Thomisus quinquetuberculatus Taczanowski, 1872: 100 Stephanopis acanthogastra Mello-Leitão, 1929: 53 Stephanopis quinquetuberculata Keyserling 1880: 171 Mello-Leitão 1929: 62 Tobias quinquetuberculatus Simon 1895: 1053 Kryptochroma: a new genus of bark-dwelling crab spiders (Araneae, Thomisidae) Machado, Miguel Viecelli, Rafaela Guzati, Catherine Grismado, Cristian J. Teixeira, Renato A. European Journal of Taxonomy 2021 2021-11-03 778 26 70 98YQ7 (Taczanowski, 1872) Machado & Viecelli & Guzati & Grismado & Teixeira 2021 Taczanowski 1872 [346,1019,512,539] Arachnida Thomisidae Kryptochroma Animalia Araneae 25 51 Arthropoda species quinquetuberculata gen. et comb. nov.   Figs 14, 18, 19C      Thomisus quinquetuberculatus Taczanowski, 1872: 100.     Stephanopis acanthogastra Mello-Leitão, 1929: 53, fig. 135. syn. nov.    Stephanopis quinquetuberculata–  Keyserling 1880: 171, pl. 3 fig. 94. —  Mello-Leitão 1929: 62.  Tobias quinquetuberculatus–  Simon 1895: 1053.    Notes According to Keyserling (1880), the description of the species in his work was based on the holotypeoriginally proposed by Taczanowski (1872), from Cayenne ( French Guiana), which is presumably lost. We had access to a specimen from the Natural History Museum of London (BMNH 1890.7.1.3479) that is labeled as “type” ( Fig. 14G–H); however, it is clearly not the holotypebecause it was collected after the original description. Since this specimen is strikingly similar to the one represented in the original illustrations (where the author highlighted the diagnostic features of the male palpus: sinuous RTA and the tip-narrowed tegulum) and was found near the original locality ( New Granada, currently corresponding to the Colombian territory), we propose it as the neotypefor  K. quinquetuberculata comb. nov.( Fig. 14G–H).    Diagnosis The male of  K. quinquetuberculataresembles that of  K. pentacanthaby its stout opisthosomal projections and by the presence of a dorsal white macula on the opisthosoma ( Fig. 14A). However, it can be recognized by the tegulum in the shape of a water drop, bearing a straight and tip-narrowed embolus instead of a curved termination observed in its closest correlated species ( Figs 14C, E, 19C). Moreover,  K. quinquetuberculatahas a remarkable RTA which is large at the basis, sharpen abruptly at the tip, and sinuous, pointing towards to the dorsum of the cymbium ( Figs 14D, F, 19C).    Material presumably lost    Holotype FRENCH GUIANA• ♂; Cayenne; MZPW.   Material examined    Neotype COLOMBIA• ♂; BMNH 1890.7.1.3479.    Holotypeof  S. acanthogastra syn. nov. BRAZIL• ♂; Bahia; MNHN 17126.  Additional material   FRENCH GUIANA• 1 ♂; Maripasoula, Massif du Mitaraka; 02°17′26″ N, 54°31′18″ W; 2015; Vedelleg.; MCTP 42644•  1 ♂; Maripasoula, Sommet Tabulaire, Mont Itoupe; 03°32′13″ N, 53°34′25″ W;  13 Nov. 2014; Vedeland Lalagueleg.; MCTP 42645.   