Urban Scatopsidae (Diptera) of Los Angeles, California, United States Amorim, Dalton de Souza Brown, Brian V. Insect Systematics and Diversity 2020 2020-01-31 4 1 1 1 41 WK54 (Meigen) Meigen 1830 [818,1127,541,564] Insecta Scatopsidae Coboldia Animalia Diptera 23 24 Arthropoda species fuscipes  ( Figs. 7, 8, 19, 20, 62, and 66)   Coboldia fuscipes( Meigen, 1830): 314 (  Scatopse). Typelocality: ‘Europe’. Ref.–Hardy, 1956: 99 (as  Scatopse); Cook, 1957: figs. 3A–B (male terminalia), 3D (spiracular sclerite), 3N (female sternite 7), 5D (female terminalia), 5F (male segment 7), 6B (wing), 6G (male tergite 8); Cook, 1963: 17, Figs. 47(female terminalia), 48 (male segment 7), 49 (male terminalia), 59 (wing); Rabello and Forattini, 1962: 303(as  Rhexoza coutinhoi) (immature, biology); Cook, 1974: 70, Figs. 78(male terminalia); 79 (female terminalia); Cook, 1967(Neotropical catalog); Haenni and Vaillant, 1994(biology). Type, MNHNP ( Duda 1928a: 22).   Diagnosis.Orange-brown. Male tergite 8 with a medial elongate, asymmetrical spatulate projection, aedeagus very long, coiled; female sternite 7 with a medial bare emargination on posterior margin.   Material Examined.  1 ♂, United States, California, Los Angeles Co., Los Angeles, University Park, 34.034°N 118.281°W, BioSCAN site 3,  30 April–7 May 2014, Malaise trap,  63 m, Coll. Peggy Hentschke, BioSCAN sample 16068 ( LACM);  2 ♂♂, 1 ♀, United States, California, Los Angeles Co., Los Angeles, Silverlake, 34.093°N 118.274°W, BioSCAN site 5,  29 March–5 April 2014, Malaise trap,  166.1 m, Coll. Walter Renwick, BioSCAN sample 15916 ( LACM);  1 ♂, same data, but  1–8 February 2014, BioSCAN sample 15589 ( LACM);  3 ♂♂, 1 ♀, same data, but  1–8 March 2014, BioSCAN sample 15735 ( LACM);  1 ♂, 1 ♀, United States, California, Los Angeles Co., Los Angeles, Korea Town, 34.072°N 118.291°W, BioSCAN site 11,  5–12 April 2014, Malaise trap,  85.6 m, Coll. Peter Ralph, BioSCAN sample 16056 ( LACM);  1 ♀, same data, but  2–9 March 2014, BioSCAN sample 15796 ( LACM);  1 ♂, same data, but BioSCAN sample 16482 ( LACM);  1 ♂, United States, California, Los Angeles Co., Glendale, 34.149°N 118.218°W, BioSCAN site 13,  29 November–6 December 2014, Malaise trap,  245.4 m, Coll. Karen Hoffman, BioSCAN sample 16893 ( LACM);  1 ♀, United States, California, Los Angeles Co., Los Angeles, Hollywood, 34.095°N 118.334°W, BioSCAN site 16,  2–9 April 2014, Malaise trap,  100 m, Coll. Tony Hein, BioSCAN sample 15980 ( LACM);  3 ♀♀, same data, but BioSCAN sample 16345 ( LACM);  1 ♀, same data, but  1–18 January 2014, BioSCAN sample 15608 (on slide) ( LACM);  1 ♀, same data, but BioSCAN sample 16341 ( LACM);  1 ♂, United States, California, Los Angeles Co., Glendale, 34.159°N 118.247°W, BioSCAN site 17,  29 March–5 April 2014, Malaise trap,  190 m, Coll. Celeste Armstrong, BioSCAN sample 15911 ( LACM);  1 ♂, same data, but  3–10 May 2014, BioSCAN sample 16013 ( LACM);  1 ♂, United States, California, Los Angeles Co., Los Angeles, Carthay, 34.059°N 118.369°W, BioSCAN site 19,  2–9 December 2014, Malaise trap,  41.1 m, Coll. Teresa Dahl, BioSCAN sample 16876 ( LACM);  7 ♀♀, United States, California, Los Angeles Co., Burbank, 34.17°N 118.308°W, BioSCAN site 21,  1–8 February 2014, Malaise trap,  158 m, Coll. Chris Cianci, BioSCAN sample 15664 ( LACM);  1 ♀, same data, but  25 January–1 February 2014, BioSCAN sample 15554 ( LACM);  1 ♂, same data, but  1–8 March 2014, BioSCAN sample 15668 ( LACM);  1 ♂, United States, California, Los Angeles Co., Los Angeles, Echo Park, 34.074°N 118.264°W, BioSCAN site 22,  21–28 January 2014, Malaise trap,  133.5 m, Coll. Erin Johnson, BioSCAN sample 15576 ( LACM);  1 ♀, United States, California, Los