A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae) Pulawski, Wojciech J. Proceedings of the California Academy of Sciences 2018 2018-12-17 65 1 584 Pulawski Pulawski 2018 [123,432,1179,1202] Insecta Crabronidae Pison Animalia Hymenoptera 461 462 Arthropoda species translucens sp. nov.    NAME DERIVATION.–  Translucens, Latinfor translucent; with reference to the translucent apical lamella of male tergum VII.   RECOGNITION. –  Pison translucenshas three submarginal cells, the second recurrent vein interstitial with the second intersubmarginal vein or nearly so, tegula partly impunctate and asetose, and setae appressed on tergum I. The gaster is all black (except tergum VII apically in the male), but the apical depressions of terga are brown and tergal setae are golden, forming golden fasciae on the apical depressions. The female is mainly characterized by the absence of specializations found in other species: the clypeus is the usual shape, with a roundly arcuate lamella that is longer mesally than laterally, with a deeply concave free margin of the lateral clypeal section, and the surface not concave above the lamella, the ocellocular distance is 0.9-1.0 × hindocellar diameter, the gena is punctate and setose on each side of the oral fossa (setae sinuous, as long as 1.5 × midocellar diameter), the tegula is largely impunctate and asetose, the propodeum is ridged and punctate on the dorsum and has a carina separating dorsum and posterior surface from the side and extending from the gastral socket area toward the spiracle, and sterna II-IV are punctate throughout. It closely resembles on the inner mandibular margin (tooth absent in  P. translucens); also, the setae of the lower gena are curved apically in  P. decipiens, while sinuous in  P. translucens, and the ocellocular distance is 0.9-1.1 × midocellar diameter in  P. translucens, but 1.1-1.5 in  P. impressiventre.   FIGURES1126-1128.  Pison translucensPulawski, sp. nov. (1126) Female clypeus and mandibles; (1127) Male clypeus and mandibles; (1128) Male apical terga in dorsal view. The male is easily recognized by its translucent, medially narrowed lamella along the apical margin of tergum VII (Fig. 1128), in combination with the setae of lower gena sinuous, as long as 1.0-1.2 × midocellar diameter. The shape of sternum VIII is a useful subsidiary recognition feature (Fig. 1129). The male of  P. aridumis similar, but differs in having the posterior margin of the black, sclerotized portion of tergum VII (adjacent to the yellowish portion) acutely angulate ( Fig. 114), and the genal setae straight, shorter than midocellar diameter; also the shape of sternum VIII is different (compare Figs. 1129 and 115).   DESCRIPTION.– Frons dull, punctures of upper frons (above midfrontal carina) compressed against each other. Occipital carina joining hypostomal carina. Gena in female narrow in dorsal view. Labrum shallowly emarginate. Anteromedian pronotal pit transversely elongate, about as long as midocellar diameter. Propleuron closely punctate in most specimens, but punctures several diameters apart in outer half in some. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures well defined, interspaces linear. Tegula enlarged, extending slightly beyond anterior margin of axilla, its outer margin straight or slightly convex (except anteriorly and posteriorly). Mesopleural punctures less than one diameter apart. Postspiracular carina present but ill defined, varying from about half diameter to entire diameter of midocellus. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum with short transverse ridges emerging from middle carina, otherwise closely punctate, interspaces merging into irregular ridges; side finely, irregularly ridged, punctate between ridges; posterior surface ridged, punctate between ridges. Posteroventral forefemoral surface finely punctate, punctures about one diameter apart. Hindcoxal dorsum with outer margin sharply carinate (except basally). Punctures of tergum I less than one diameter apart. Sterna punctate throughout.   FIGURES1129-1132.  Pison translucensPulawski, sp.nov., male. (1129) Sternum VIII (ventral surface); (1130) Sternum VIII in lateral view; (1131) Genitalia in dorsal view; (1132) Genitalia in lateral slightly oblique view. Setae silvery on head and thorax except golden on scutum, also golden on propodeal dorsum and terga (forming conspicuous fasciae on apical depressions), golden or with golden tinge on upper frons; both appressed and erect on frons and scutum; appressed on tergum I; oriented laterally above dorsal end of midfrontal carina and oriented ventrally beneath midocellus, partially concealing integument on clypeus in female, completely so in male (except lamella); setae of lower gena sinuous, as long as 1.0-1.2 × midocellar diameter. Head, thorax, propodeum, and gaster black (apex of tergum VII yellowish in male); mandible ferruginous except black basally and apically; apical depressions of terga brown. Femora in female all black or apex of hindfemur ferruginous; hindtibia ferruginous, also midtibia in many specimens and foretibia in some; tarsi all or largely black, all ferruginous in some specimens. In male forefemur black or ferruginous in apical third, midfemur black or ferruginous in apical half, hindfemur ferruginous (only apically so in some specimens); tibiae ferruginous or foretibia black; tarsi ferruginous.  ♀.– Upper interocular distance equal to 0.62-0.64 × lower interocular distance; ocellocular distance equal to 0.9-1.0 × hindocellar diameter, distance between hindocelli equal to 1.0-1.3 × hindocellar diameter; eye height equal to 0.82-0.87 × distance between eye notches. Free margin of clypeal lamella obtusely arcuate, slightly more than in  P. angulare(compare Figs. 1126 and 43). Dorsal length of flagellomere I 2.2-2.4 × apical width, of flagellomere IX 1.0-1.1 × apical width. Mandible: trimmal carina with small incision shortly beyond midlength. Length 9.5-11.3 mm; head width 3.0- 3.5 mm.  ♂.