Ledocarpon meyenianum (Klotzsch) Steudel (1840: 20) Balbisia meyeniana Balbisia meyeniana The genus Balbisia (Vivianiaceae, Geraniales) in Peru, Bolivia and northern Chile Weigend, Maximilian Phytotaxa 2011 2011-04-11 22 1 47 56 5WCRR Klotzsch (1836: 432) Klotzsch 432 1836 [183,730,1915,1944] Magnoliopsida Vivianiaceae Balbisia Plantae Geraniales 2 49 Tracheophyta species meyeniana    Type: [ PERU. Depto. Tacna:], Cordillerabei Tacna,  Meyens.n. anno 1831( B†), Laguna de Titicaca,  v. Besser54( B†).— Neotype(here designated): PERU. Depto Tacna: Prov. Tarata, between Tacnaand Tarata on Wside of pass, 1 km  below Palquilla,  3370 m,  28 November 1964,  Hutchison& Wright7190( USM!, isoneotypes F!, MO, NY, UC, US).   Ξ   Ledocarpon meyenianum(Klotzsch) Steudel (1840: 20).   Balbisia meyenianais here reported from Peruand Bolivia. In Peruthe species appears to be restricted to the two southwestern Departments Moqueguaand Tacnaand has not so far been found further north, in spite of copious collections now available from Arequipa. The leaves are glabrous to puberulent and glaucous in the living state ( Fig. 1 D) and lack both dense, sericeous pubescence and a revolute leaf margin (present in all the other species here treated).  Balbisia meyenianaoften has a distinctly red calyx ( Fig. 1 F) and a very sparsely branched, glabrous or puberulent epicalyx ( Figs. 1 F, 2 F).  The northernmost Peruvian collection ( Weigend & Baldárrago 9366, Fig. 1 B )was found only a few kilometers south of the village of Puquina while just north of that village is the southernmost population of  B. verticillata( Weigend & Baldárrago 9367, Fig. 1 A). Which factors maintain this well-defined distribution limit is entirely unclear. The Bolivian specimens seen differ in being slightly more pubescent, especially the trichomes on the epicalyx are long (and equal the diameter of the epicalyx segments). However, they lack the sericeous pubescence and recurved leaf margins typical of  B. verticillataand have to be referred to  B. meyenianafor the time being.    Representative specimens:  PERU. Depto. Moquegua: Prov. Moquegua,  50–57 km NE Moqueguaon road to Carumas,  3000–3200 m,  15 November 1986,  Dillonet al. 4802( F).  Carumas,  2700–2800 m,  21 February– 6 March 1925,  Weberbauer7271( F).  Prov. General Sánchez Cerro, between Puquinaand Omate, S 16° 37´42.2´´, W 071° 04´17.6´´,  3003 m,  20 April 2004,  Weigendet al. 8326( B, F, HUT, M, USM).  Prov. General Sánchez Cerro, Puquina, a few kilometers before Puquinaon road from Omate, S 16° 38´21.4´, W 071° 09´21.4´´,  3142 m,  21 July 2010,  Weigend& Baldárrago9366( B, F, HUSA, M, USM).  Roadfrom Arequipato Omate, just above Omate, S 16° 37´29.0´´, W 71° 03´48.4´´,  2800 m,  7 December 2006,  Ackermann& Cáceres H. 658( B, HUSA, M, USM).  Roadfrom Omateto Carabaya, above Carabaya, S 16° 37' 51.1'' W 070° 57' 59.3'',  2799 m,  Weigend& Ackermann9266( B, HUSA, M, USM).  Depto. Tacna: Prov. Tarata, Distr. Ticaco,  3069 m,  12 November 2006, Cáceres H . &   Fernández5860( B, HUSA). Prov. Tacna, 67 kmtowards Torata,  3250 m,  26 June 1979, Beck 2030( LPB, M).   BOLIVIA: Depto. Potosí: Licori,  3300 m,  1 November 1966,  Guzmáns.n.( LPB).  Prov. Sud Chinchas, Quebrada Chiriyoj Waykhow Tupiza, S 21° 26´, W 065° 47´,  3400 m,  15 February 2002,  Müller9008( LPB).  Licori,  3300 m,  1 November 1966,  Guzmáns.n.( LPB).  Prov. Sud Chinchas, road from Tupizavia Quebrada Palalato El Sillar,  3200 m,  29 March 1997,  Wood11953( LPB).  Salasala–Tasnapalca[or Tamapalca],  3200 m,  12 May 1927,  Trolls.n.( B).  Depto. Tarija: Prov. José Maria Aviléz, Higuerasto Villazón, S 21° 54´, W 065° 27´,  3200 m,  9 March 1998, Beck et al. 23835( LPB).  Jaipa,  3400 m,  19 April 1904, Fiebrig 3094( B, 3 x F, K). B, W Type & Depto. & Cordillera & Meyen & v. Besser Peru Laguna de Titicaca 2 49 1 Tacna neotype [199,1432,151,175] 1964-11-28 USM, F, MO, NY, UC Hutchison & Wright United States of America 3370 US 3 50 1 1986-11-15 NE, F Dillon Peru Depto. 3100 Carumas Prov. 3 50 1 Moquegua 1925-02-21 1925-03-06 1925-02-21 F Weberbauer Peru 2750 Carumas 3 50 1 2004-04-20 B, F, HUT, M, USM Weigend Peru Prov. General Sanchez Cerro 3003 -16.62839 Omate 1 -71.071556 Puquina 3 50 1 2010-07-21 B, F, HUSA, M, USM Weigend & Baldarrago Peru Prov. General Sanchez Cerro 3142 -16.639278 Puquina 1 -71.155945 Puquina 3 50 1 2006-12-07 H, B, HUSA, M, USM Ackermann & Caceres Peru Road 2800 -16.624722 Omate 1 -71.063446 Omate 3 50 1 Arequipa B, HUSA, M, USM Weigend & Ackermann Peru Road 2799 -16.630861 Carabaya 1 -70.96647 Omate 3 50 1 [198,1160,1105,1131] 2006-11-12 H 3069 Tacna 3 50 1 Tacna 1979-06-26 B, HUSA, LPB, M 3250 Torata Fernandez 3 50 1 Tacna [198,1181,1182,1208] 1966-11-01 LPB Guzman Bolivia 3300 Licori Depto. 3 50 1 Potosi 2002-02-15 LPB Muller Bolivia Prov. Sud Chinchas 3400 -21.433332 Tupiza 1264 -65.78333 Quebrada Chiriyoj Waykho 3 50 1 [151,804,1260,1286] 1966-11-01 LPB Guzman Bolivia 3300 Licori 3 50 1 1997-03-29 LPB El Sillar & Wood Bolivia Prov. Sud Chinchas 3200 Quebrada Palala Tupiza 3 50 1 1927-05-12 B Troll Bolivia 3200 Salasala - Tasnapalca 3 50 1 1998-03-09 LPB 3200 -21.9 Villazon 1262 -65.45 Higueras 3 50 1 Tarija [523,1191,1414,1440] 1904-04-19 B, F, K 3400 Jaipa 3 50 1 Tarija