566
Leslie Dam
12
413
1
1
paratype
1502352859
2001-12-01
Open
Australia
-28.252167
Warwick
12
151.92783
Leslie Dam
12
413
1
1
paratype
1502352851
2002-01-09
N. Chinchilla & Brigalow, A. E
Australia
-26.663166
Allinga'
12
150.63434
12
413
1
1
paratype
1502352852
[674,1174,1412,1437]
1999-12-16
M. Coombs
Australia
Anstead Qld
12
413
1
1
paratype
1502352856
1999-10-31
G. Maywald
Australia
-26.216667
Glastonbury Qld
1243
152.53334
12
413
1
1
paratype
1502352842
[687,1209,1447,1472]
2001-12-11
M. Coombs
Australia
Eumundi Qld
12
413
1
1
paratype
1502352861
2001-12-14
M. Coombs
Australia
Gunalda Qld
12
413
1
1
paratype
1502352853
[560,1121,1484,1509]
1993-11-07
R. Eastwood
Australia
Blackbutt
Benarkin
12
413
1
1
paratype
1502352847
2004-01-02
Cooley & Hill & Marshall & Moulds
Australia
223
-28.3
Boggabilla
123
150.295
Yarrill Ck
12
413
1
1
paratype
1502352839
1987-12-19
T. A. Moulds
Australia
Miles
12
413
1
1
paratype
1502352845
[451,1230,1556,1581]
1976-12-19
M. S. & B. J. Moulds
Australia
Goomeri
Kinbombi Falls
12
413
2
2
paratype
1502352860
1979-12-17
Jimna, N. & A. Hiller
Australia
Kilcoy
12
413
4
2
2
paratype
1502352841
[527,973,1591,1616]
1989-10-22
R. Eastwood
Iceland
Isla
12
413
1
1
paratype
1502352855
1975-01-06
M. S. & B. J. Moulds
Iceland
Monto
Coominglah Range
12
413
3
2
1
paratype
1502352844
[849,1388,1628,1653]
1980-08-10
I. Gynther
Iceland
Brisbane
St Lucia
12
413
1
1
paratype
1502352854
1984-10-26
J. North
Iceland
Kenmore
12
413
1
1
paratype
1502352840
[259,643,1700,1725]
1983-09-24
J. North
Iceland
St Lucia
12
413
1
1
paratype
1502352850
[655,1265,1700,1725]
1987-10-10
R. Van Barneveld
Iceland
Brisbane
St Lucia
12
413
1
1
paratype
1502352849
1983-10-05
J. T. North
Iceland
Brisbane
Ashgrove
12
413
1
1
paratype
1502352837
[517,1090,1735,1760]
1993-01-01
R. Eastwood
Iceland
Warwick
Leslie Dam
12
413
4
4
paratype
1502352857
1990-01-07
R. Eastwood
Iceland
Warwick
Leslie Dam
12
413
1
1
paratype
1502352858
[372,1388,1772,1797]
1986-10-25
G. and A. Daniels
Iceland
-27.35
Dalby
1237
151.1
Lake Broadwater
12
413
1
1
paratype
1502352838
1986-12-26
G. and A. Daniels
Iceland
-27.35
Dalby
1237
151.1
Lake Broadwater
12
413
1
1
paratype
1502352874
[579,1104,1807,1832]
1987-01-30
G. and A. Daniels
Iceland
-27.35
Dalby
1237
151.1
Lake Broadwater
12
413
1
1
paratype
1502352869
1987-02-30
G. and A. Daniels
Iceland
-27.35
Dalby
1237
151.1
Lake Broadwater
12
413
1
1
paratype
1502352870
[279,699,1844,1869]
1983-12-09
C. E. Hagan
Iceland
St Lucia
12
413
1
1
paratype
1502352884
[710,1325,1844,1869]
1993-01-13
R. Eastwood
Iceland
Warwick
Canningvale
12
413
1
1
paratype
1502352879
1993-02-06
R. Eastwood
Iceland
Warwick
Canningvale
12
413
1
1
paratype
1502352878
2002-01-10
Cooley & Hill & Marshall & Cowan & Moulds
Iceland
357
-27.3141
Lake Broadwater
1
151.09937
12
413
1
1
paratype
1502352876
2002-01-09