– Upper interocular distance equal to 0.90-0.96 × lower interocular distance; ocellocular distance equal to 1.6 × hindocellar diameter, distance between hindocelli equal to 1.5 × hindocellar diameter; eye height equal to 0.86-0.88 × distance between eye notches. Middle clypeal lobe markedly convex ventrally, base of lamella markedly below main clypeal surface; free margin of lamella sharply angulate. Dorsal length of flagellomere I 1.9-2.0 × apical width, of flagellomere X 0.9-1.0 × apical width. Tergum VII with translucent, medially narrowed lamella along apical margin, posterior margin of black, sclerotized portion of tergum (adjacent to lamella) broadly, obtusely tridentate (Fig. 1128). Apical margin of sternum VII broadly, shallowly concave; sternum VIII with impunctate, glabrous swelling basomedially, its apical margin slightly convex medially and slightly concave laterally; apicolateral arm broadly, obtusely angulate (Fig. 1129); lateral view: Fig. 1130. Genitalia: Figs. 1131, 1132. Length 8.8-11.6 mm; head width 2.9-3.3 mm.   GEOGRAPHIC  DISTRIBUTION(Fig. 1133).–  All Australiaexcept Tasmaniaand Victoria.   RECORDS.–   HOLOTYPE: ♂,   AUSTRALIA:   South    Australia: Wilpena in Flinders Ranges National Park at 31°31.7ʹS 138°36.2ʹE, 22 Dec 2010, V. Ahrens and W.J. Pulawski ( SAM).    PARATYPES:   AUSTRALIA:  New South Wales:  13 mi.N Broken Hill, 3 Apr 1963, K. Dansie ( 2 ♀,   1 ♂, SAM); 17 kmNE Broken Hillat 31°47ʹS 141°31ʹE, J. Carpenter and A. Davidson ( 1 ♂,   AMN); Coleambally Irrigation Areaat 34°56ʹ10ʺS  145°46ʹ51ʺE, 14 Dec 1998, L. Wilkie and S. Pride ( 1 ♂, AMS);  1 kmW Eumungerieat 31°56.7ʹS 148°36.9ʹE, 15, 19, and 20 Dec 2009, V. Ahrens and W.J. Pulawski ( 3 ♂, CAS); Fowlers Gap Research FIGURE1133. Collecting localities of  Pison translucensStation at 31°05ʹS 141°42ʹE, 29 Nov – 2 Dec 1981, Pulawski, sp. nov.   I.D. Naumannand J.C. Cardale( 4 ♀, 2 ♂, ANIC);  Menindee,  2 Dec 1992, N.W. Rodd( 1 ♀, AMS);  Springs Creek  68 kmSW Wilcanniaat 31°44ʹS 142°41ʹE,  29 Nov 1981, J.C. Cardaleand I.D. Naumann( 1 ♀, 1 ♂, ANIC);  Warrensburg National Park,  20 Dec 1987, M.E. Irwin( 1 ♀, UCD);   87 kmE Wilcanniaat 31°42.8ʹS 144°08.6ʹE, V. Ahrensand W.J. Pulawski,  21 Dec 2011( 10 ♀, 61 ♂, CAS) and  23 Dec 2011( 18 ♀, 65 ♂, CAS; 1 ♀, 1 ♂, NHMW);  Warrumbungle National Parkat 31°16ʹS 148°57ʹE,  17 Dec 1995, M.E. Irwin( 1 ♀, 1 ♂, MNKB).   Northern Territory:  30 kmWNW Alice Springsat 23°32ʹS 133°38ʹE,  7 Oct 1978, J.C. Cardale( 2 ♂, ANIC);  32 kmWNW Alice Springsat 23°36ʹS 133°35ʹE,  8 Oct 1978, J.C. Cardale( 6 ♂, ANIC);  Trephina Gorge Nature Parkat John Hayes Rockholeat 23°32ʹS 134°21ʹE,  10 Apr 1981, M. Malipatiland J. Hawkins( 1 ♂, NTM);  Trephina Gorge Nature Parkat Waterhouse Range  39 kmSSW Alice Springsat 23°59ʹS 133°38ʹE,  11 Oct 1978, J.C. Cardale( 2 ♂, ANIC);  West McDonnell National Parkca 3 kmW road to Simpson Gapat 23°41.8ʹS 133°41.7ʹE,  27 Jan – 27 Feb 2008, Ch.M. Palmer( 1 ♂, NTM).   Queensland: Coenat 13°57ʹS 143°12ʹE,  16 Aug – 13 Sept 1993, P. Zborowskiand S. Shattuck( 1 ♀, ANIC) and  17 Dec 1993–  13 Jan 1994, P. Zborowskiand E.D. Edwards( 1 ♀, ANIC);  Hann Riverat 15°11ʹS 143°52ʹE,  20 Mar – 24 Apr 1994, P. Zborowskiand G. Turner( 1 ♀, ANIC);   48 kmE Mount Surpriseat 18°09.0ʹS 144°43.6ʹE, V. Ahrensand W.J. Pulawski,  21 Nov 2012( 1 ♀, 12 ♂, CAS),   22 Nov 2012( 3 ♀, 16 ♂, CAS);  9 kmNW Mount Tozerat 12°44ʹS 143°08ʹE,  30 June – 7 July 1986, J.C. Cardale( 1 ♀, ANIC);  2 kmN Rokeby,  17 Dec 1993–  17 Jan 1994( 1 ♀, ANIC);  Split Rockat 15°39ʹS 144°31ʹE,  27 Apr – 28 May 1993, P. Zborowskiand A. Roach( 1 ♀, ANIC), and  28 May – 26 June 1993, P. Zborowskiand I.D. Naumann( 1 ♀, ANIC).   