Revision of the Australian Bee Genus Trichocolletes Cockerell (Hymenoptera: Colletidae: Paracolletini) Batley, Michael Records of the Australian Museum 2012 Rec. Aust. Mus. 2012-05-23 64 1 1 50 Batley, 2012 Batley 2012 [894,1253,1656,1682] Insecta Colletidae Trichocolletes Animalia Hymenoptera 26 26 Arthropoda species macrognathus sp. nov.   Figs 16–18, 35, 65, 103   Type.  Holotype ♂,  13 kmS Wanoo, Western Australia[ 26.49S, 114.37E],  24–28 Aug. 1984, T. F. Houston& B. P. Hanich, on flowers of  Baeckea, in WAM13671. Theco-ordinates on the label refer to the location of Wanooat 26°49'S 114°37'E.  Specimens examined. The holotypeand the following.  Western Australia:  ♀, 11 kmENE Anketell,  4–6 Sep. 1981, T. F. Houston, on flowers of Cassia phyllodineaor charlesiana, WAM(13659); ♂, Denham-Hamelin Rd NW   Tamalaturnoff,  28 Aug. 1997, T.  F. Houston& P. Mathiasen, on flowers of  Malleostemon roseus, WAM(13680); 4♂, ♀, East Yuna Reserve,  34 kmWNW Mullewa,  16–29 Aug. 1984, T.  F. Houston& B. P. Hanich, on flowers of  Thryptomene hyporhytis, WAM(13673–77); 3♀, 10 miE of Meekatharra,  31 Aug. 1971, T. F. Houston, SAM(32-024663–65);  2♀, 8 kmN Nerren NerrenHS,  25 Aug. 1985, R.  P. McMillan, on  Prostanthera wilkeana, WAM(13678–79); 3♂, 9♀, 13 kmS Wanoo,  24–28 Aug. 1984, T. F. Houston& B. P. Hanich, on flowers of  Baeckea& burrowing into walls of freshly dug pit in sand, WAM(13661–70,72). Diagnosis Eyes not hairy; metasomal bands wide, gold; malar space very long. Male S3 with distinctive hair plume. Female scutumwith transverse band of dark hair. Unlike any other species. Descriptions Male ( WAM21597).— Headwidth 4.35 mm, body length 12.0 mm. Relative dimensions: HW 50, HL 39, UID 36, UFW 35, LID 32, DMA 22, HVO 5, WOC 15, MOD4, OOD 10, IAD 10, ASD 3, AOD8, ML20, BMW 6, MSL 7.5, SL 13, SW 3, FL c. 59.— Eyesnot hairy, much shortened leaving malar area almost twice as long as wide ( Fig. 17); flagellum longer than head width; all flagellomeres c. 1.5× as long as wide. Legs slender; hind tarsus 1.3× as long as hind tibia; hind basitarsus 5.4× as long as wide; posterior carina of basitibial area extends past apex. Gonoforceps transversely truncate with long apical hairs (Fig. 35); lateral lobes of S7 fused with posterior projections and basal teeth ( Fig. 65).— Mandible, scape, tarsi, fore and hind tibiae, anterior faces of mid tibia and all femora, flagellomeres F2–8 ventrally, orange-brown; remainder of flagellum brown. T1–5 with wide, deep gold bands; T7, S6 laterally, orangebrown.—  Scutumwith dull sheen and moderately strong pit-reticulation. Scutumclosely and face densely covered with bright orange hair, erect on clypeus, fore basitarsus with brush of relatively short golden hair; S3 with dense medial plume of long white hair ( Fig. 18). Female ( WAM13661).— Headwidth 4.85 mm, body length 13.6 mm. Relative dimensions: HW 50, HL 37, UID 31, UFW 34, LID 32, DMA 27, HVO 4, WOC 14, MOD3, OOD 9, IAD 8, ASD 3, AOD10, ML19, BMW 8, MSL 5.0, SL 13, SW 3, FL c. 36.— Eyesnot hairy; face quadrate; length malar space c. 0.7× basal mandibular width ( Fig. 16); clypeus uniformly convex transversely; middle flagellomeres c. 1.5× as long as wide; inner hind tibial spur with c. 5 wellspaced teeth; tarsal claws with sharp inner ramus about half length of outer; pygidial plate broad with truncate, upturned margin.— Mandiblemedially orange-brown; legs dark brown. T1–4 with wide, bright gold bands.— Clypeuspolished with open to close large punctures; scutumpolished with close punctures.— Lower facewith long, plumose, white hair, dense in paraocular areas, sparse to open on clypeus and supraclypeal area; frons densely covered with plumose, orange-brown hair, darker towards ocelli. Scutumcovered with close to dense, plumose hair, anterior 20% and thin line around lateral and posterior margins pale orangebrown, remainder dark brown; semi-erect hair on terga strongly reflective giving metasoma a golden sheen, more pronounced apically.   Remarks. This species is referred to as F247/M 220 in Houston (2000).   Etymology. The specific name, used as a noun in apposition, is derived from the Greek words for large and jaw, referring to the greatly elongated malar space of this species.   Distribution. Area around Geraldton, Western Australia ( CAR, GS, MUR, YAL) ( Fig. 103). WAM 13671 1984-08-24 1984-08-28 1984-08-24 T, WAM F. Houston & B. P. Hanich & The -26.816668 Wanoo 1240 114.61667 26 26 1 1 Western Australia holotype 1981-09-04 1981-09-06 1981-09-04 T, WAM F. Houston Anketell 26 26 1 1 [166,449,154,173] 1997-08-28 T Tamala 27 27 1 1984-08-16 1984-08-29 1984-08-16 T F. Houston & P. Mathiasen East Yuna Reserve 27 27 5 1 4 1971-08-31 T F. Houston & B. P. Hanich Meekatharra 27 27 3 3 [437,735,250,269] SAM Houston 27 27 1 1985-08-25 R Nerren Nerren 27 27 2 2 1984-08-24 1984-08-28 1984-08-24 T P. McMillan Wanoo 27 27 12 9 3 WAM Houston & B. P. Hanich 27 27 1