BRAZIL–  Pernambuco State• 1 ♂, 1 juv.; Recife, Horto Dois Irmãos; 8°00′35″ S, 34°56′51″ W;  13 Sep. 1999; M. Peresleg.; IBSP 38639. –   Bahia State• 1 ♂; Una, Reserva Biológica de Una; 15°10′42″ S, 39°06′13″ W;  15–28 Nov. 2000; A.D. Brescovitleg.; IBSP 46738• 1 ♂; same collection data as for preceding; IBSP 46712• 1 ♂; same collection data as for preceding; IBSP 46714• 1 ♂; same collection data as for preceding; IBSP 46724• 1 ♂; same collection data as for preceding; IBSP 46726• 1 ♂; same collection data as for preceding; IBSP 46740•  1 ♂; Reserva Zoobotânica CEPLAC; 14°46′22″ S, 39°13′13″ W;  8 Dec. 2010; G.H.F. Azevedoand A.J. Santosleg.; UFMG 9484.    Description  Male(IBSP 46712) PROSOMA. Anterior eye row strongly recurved, posterior slightly procurved; carapace predominantly dark-brown with discrete MS; clypeus lighter than the rest of carapace with light spots at its lateral margins ( Fig. 14B). Labium, endites and chelicerae dark-brown. LEGS. Anterior legs (I and II) dark-brown and posterior femora (III and IV) presenting their proximal half yellow. OPISTHOSOMA. Predominantly brown with a dorsal white macula at its median region and a light transversal line connecting the anterolateral projections ( Fig. 14A). MEASUREMENTS. Eye diameters and eye interdistances: AME 0.06, ALE 0.11, PME 0.09, PLE 0.09, AME–AME 0.06, AME–ALE 0.08, PME–PME 0.16, PME–PLE 0.04, MOQ length 0.32, MOQ width 0.35; leg formula: 1-2-4-3: leg I – femur 1.61/ patella 0.88/ tibiae 1.19/ metatarsus 0.88/ tarsus 0.56/ total 5.12; II – 1.45/ 0.72/ 0.96/ 0.72/ 0.48/ 4.33; III – 0.88/ 0.48/ 0.72/ 0.48/ 0.37/ 2.93; IV – 1.01/ 0.48/ 0.80/ 0.48/ 0.40/ 3.17. Total body length 4.39; carapace 2.12 length, 2.07 width; opisthosoma length 2.22; clypeus 0.24 height; sternum 0.85 length, 0.72 width; endites 0.32 length, 0.25 width; labium 0.16 length, 0.29 width.  Female Unknown.    Distribution  French Guiana(Cayenne and Maripasoula) and Brazil( Bahiaand Pernambuco) ( Fig. 18). 3400646489 MZPW French Guiana Cayenne 25 51 1 1 holotype 3400646500 BMNH Colombia COLOMBIA 25 51 BMNH 1890.7.1.3479 1 1 neotype 3400646405 MNHN Brazil Bahia 25 51 MNHN 17126 1 1 Bahia holotype 3400646502 MCTP Vedel French Guiana 2.2905555 Maripasoula 21 -54.521667 25 51 MCTP 42644 1 1 3400646318 2014-11-13 MCTP Vedel & Lalague French Guiana 3.5369444 Maripasoula 21 -53.57361 25 51 MCTP 42645 1 1 3400646446 1999-09-13 2000-11-28 1999-09-13 BMNH M. Peres & A. D. Brescovit & Fig. & Male & A - F. Specimen & A. Habitus & B. Front. & C - F. Left & C, E. Ventral & D, F. Retrolateral Brazil Recife -15.178333 Reserva Biologica de Una 21 -39.10361 Horto Dois Irmaos 26 52 IBSP 38639, IBSP 46738, IBSP 46712, IBSP 46714, IBSP 46724, IBSP 46726 8 7 Pernambuco neotype 3400646505 1999-09-13 2000-11-28 1999-09-13 BMNH M. Peres & A. D. Brescovit & Fig. & Male & A - F. Specimen & A. Habitus & B. Front. & C - F. Left & C, E. Ventral & D, F. Retrolateral Brazil Recife 27 53 -15.178333 Reserva Biologica de Una 21 -39.10361 Horto Dois Irmaos 26 52 IBSP 38639, IBSP 46738, IBSP 46712, IBSP 46714, IBSP 46724, IBSP 46726 8 7 Pernambuco neotype 3400646524 2010-12-08 UFMG G. H. F. Azevedo & A. J. Santos -14.772778 Reserva Zoobotanica CEPLAC 21 -39.22028 27 53 UFMG 9484 1 1