South Australia: Calperum Station  16 kmN Renmarkat 34°02.9ʹS 140°42.2ʹE,  4 Dec 2010, V. Ahrensand W.J. Pulawski( 1 ♂, CAS);  Calperum Station  79 kmN Renmarkat 33°31ʹS 140°24ʹE,  11 Oct – 9 Nov 1995, K. Pullen( 1 ♀, ANIC);  8 kmE Cedunaat 32°07.8ʹS 133°46.0ʹE,  8 Jan 2011, V. Ahrensand W.J. Pulawski( 2 ♀, 1 ♂, CAS);  Chowilla Game Reserve24 air km N Renmarkat 33°58.0ʹS 140°48.8ʹE,  5 Dec 2010, V. Ahrensand W.J. Pulawski( 1 ♂, CAS);  43 kmNNE Cowellat 33°20ʹS 137°06ʹE,  28 Nov 1992, I.D. Naumannand J.C. Cardale( 1 ♂, ANIC);  Dingly Dell Camp on Oraparina Creekat 31°21ʹS 138°42ʹE, I.D. Naumannand J.C. Cardale,  4 Nov 1987( 4 ♂, ANIC),   4-10 Nov 1987( 3 ♂, ANIC), and  7 Nov 1987( 4 ♂, ANIC);  Gawler National Parkat 32°35.1ʹS 135°26.3ʹE, V. Ahrensand W.J. Pulawski,  5 Jan 2011( 1 ♀, 11 ♂, CAS) and   7Jan 2011( 1 ♀, 2 ♂, CAS);  Gawler National Parkat 32°35.4ʹS 135°21.1ʹE,  7 Jan 2011, V. Ahrensand W.J. Pulawski( 2 ♀, 10 ♂, CAS);  28 kmS Gladstone,  11 Jan 1982, R.W. Thorp( 1 ♀, UCD);  9.7 kmN Hawk- er,  1 Mar 1972, E. Matthews( 1 ♀, SAM);   10 kmNNW Penongat 31°50.3ʹS 132°57.9ʹE, V. Ahrensand W.J. Pulawski,  16 Jan 2011( 5 ♀, 2 ♂, CAS) and  18 Jan 2011( 6 ♀, 7 ♂, CAS);  Port Clinton Conservation Parkat 34°09.4ʹS 138°03.2ʹE,  14 Dec 2010( 1 ♀, 4 ♂, CAS) and  16 Dec 2010( 2 ♀, 6 ♂, CAS),  V. Ahrensand W.J. Pulawski;  19 kmN Renmarkat 34°00ʹS 140°47ʹE, R K. Pullen,  10 Oct – 9 Nov 1995( 1 ♂, ANIC) and  8 Nov – 4 Dec 1995( 1 ♀, ANIC);  Sheoak Hill Conservation Reserve  38 kmNNW Covilleat 33°22.6ʹS 136°47.4ʹE,  29 Dec 2010, V. Ahrensand W.J. Pulawski( 1 ♀, 6 ♂, CAS);  Wilpena in Flinders Ranges National Parkat 31°31.7ʹS 138°36.2ʹE, V. Ahrensand W.J. Pulawski,  21 Dec 2010( 1 ♀, 1 ♂, CAS),   22 Dec 2010( 2 ♀, 8 ♂, CAS),   23 Dec 2010( 2 ♂, CAS),   27 Jan 2011( 1 ♀, 5 ♂, CAS), and 28 Jan1011( 1 ♂, CAS);   3 kmENE Wilpenaat 31°31.0ʹE 138°36.6ʹE, V. Ahrensand W.J. Pulawski,  23 Dec 2010( 2 ♂, CAS),   26 Jan 2011( 1 ♀, 2 ♂, CAS) and  27 Jan 2011( 2 ♀, 7 ♂, CAS);  34 kmS Wilpena,  4 Jan 1980, R.M. Bohart( 1 ♀, 13 ♂, UCD);  Wirreanda Creek  28 kmSW Hawkerat 32°05.9ʹS 138°17.7ʹE,  26 Jan 2011, V. Ahrensand W.J. Pulawski( 1 ♂, CAS).   Western Australia:  55 kmN Esperance,  25 Nov 1979, R.M. Bohart ( 1 ♀, UCD);  Ethel Creekat 22°54ʹS 120°10ʹE,  28 Nov 1971, N.S. ExpeditionIV ( 2 ♂, WAM);  12 kmNE Giles in Rawlinson Rangeat 25°02ʹS 128°18ʹE,  14 Jan 1990, T.F. Houston( 1 ♀, WAM);  Irrunytju Rockhole in Hinckley Rangeat 26°07ʹS 128°58ʹE,  19-21 Jan 1990, T.F. Houstonand M.S. Harvey( 1 ♂, WAM);  Kathleen Valley, 1963, T. Moriarty( 1 ♀, WAM);  Karijini National Parkat 22°30.1ʹS 118°24.4ʹE,  21-23 May 2003, M.E. Irwinand F.D. Parker( 1 ♂, CAS);   28 mi.E Leonora,  18 Sept 1962, E.S. Rossand D.Q. Cavagnaro( 3 ♀, 19 ♂, CAS);  Mount Gibson Station,  26 Feb 2000, S.R. Patterson( 1 ♂, WAM);  Nanutarra - Wittenoomroad 25 