Cooley & Cowan & Hill & Marshall
Iceland
-26.889
Blackbutt
12
152.21466
12
413
3
1
2
paratype
1502352866
2009-01-04
SUN
Sundown N. P. & Hill & Marshall & Moulds & Owen
Iceland
385
-28.919733
Texas
81
151.58167
12
413
1
1
Texas
paratype
1502352897
2009-01-03
Hill & Marshall & Moulds & Owen
Iceland
347
13
414
-28.672817
Inglewood
1
151.1493
12
413
1
1
Texas
paratype
1502352880
[151,686,188,213]
1993-01-14
R. Eastwood
Iceland
Warwick
Wildash
13
414
3
3
Texas
paratype
1502352867
[699,1124,188,213]
1995-02-20
R. Eastwood
Iceland
Warwick
Wildash
13
414
1
1
Texas
paratype
1502352871
1995-02-22
R. Eastwood
Iceland
Warwick
Wildash
13
414
1
1
Texas
paratype
1502352865
[284,727,223,248]
1994-12-09
R. Eastwood
Iceland
Warwick
Wildash
13
414
1
1
Texas
paratype
1502352883
[743,1195,223,248]
1989-01-21
R. Eastwood
Iceland
Blackbutt
13
414
1
1
Texas
paratype
1502352873
1990-01-10
R. Eastwood
Iceland
Warwick
Connolly Dam
13
414
1
1
Texas
paratype
1502352877
1998-02-29
G. Daniels
Iceland
-27.5
Brisbane
1236
153.1
St Lucia
13
414
1
1
Queensland
paratype
1502352868
2002-01-08
MSM
Cooley & Cowan & Hill & Marshall & Moss & Moulds
Iceland
Brisbane
Capalaba
13
414
1
1
NEW SOUTH WALES
paratype
1502352881
[361,1389,332,357]
2003-11-01
L. Popple & R. MacSloy
Australia
Legume
Oaky Hill
13
414
1
1
paratype
1502352863
2016-01-04
NSW
L. W. Popple & Recorded
Australia
-29.35249
Stannum
1
151.80315
13
414
1
1
paratype
1502352875
1992-01-16
1995-02-04
1992-01-16
NSW
M. Coombs & M Inverell
Australia
Tamworth
13
414
3
3
paratype
1502352872
[227,1254,439,464]
2015-02-16
N. & D. Emery
Australia
420
-31.487
Warrumbungle
12
149.01266
13
414
5
1
4
paratype
1502352862
2014-01-12
C. & D. Emery
Australia
680
-31.302334
Warrumbungle
12
149.03416
Whitegum Lookout
13
414
1
1
paratype
1502352888
Australia
-31.651833
Spring Ridge
13
414
1
1
paratype
1502352882
2013-01-04
N., C. & D. Emery
Australia
654
-30.068
Ebor
12
152.63467
13
414
1
1
paratype
1502352864
2013-12-02
N., C. & D. Emery
Australia
600
-30.135334
Bingara
12
150.07217
Killarney Gap
13
414
3
1
2
paratype
1502352896
[254,1053,620,645]
2013-12-02
NSW
N., C. & D. Emery
Australia
600
-30.135334
Bingara
12
150.07217
Killarney Gap
13
414
1
1
paratype
1502352893
2013-12-02
SIMON, LAB
N., C. & D. Emery
Australia
600
-30.135334
Bingara
12
150.07217
Killarney Gap
13
414
LAB 05
1
1
paratype
1502352891
2005-01-01
NSW
Warialda, N. S. W & Hill & Marshall & Moulds
Australia
591
-29.551266
Warialda
1
150.58772
13
414
1
paratype
1502352920
1974-01-01
M. S. & B. J. Moulds
Australia
Baradine
13
414
6
6
paratype
1502352901
[610,1290,799,824]
1975-01-10
M. S. & B. J. Moulds
Australia
Kyogle
Afterlee
13
414
5
3
2
paratype
1502352900
1985-01-02
M. S. & B. J. Moulds
Australia
Inverell
13
414
1
1
paratype
1502352903
[498,956,836,861]
1985-01-07
MSM
W. Rixon
Australia
Kyogle
13
414
1
1
paratype