Angeles Co., Los Angeles, Atwater Village, 34.114°N 118.251°W, BioSCAN site 23,  2–10 April 2014, Malaise trap,  119 m, Coll. Sidney Higgins, BioSCAN sample 15858 ( LACM);  1 ♂, same data, but  26 November–10 December 2014, BioSCAN sample 16887 ( LACM);  1 ♂, United States, California, Los Angeles Co., Los Angeles, Eagle Rock, 34.136°N 118.194°W, BioSCAN site 24,  1–8 February 2014, Malaise trap,  215 m, Coll. Jim Hogue, BioSCAN sample 15651 ( LACM);  2 ♂♂, 3 ♀♀, same data, but  1–8 March 2014, BioSCAN sample 15842 ( LACM);  1 ♀, United States, California, Los Angeles Co., Los Angeles, Eagle Rock, 34.136°N 118.194°W, BioSCAN site 24,  3–10 May 2014, Malaise trap,  215 m, Coll. Jim Hogue, BioSCAN sample 16018 ( LACM);  1 ♂, United States, California, Los Angeles Co., Los Angeles, Pico–Union, 34.046°N 118.275°W, BioSCAN site 25,  1–8 February 2014, Malaise trap,  70 m, Coll. Humberto Marquez, BioSCAN sample 15689 ( LACM);  1 ♀, same data, but  28 December 2013–  4 January 2014, BioSCAN sample 15505 ( LACM);  1 ♀, United States, California, Los Angeles Co., Los Angeles, Highland Park, 34.123°N 118.193°W, BioSCAN site 29,  1–8 February 2014, Malaise trap,  205.6 m, Coll. Natalie Brejcha, BioSCAN sample 15703 ( LACM);  1 ♀, United States, California, Los Angeles Co., Los Angeles, Highland Park, 34.125°N 118.189°W, BioSCAN site 30, Malaise trap,  227.8 m, Coll. Candace Franco, BioSCAN sample 16235 ( LACM);  1 ♀, United States, California, Los Angeles Co., Los Angeles, Rosemead, 34.05°N 118.108°W, BioSCAN site 34,  1–8 April 2016, Malaise trap,  135 m, Coll. Deniece Lascano, BioSCAN sample 18230 ( LACM);  1 ♂, United States, California, Los Angeles Co., Monrovia. 34.158°N 117.999°W, BioSCAN site 35,  1–15 June 2016, Malaise trap,  214 m, Coll. Brian Brown, BioSCAN sample 18277 ( LACM);  1 ♂, 3 ♀♀, same data, but  1–8 May 2016, BioSCAN sample 18212 ( LACM);  1 ♀, United States, California, Los Angeles Co., Claremont, 34.09°N 117.711°W, BioSCAN site 37,  1–8 June 2016, Malaise trap,  341 m, Coll. Kathryn Turner, BioSCAN sample 18337 ( LACM);  1 ♂, same data, but  1–8 March 2016, BioSCAN sample 18164 ( LACM);  1 ♂, same data, but BioSCAN sample 18250 ( LACM);  1 ♂, same data, but  1–8 September 2016, BioSCAN sample 18399 ( LACM);  1 ♂, United States, California, Los Angeles Co.,  El Segundo. 33.925°N 118.425°W, BioSCAN site 40,  1–8 May 2016, Malaise trap,  35 m, Coll. Jack Fulmer, BioSCAN sample 18240 ( LACM);  1 ♀, United States, California, Los Angeles Co.,  La Mirada, 33.914°N 118.005°W, BioSCAN site 43,  1–8 June 2016, Malaise trap,  63 m, Coll. Carmen Gendusa, BioSCAN sample 18321 ( LACM);  1 ♀, same data, but  1–8 April 2016, BioSCAN sample 18265 ( LACM);  2 ♂♂, 2 ♀♀, same data, but  1–8 May 2016, BioSCAN sample 18266 ( LACM);  1 ♀, United States, California, Riverside Co., Norco. 33.912°N 117.57°W, BioSCAN site 46,  1–8 March 2016, Malaise trap,  180 m, Coll. Sara Taylor, BioSCAN sample 18177 ( LACM);  1 ♀, United States, California, Riverside Co., Riverside, 33.984°N 117.313°W, BioSCAN site 47,  1–8 March 2016, Malaise trap,  375 m, Coll. Justin Betz, BioSCAN sample 18179 ( LACM).   Distribution.Cosmopolitan.   Remarks.  Coboldia fuscipesis the most widespread scatopsid species known. Its larvae have been found in very different kinds of environments: there are records of larvae connected to a variety of decaying organic matter of vegetal and animal origin ( Haenni and Vaillant, 1994). The larvae, pupae, and biology were described in detail by Rabello and Forattini (1962). Cook (1975)refers to adults of this species reared from  Echinocactus polycephalus.   