kmNE railway crossing at 22°21ʹ21ʺS 117°54ʹ16ʺE,  16-20 Feb 2005, M. Bulbertand S. Ginn( 1 ♀, AMS);   158 kmS Newman(= 9 kmN Kumarina Roadhouse) at 24°37.8ʹS 119°36.8ʹE,  18-21 May 2003, M.E. Irwinand D.D. Parker( 1 ♂, CAS);  Thomas River  23 kmWNW Mount Aridat 33°51ʹS 123°00ʹE,  4-7 Nov 1977, M.S. Upton( 1 ♂, ANIC);  Youanmiat 28°37ʹS 118°50ʹE,  13 Oct 1974, A.M. and M.J. Douglas( 2 ♂, WAM);  Yundamindra Homesteadat 29°15ʹS 122°06ʹE,  16 Mar 1979, T.F. Houston( 2 ♂, WAM). [123,588,827,850] Australia Tasmania 464 465 1 Tasmania [290,625,859,880] Australia South 464 465 1 1 holotype [171,618,975,996] Australia New South Wales 464 465 1 New South Wales paratype [123,625,1032,1053] SAM Broken Hill 464 465 1 1 [123,625,1090,1111] AMN -34.93611 Coleambally Irrigation Area 464 465 1 [269,625,1147,1168] Eumungerie 464 465 1 [123,610,1263,1284] ANIC I. D. Naumann & Cardale 464 465 6 4 2 [620,1089,1263,1284] 1992-12-02 AMS N. W. Rodd Menindee 464 465 1 1 1981-11-29 ANIC J. C. Cardale & I. D. Naumann Springs Creek 464 465 2 1 1 [202,850,1320,1341] 1987-12-20 UCD M. E. Irwin Warrensburg National Park 464 465 1 1 2011-12-21 CAS Pulawski Ahrens 464 465 71 10 61 2011-12-23 NHMW 464 465 85 19 66 1995-12-17 MNKB M. E. Irwin Warrumbungle National Park 464 465 2 1 1 1978-10-07 ANIC J. C. Cardale 30 km WNW Alice Springs 464 465 2 2 Northern Territory [255,1094,1435,1456] 1978-10-08 ANIC J. C. Cardale Alice Springs 464 465 6 6 Northern Territory 1981-04-10 NTM M. Malipatil & J. Hawkins John Hayes Rockhole Nature Park 464 465 1 1 Northern Territory 1978-10-11 ANIC J. C. Cardale Waterhouse Range Nature Park 464 465 2 2 Northern Territory 2008-01-27 2008-02-27 2008-01-27 NTM Ch. M. Palmer Simpson Gap West McDonnell National Park 464 465 1 1 Northern Territory 1993-08-16 1993-09-13 1993-08-16 ANIC P. Zborowski & S. Shattuck Coen 464 465 1 1 Queensland 1993-12-17 1994-01-13 1993-12-17 ANIC Zborowski & E. D. Edwards 464 465 1 1 Queensland 1994-03-20 1994-04-24 1994-03-20 ANIC P. Zborowski & G. Turner Hann River 464 465 1 1 Queensland 2012-11-21 CAS Pulawski Ahrens 464 465 13 1 12 Queensland [747,1064,1666,1687] 2012-11-22 CAS true Pulawski Ahrens 464 465 19 3 16 Queensland 1986-06-30 1986-07-07 1986-06-30 ANIC J. C. Cardale Mount Tozer 464 465 1 1 Queensland 1993-12-17 1994-01-17 1993-12-17 ANIC 465 466 Rokeby 464 465 1 1 Queensland 1993-04-27 1993-05-28 1993-04-27 ANIC P. Zborowski & A. Roach Split Rock 465 466 1 1 Queensland [361,1103,249,270] 1993-05-28 1993-06-26 1993-05-28 ANIC Zborowski & I. D. Naumann 465 466 1 1 Queensland 2010-12-04 CAS V. Ahrens & W. J. Pulawski Calperum Station 465 466 1 1 South Australia 1995-10-11 1995-11-09 1995-10-11 ANIC K. Pullen Calperum Station 465 466 1 1 South Australia 2011-01-08 CAS V. Ahrens & W. J. Pulawski Ceduna 465 466 3 2 1 South Australia 2010-12-05 CAS V. Ahrens & W. J. Pulawski Renmark Chowilla Game Reserve 465 466 1 1 South Australia 1992-11-28 ANIC I. D. Naumann & J. C. Cardale Cowell 465 466 1 1 South Australia 1987-11-04 ANIC I. D. Naumann Cardale Dingly Dell Camp on Oraparina Creek 465 466 4 4 South Australia [603,878,453,474] 1987-11-04 1987-11-10 