Three new species of Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae) from Amazonian Peru and a checklist of Cyclocephala species in Peru Paz, Fernando Ratcliffe, Brett C. Figueroa, Luis Zootaxa 2022 2022-01-07 5087 3 427 440 Paz & Ratcliffe Paz & Ratcliffe 2022 [151,601,1734,1761] Insecta Dynastidae Cyclocephala Animalia Coleoptera 1 428 Arthropoda species ukuku sp. nov.  ( Figs. 1–6, 20–21)   Type material:  Holotypemale labeled: “ PERU, CU, RC / Machiguenga, Campto/ Mapi, 35.5 Km O de Nuevo/ Mundo,   741 m.73° / 28'30,46"/ 11°31'30,24", 13- /  18.i.2010 J. Grados” and with our red holotypelabel.  Allotypefemale labeled: “ PERU, CU. Zonade / amortiguamiento / Megantoni 12°34'18.6"S/ 73°5'13.18"W  1708 m. /  27.ix.2010 M. Alvaradoy / J. Peralta” and with our red allotypelabel. Two paratypemales with same data as allotype.  Holotype, allotype, and 1 paratypedeposited in MUSM, and 1 paratypein BCRC.   Description. Holotype,male. ( Fig. 1). Length 18.5 mm; width 9.1 mm. Color dark reddish brown, shiny, except for black tarsomeres and black on posterior half of frons. Head:Frons and clypeus with punctures moderate in size and density, setigerous; setae minute, tawny. Clypeal apex broadly subtruncate, slightly reflexed. Interocular width equals 3.1 transverse eye diameters. Antenna with 10 antennomeres, club subequal in length to antennomeres 2–7. Pronotum:Surface similar to that of frons; setae short, tawny. Base lacking marginal bead. Elytra:Surface weakly punctate-striate, transversely wrinkled, punctures slightly larger and denser than those of pronotum, setigerous; setae short, tawny. Pygidium:Surface with large, dense, setigerous punctures; setae long, tawny. In lateral view, surface weakly convex. Legs:Protibia slender, tridentate, basal tooth reduced to a rounded prominence. Protarsomere 5 enlarged, slightly bent, venter concave; medial claw large, strongly bent, apex split ( Fig. 5). Metatarsus slightly shorter than metatibia. Venter:Prosternal process long, columnar, apex obliquely flattened into transversely oval disc with transverse, elevated “button” on anterior half. Parameres:In caudal view, form elongate, subparallel, apices rounded ( Fig. 3). In lateral view, basal piece subequal in length to parameres ( Fig. 4).   Allotype,female ( Fig. 2). Length 19.3 mm; width across humeri 9.0 mm. As holotypeexcept in the following respects: Elytra:Epipleuron (ventral view) gradually tapering beginning at level of abdominal sternite 2 ( Fig. 6). In dorsal view, lateral margin simple, evenly rounded. Pygidium: In lateral view, surface weakly convex. Legs:Protibia distinctly tridentate. Protarsus simple, not enlarged.  Variation.Males ( 2 paratypes). Length 18.0– 19.5 mm; width 8.5–9.0 mm. The paratypesdo not differ from the holotypeexcept in size.   Etymology.  Cyclocephala ukukuis from Peruvian mythology and refers to a being from the mountains that is half bear and half human. The creature is known for being a dancer and guardian of the snowy mountains. This name should be treated as a noun in apposition.   Distribution. 4 specimensexamined ( Fig. 20). PERU(4): CUSCO(4): Reserva Comunal Machiguenga, Campamento Mapi, 35.5 kmW Nuevo Mundo (1); Zona de Amortiguamiento Megantoni (3).  Temporal distribution.January (1), September (3).   Diagnosis.  Cyclocephala ukukuis characterized by a distinctive, shiny, dark reddish-brown color ( Figs. 1–2); presence of small, abundant setae on the head, pronotum, and elytra; broadly subtruncate clypeus with a weakly emarginate apex; basal bead of the pronotum lacking; tridentate protibia; male large protarsal claw split at its apex ( Fig. 5); and uniquely shaped parameres ( Figs. 3–4). The female epipleuron is simple and gradually tapering ( Fig. 6).  Natural history.Specimens were collected at elevations of 741 mand 1708 min pristine forest ( Fig. 21), but nothing else is known of their life history. 3427663320 2010-01-18 J. Grados Peru Mundo 741 Machiguenga Km 1 428 741 holotype 3427663321 2010-09-27 M. Alvarado & J. Peralta Peru Mundo 1708 -12.571834 Zona 1 -73.087 Megantoni 1 428 3 allotype 3427663329 [235,861,1987,2013] MUSM Peru 1 428 1 holotype 3427663322 [872,1151,1987,2013] BCRC Peru 1 428 1 holotype