Coboldia fuscipesshowed up in the samples of 21 out of the 47 BioSCAN collecting sites. There is an obvious peak of emergence in mid-spring, in April and May, with some additional catches in late autumn. The Bold Systems database has 52 sequenced specimens of  C. fuscipes, from North America and Europe, all belonging to a single cluster (http://boldsystems.org/index.php/ Public_SearchTerms?query=%22 Coboldia%20fuscipes%22[tax]). 2618789725 2014-04-30 2014-05-07 2014-04-30 LACM Coll. Peggy Hentschke United States of America Los Angeles Co. 63 34.034 University Park 72 -118.281 Los Angeles 23 24 1 1 California 2618789325 2014-03-29 2014-04-05 2014-03-29 LACM Coll. Walter Renwick United States of America Los Angeles Co. 166 34.093 Silverlake 72 -118.274 Los Angeles 23 24 3 1 2 California 2618789521 2014-02-01 2014-02-08 2014-02-01 LACM Coll. Walter Renwick United States of America Los Angeles Co. 166 34.093 Silverlake 72 -118.274 Los Angeles 23 24 1 1 California 2618789666 2014-03-01 2014-03-08 2014-03-01 LACM Coll. Walter Renwick United States of America Los Angeles Co. 166 34.093 Silverlake 72 -118.274 Los Angeles 23 24 4 1 3 California 2618789387 2014-04-05 2014-04-12 2014-04-05 LACM Coll. Peter Ralph United States of America Los Angeles Co. 86 34.072 Town 72 -118.291 Los Angeles 23 24 2 1 1 California 2618789532 2014-03-02 2014-03-09 2014-03-02 LACM Coll. Peter Ralph United States of America Los Angeles Co. 86 34.072 Town 72 -118.291 Los Angeles 23 24 1 1 California 2618789593 2014-03-02 2014-03-09 2014-03-02 LACM Coll. Peter Ralph United States of America Los Angeles Co. 86 34.072 Town 72 -118.291 Los Angeles 23 24 1 1 California 2618789678 2014-11-29 2014-12-06 2014-11-29 LACM Coll. Karen Hoffman United States of America Los Angeles Co. 245 34.149 Glendale 72 -118.218 23 24 1 1 California 2618789705 2014-04-02 2014-04-09 2014-04-02 LACM Coll. Tony Hein United States of America Los Angeles Co. 100 34.095 Hollywood 72 -118.334 Los Angeles 23 24 1 1 California 2618789562 2014-04-02 2014-04-09 2014-04-02 LACM Coll. Tony Hein United States of America Los Angeles Co. 100 34.095 Hollywood 72 -118.334 Los Angeles 23 24 3 3 California 2618789491 2014-01-01 2014-01-18 2014-01-01 LACM Coll. Tony Hein United States of America Los Angeles Co. 100 34.095 Hollywood 72 -118.334 Los Angeles 23 24 1 1 California 2618789470 [818,1323,1760,1780] 2014-01-01 2014-01-18 2014-01-01 LACM Coll. Tony Hein United States of America Los Angeles Co. 100 34.095 Hollywood 72 -118.334 Los Angeles 23 24 1 1 California 2618789629 2014-03-29 2014-04-05 2014-03-29 LACM Coll. Celeste Armstrong United States of America Los Angeles Co. 190 34.159 Glendale 72 -118.247 23 24 1 1 California 2618789672 2014-05-03 2014-05-10 2014-05-03 LACM Coll. Celeste Armstrong United States of America Los Angeles Co. 190 34.159 Glendale 72 -118.247 23 24 1 1 California 2618789591 2014-12-02 2014-12-09 2014-12-02 LACM Coll. Teresa Dahl United States of America Los Angeles Co. 41 24 25 34.059 Carthay 72 -118.369 Los Angeles 23 24 1 1 California 2618789477 2014-02-01 2014-02-08 2014-02-01 LACM Coll. Chris Cianci United States of America Los Angeles Co. 158 34.17 Burbank 557 -118.308 24 25 7 7 California 2618789499 2014-01-25 2014-02-01 2014-01-25 LACM Coll. Chris Cianci United States of America Los Angeles Co. 158 34.17 Burbank 557 -118.308 24 25 1 1 California 2618789549 2014-03-01 2014-03-08 2014-03-01 LACM Coll. Chris Cianci United States of America Los Angeles Co. 158 34.17 Burbank 557 -118.308 24 25 1 1 California 2618789407 2014-01-21 2014-01-28 2014-01-21 LACM Coll. Erin Johnson United States of America Los Angeles Co. 134 34.074 Echo Park 72 -118.264 Los Angeles 24 25 1 1 California 2618789379 2014-04-02 2014-04-10 2014-04-02 LACM Coll. Sidney