1987-11-04 ANIC I. D. Naumann true Cardale Dingly Dell Camp on Oraparina Creek 465 466 3 3 South Australia [887,1170,453,474] 1987-11-07 ANIC 465 466 4 4 South Australia [123,1174,482,503] 2011-01-05 CAS V. Ahrens Pulawski Gawler National Park 465 466 12 1 11 South Australia [123,442,511,532] 2011-01-07 CAS 465 466 3 1 2 South Australia 2011-01-07 CAS V. Ahrens & W. J. Pulawski Gawler National Park 465 466 12 2 10 South Australia [446,1003,540,561] 1982-01-11 UCD R. W. Thorp Gladstone 465 466 1 1 South Australia 1972-03-01 SAM Matthews 465 466 1 1 South Australia 2011-01-16 CAS Pulawski Ahrens 465 466 7 5 2 South Australia [511,837,598,619] 2011-01-18 CAS 465 466 13 6 7 South Australia 2010-12-14 CAS Port Clinton Conservation Park 465 466 5 1 4 South Australia [643,979,628,649] 2010-12-16 CAS 465 466 8 2 6 South Australia 1995-10-10 1995-11-09 1995-10-10 ANIC V. Ahrens & W. J. Pulawski Pullen 465 466 1 1 South Australia 1995-11-08 1995-12-04 1995-11-08 ANIC 465 466 1 1 South Australia 2010-12-29 CAS V. Ahrens & W. J. Pulawski Sheoak Hill Conservation Reserve 465 466 7 1 6 South Australia 2010-12-21 CAS Wilpena in Flinders Ranges National Park Pulawski Ahrens 465 466 2 1 1 South Australia 2010-12-22 CAS Wilpena in Flinders Ranges National Park true Pulawski Ahrens 465 466 10 2 8 South Australia [189,427,773,794] 2010-12-23 CAS 465 466 2 2 South Australia [436,713,773,794] 2011-01-27 CAS 465 466 6 1 5 South Australia [722,994,773,794] CAS Jan 465 466 1 1 South Australia 2010-12-23 CAS V. Ahrens Pulawski 465 466 2 2 South Australia 2011-01-26 CAS V. Ahrens true Pulawski 465 466 3 1 2 South Australia [183,501,832,853] 2011-01-27 CAS 465 466 9 2 7 South Australia [511,1102,832,853] 1980-01-04 UCD R. M. Bohart Wilpena 465 466 14 1 13 South Australia 2011-01-26 CAS V. Ahrens & W. J. Pulawski Wirreanda Creek 465 466 1 1 South Australia [193,1017,890,911] 1979-11-25 UCD R. M. Bohart 55 km N Esperance 465 466 1 1 Western Australia 1971-11-28 WAM N. S. Expedition Ethel Creek 465 466 2 2 Western Australia 1990-01-14 WAM T. F. Houston Giles in Rawlinson Range 465 466 1 1 Western Australia 1990-01-19 1990-01-21 1990-01-19 WAM T. F. Houston & M. S. Harvey Irrunytju Rockhole in Hinckley Range 465 466 1 1 Western Australia WAM T. Moriarty Kathleen Valley 465 466 1 1 Western Australia 2003-05-21 2003-05-23 2003-05-21 CAS M. E. Irwin & F. D. Parker Karijini National Park 465 466 1 1 Western Australia [247,1049,1036,1057] 1962-09-18 CAS E. S. Ross & Cavagnaro 28 mi. E Leonora 465 466 22 3 19 Western Australia 2000-02-26 WAM S. R. Patterson Mount Gibson Station 465 466 1 1 Western Australia 2005-02-16 2005-02-20 2005-02-16 AMS M. Bulbert & S. Ginn -22.355833 Nanutarra - Wittenoom 21 117.90445 465 466 1 1 Western Australia 2003-05-18 2003-05-21 2003-05-18 CAS M. E. Irwin & D. D. Parker Kumarina Roadhouse 465 466 1 1 Western Australia 1977-11-04 1977-11-07 1977-11-04 ANIC M. S. Upton Thomas River 465 466 1 1 Western Australia [204,1032,1182,1203] 1974-10-13 WAM A. M. and M. J. Douglas Youanmi 465 466 2 2 Western Australia 1979-03-16 WAM T. F. Houston Yundamindra Homestead 465 466 2 2 Western Australia