Higgins United States of America Los Angeles Co. 119 34.114 Atwater Village 72 -118.251 Los Angeles 24 25 1 1 California 2618789616 2014-11-26 2014-12-10 2014-11-26 LACM Coll. Sidney Higgins United States of America Los Angeles Co. 119 34.114 Atwater Village 72 -118.251 Los Angeles 24 25 1 1 California 2618789493 2014-02-01 2014-02-08 2014-02-01 LACM Coll. Jim Hogue United States of America Los Angeles Co. 215 34.136 Eagle Rock 72 -118.194 Los Angeles 24 25 1 1 California 2618789495 2014-03-01 2014-03-08 2014-03-01 LACM Coll. Jim Hogue United States of America Los Angeles Co. 215 34.136 Eagle Rock 72 -118.194 Los Angeles 24 25 5 3 2 California 2618789398 2014-05-03 2014-05-10 2014-05-03 LACM Coll. Jim Hogue United States of America Los Angeles Co. 215 34.136 Eagle Rock 72 -118.194 Los Angeles 24 25 1 1 California 2618789498 2014-02-01 2014-02-08 2014-02-01 LACM Coll. Humberto Marquez United States of America Los Angeles Co. 70 34.046 Pico-Union 72 -118.275 Los Angeles 24 25 1 1 California 2618789363 2013-12-28 2014-01-04 2013-12-28 LACM Coll. Humberto Marquez United States of America Los Angeles Co. 70 34.046 Pico-Union 72 -118.275 Los Angeles 24 25 1 1 California 2618789517 2014-02-01 2014-02-08 2014-02-01 LACM Coll. Natalie Brejcha United States of America Los Angeles Co. 206 34.123 Highland Park 72 -118.193 Los Angeles 24 25 1 1 California 2618789601 LACM Coll. Candace Franco United States of America Los Angeles Co. 228 34.125 Highland Park 72 -118.189 Los Angeles 24 25 1 1 California 2618789431 2016-04-01 2016-04-08 2016-04-01 LACM Coll. Deniece Lascano United States of America Los Angeles Co. 135 34.05 Rosemead 557 -118.108 Los Angeles 24 25 1 1 California 2618789461 2016-06-01 2016-06-15 2016-06-01 LACM Coll. Brian Brown United States of America Los Angeles Co. 214 34.158 Monrovia. 72 -117.999 24 25 1 1 California 2618789590 2016-05-01 2016-05-08 2016-05-01 LACM Coll. Brian Brown United States of America Los Angeles Co. 214 34.158 Monrovia. 72 -117.999 24 25 4 3 1 California 2618789356 2016-06-01 2016-06-08 2016-06-01 LACM Coll. Kathryn Turner United States of America Los Angeles Co. 341 34.09 Claremont 557 -117.711 24 25 1 1 California 2618789369 2016-03-01 2016-03-08 2016-03-01 LACM Coll. Kathryn Turner United States of America Los Angeles Co. 341 34.09 Claremont 557 -117.711 24 25 1 1 California 2618789531 2016-03-01 2016-03-08 2016-03-01 LACM Coll. Kathryn Turner United States of America Los Angeles Co. 341 34.09 Claremont 557 -117.711 24 25 1 1 California 2618789511 2016-09-01 2016-09-08 2016-09-01 LACM Coll. Kathryn Turner United States of America Los Angeles Co. 341 34.09 Claremont 557 -117.711 24 25 1 1 California 2618789736 2016-05-01 2016-05-08 2016-05-01 LACM Coll. Jack Fulmer United States of America Los Angeles Co. 35 33.925 El Segundo. 72 -118.425 El Segundo. 24 25 1 1 California 2618789404 2016-06-01 2016-06-08 2016-06-01 LACM Coll. Carmen Gendusa United States of America Los Angeles Co. 63 33.914 La Mirada 72 -118.005 La Mirada 24 25 1 1 California 2618789487 2016-04-01 2016-04-08 2016-04-01 LACM Coll. Carmen Gendusa United States of America Los Angeles Co. 63 33.914 La Mirada 72 -118.005 La Mirada 24 25 1 1 California 2618789604 2016-05-01 2016-05-08 2016-05-01 LACM Coll. Carmen Gendusa United States of America Los Angeles Co. 63 33.914 La Mirada 72 -118.005 La Mirada 24 25 4 2 2 California 2618789367 2016-03-01 2016-03-08 2016-03-01 LACM Coll. Sara Taylor United States of America Riverside Co. 180 33.912 Norco. 464 -117.57 24 25 1 1 California 2618789567 2016-03-01 2016-03-08 2016-03-01 LACM Coll. Justin Betz United States of America Riverside Co. 375 33.984 Riverside 72 -117.313 24 25 1 1 California