Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae)
Edward, Karen L.
Harvey, Mark S.
Bulletin of the American Museum of Natural History
2014
2014-06-06
2014
389
1
144
3Q658
Edward & Harvey, 2014
Edward & Harvey
2014
[155,406,801,824]
Arachnida
Oonopidae
Ischnothyreus
Animalia
Araneae
84
84
Arthropoda
species
eacham
sp. nov.
ETYMOLOGY: The specific epithet is a noun in apposition, taken from the typelocality.
DIAGNOSIS: The male of this species exhibits a simplified male palp, with a broad, obtusely bent distal tip. A small lateral process bends backward on the retrolateral side of the embolic region (fig. 50A, B, E). The female epigynal region is equally simple, and consists of a small triangular extention pointing anteriorly and positioned at the base of a long convoluted duct (figs. 49I, 50F).
MALE (PBI_OON 5883, figs. 49A–C, G– H, 50A–E). Total length 1.63. CEPHALO- THORAX: Carapaceorange-brown, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less, anterolateral corners without extension or projections, surface of elevated portion of pars cephalica smooth, sides finely reticulate. Clypeusmargin unmodified, curved downward in front view, low, ALE separated from edge of carapace by less than their radius; setae dark. Eyes:ALE largest, ALE circular, PME oval, PLE oval; posterior eye row straight from above; ALE touching, ALE- PLE touching. Sternumlonger than wide, pale orange, uniform; setae dark, evenly scattered. Labium and endites dark red brown, chelicerae pale. Chelicerae slightly divergent, anterior face unmodified; promargin with one or two slightly larger denticles; fang shape normal, without prominent basal process; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, same as sternum in sclerotization; with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. AB- DOMEN: Ovoid; dorsum soft portions white. Book lung covers elliptical. Dorsal scutum pale orange, covering 1/2 to 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface smooth, sides smooth. Epigastric scutum small lateral sclerites present. Postepigastric scutum pale orange, covering about 2/3 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Pale orange, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-1-1; II p0-2-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur one to two times as long as trochanter; patella about as long as femur; cymbium dark red-brown; bulb dark red-brown, 1 to 1.5 times as long as cymbium, stout; embolic region stout, broad, obtusely bent, small lateral process bent backward on retrolateral side (fig. 50A, B, E). FEMALE (PBI_OON 00005854, figs. 49D– F, I, 50F). Total length 1.90. CEPHALO- THORAX: Carapacepale orange, pars cephalica slightly elevated in lateral view, surface of elevated portion of pars cephalica finely reticulate. Clypeusstraight in front view. Chelicerae, endites, and labium pale orange. ABDOMEN: Dorsal scutum between 1/4 and 1/2 abdomen width. Postepigastric scutum widely hexagonal, only around epigastric furrow. LEGS: Leg spination: femora: I p0-1-1; II p0-2-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: epigynal region simple, epigynal atrium not clearly visible but with slight depression, posterior portion of depression with small triangular extention pointing anteriorly; convoluted duct thicker than apodemes, long (figs. 49I, 50F). Fig. 49. Ischnothyreus eacham, sp. nov.Holotype male (PBI_OON 00005883): A.habitus, dorsal view; B.habitus, ventral view; C.habitus, lateral view; G.carapace, anterior view; H.sternum, ventral view. Allotype female (PBI_OON 00005854): D.habitus, dorsal view; E.habitus, ventral view; F.habitus, lateral view; I.epigynum, ventral view. OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: Curtain Fig Tree, 1 kmSW of Yungaburra, rainforest, leaf litter, 17.27472 °S, 145.57333 °E, Dec. 3, 1969( J.G. Brooks, ANIC, PBI_ OON 5839), 1 ♀; 1 Fig. 50. Ischnothyreus eacham, sp. nov.Holotype male (PBI_OON 00005883): A.left palp, prolateral view; B.left palp, retrolateral view; C.left chelicerae, anterior view; D.left chelicerae, posterior view; E.left palp, dorsal view. Allotype female (PBI_OON 00005854): F.epigynum, ventral view. Scale lines 5 0.1 mm. ♀( ANIC PBI_ OON 5839); Danbulla National Park, Mobo Creekcrater circuit walk, rainforest, leaf litter, 17.16944 °S, 145.63888 °E, Apr. 22, 2009(K. Edwardand P. Cullen, QM S95953, PBI_OON 26316), 1 ³; Dinden National Park, start of Kahlpahlimrock trail, rainforest, leaf litter, 17.03388 °S, 145.61305 °E, Apr. 11, 2009(K. Edwardand J. Waldock, QM S95954), 3 ♀; Dinden National Park, start of Kahlpahlimrock trail, rainforest, leaf litter, 17.03388 °S, 145.61305 °E, Apr. 11, 2009(K. Edwardand J. Waldock, QM S95955, PBI_OON 26318), 3 ♀; Boar Pocket Road, 5 milesN. of Gillieshighway, rainforest, leaf litter, 17.20000 °S, 145.66666 °E, Dec. 13, 1969(J.G. Brooks, ANIC, PBI_OON 25850), 1 ³, 2 ♀; Boar Pocket Road, 5 milesN. of Gillieshighway, rainforest, leaf mould, 17.20000 °S, 145.66666 °E, Nov. 15, 1969(J.G. Brooks, ANIC, PBI_OON 25872), 1 ♀; Cathedral Tree, 720 m, 17.18333 °S, 145.65000 °E, Feb. 7, 1996(G. Monteith, QM S43828, PBI_OON 22053), 1 ♀; Crater Lakes National Park, Lake Barinewalk trail, rainforest, leaf litter, 17.23722 °S, 145.64083 °E, Apr. 23, 2009(K. Edwardand P. Cullen, QM S95956, PBI_ OON 5562), 1 ♀; Crater Lakes National Park, Lake Eachamwalk, rainforest, leaf litter, 17.28805 °S, 145.62166 °E, May 18, 2007(K. Edwardand K. Pitz, QM S95957), 1 ♀; Crater Lakes National Park, Lake Eachamwalk, rainforest, leaf litter, 17.28805 °S, 145.62166 °E, May 18, 2007(K. Edwardand K. Pitz, QM S95958, PBI_OON 25719), 2 ♀; Crater Lakes National Park, Lake Eachamwalk, rainforest, leaf litter, 17.28805 °S, 145.62166 °E, May 18, 2007(K. Edwardand K. Pitz, QM S95959, PBI_OON 25720), 1 ³; Davies CreekRoad, 20 kmESE Mareeba, 750 m, Dec. 4–13, 1988(G. Monteith, G. Thompson, QMS49364, PBI_OON 22080), 1 ♀; Eacham National Park, rainforest, leaf litter, 760 m, 17.2833 °S, 145.6167 °E, Oct. 1–7, 1972(R.W. Taylor, ANIC, PBI_ OON 5842, 5854), 4 ♀; Eacham National Park, rainforest, 760 m, 17.2833 °S, 145.6167 °E, Feb. 16, 1973(R.W. Taylor, ANIC, PBI_ OON 5859), 1 ♀; Eacham National Park, rainforest, 760 m, 17.2833 °S, 145.6167 °E, Mar. 23, 1975(R.W. Taylor, ANIC, PBI_ OON 5862), 1 ³, 2 ♀; Eacham National Park, rainforest, leaf litter, 760 m, 17.2833 °S, 145.6167 °E, Oct. 1–7, 1972(R.W. Taylor, ANIC, PBI_OON 25851), 1 ♀; Eacham National Park, rainforest, leaf litter, 760 m, 17.2833 °S, 145.6167 °E, Mar. 21, 1975(R.W. Taylor, ANIC, PBI_OON 25854), 1 ♀; Gadgarra Road, 5 kmE Lake Eacham, 700 m, 17.26667 °S, 145.66670 °E, Dec. 9, 1989(G. Monteith, G. Thompson, H. Janetzki, QM S31926, PBI_OON 22232), 1 ³, 2 ♀; Mount Edith Road, rainforest, leaf litter, 1000 m, 17.06777 °S, 145.57305 °E, Apr. 9, 2009(K. Edwardand J. Waldock, QM S95960, PBI_ OON 5533), 1 ³, 1 ♀; Mount Edith Road, rainforest, leaf litter, 1000 m, 17.06777 °S, 145.57305 °E, Apr. 9, 2009(K. Edwardand J. Waldock, QM S95961, PBI_OON 26317), 2 ³, 3 ♀; same data ( WAM T130762), 2 ³, 2 ♀; North Bell Peak, Malbon Thompson Range, 17.11667 °S, 145.88330 °E, Nov. 20–22, 1990(G. Monteith, G. Thompson, QM S47016, PBI_OON 22066), 1 ♀; Tolga, near Atherton, rainforest, leaf litter, 17.17805 °S, 145.45916 °E, June 14, 1978(A. and M. Walford-Huggins, ANIC, PBI_OON 25806), 1 ³; WongabelSF, rainforest, 17.32167 °S, 145.49080 °E, Nov. 05, 1991, to July 23, 1992(R. Raven, P. Lawless, E. Lawless, M. Shaw, QM, S24209, PBI_OON 21747), 2 ♀; WongabelSF, 17.32167 °S, 145.49080 °E, July 23–Nov. 26, 1992(R. Raven, P. Lawless, E. Lawless, M. Shaw, QM S22607, PBI_OON 22098), 1 ³; WongabelSF, forest walk, rainforest, under bark, 17.32500 °S, 145.49583 °E, Apr. 22, 2009(K. Edwardand P. Cullen, QM S95962, PBI_ OON 5561), 1 ♀; Wooroonooran National Park, Tchupala Fallstrack, rainforest, leaf litter, 17.61000 °S, 145.77722 °E, Apr. 27, 2009(K. Edwardand P. Cullen, QM S95963, PBI_ OON 5557), 1 ³; Wooroonooran National Park, Tchupala Fallstrack, rainforest, leaf litter, 17.61000 °S, 145.77722 °E, Apr. 27, 2009(K. Edwardand P. Cullen, QM S95964, PBI_OON 26307), 2 ³.
DISTRIBUTION: This species is known only from three central subregions of the Wet Tropics Bioregion (AU, LU, and MT), in northeastern Queensland(map 8).
1973-02-20
ANIC, PBI
R. W. Taylor
Australia
760
-17.2833
Eacham National Park
7
145.6167
Male
84
84
OON 00005883, OON 00005854
5
2
Queensland
holotype
1969-12-03
ANIC, PBI
J. G. Brooks
-17.27472
Curtain Fig Tree
1
145.57333
85
85
OON 5839
1
1
Queensland
[93,431,1498,1524]
ANIC, PBI
86
86
OON 5839
1
1
S95953, OON 26316
2009-04-22
QM, PBI
Edward & P. Cullen
-17.16944
Danbulla National Park
1
145.63889
Mobo Creek
86
86
1
S95954
2009-04-11
QM
Edward & J. Waldock
-17.03388
Dinden National Park
1
145.61305
Kahlpahlim
86
86
3
3
S95955, OON 26318
2009-04-11
QM, PBI
Edward & J. Waldock
-17.03388
Dinden National Park
1
145.61305
Kahlpahlim
86
86
3
3
OON 25850
1969-12-13
ANIC, PBI
Brooks
-17.2
Boar Pocket Road
1
145.66666
Gillies
86
86
2
2
OON 25872
1969-11-15
ANIC, PBI
Brooks
87
87
-17.2
Boar Pocket Road
1
145.66666
Gillies
86
86
1
1
S43828, OON 22053
1996-02-07
QM, PBI
Monteith
720
-17.18333
Cathedral Tree
1
145.65
87
87
1
1
S95956
2009-04-23
QM, PBI
Edward & P. Cullen
-17.23722
Crater Lakes National Park
1
145.64082
Lake Barine
87
87
OON 5562
1
1
S95957
2007-05-18
QM
Edward & K. Pitz
-17.28805
Crater Lakes National Park
1
145.62166
Lake Eacham
87
87
1
1
S95958, OON 25719
2007-05-18
QM, PBI
Edward & K. Pitz
-17.28805
Crater Lakes National Park
1
145.62166
Lake Eacham
87
87
2
2
S95959, OON 25720
2007-05-18
QM, PBI
Edward & K. Pitz
-17.28805
Crater Lakes National Park
1
145.62166
Lake Eacham
87
87
1
S49364, OON 22080
1988-12-04
1988-12-13
1988-12-04
QM, PBI
Monteith & Thompson
750
Davies Creek
87
87
1
1
1972-10-01
1972-10-07
1972-10-01
ANIC, PBI
Taylor
760
-17.2833
Eacham National Park
7
145.6167
87
87
OON 5842, 5854
4
4
1973-02-16
ANIC, PBI
Taylor
760
-17.2833
Eacham National Park
7
145.6167
87
87
OON 5859
1
1
1975-03-23
ANIC, PBI
Taylor
760
-17.2833
Eacham National Park
7
145.6167
87
87
OON 5862
2
2
OON 25851
1972-10-01
1972-10-07
1972-10-01
ANIC, PBI
Taylor
760
-17.2833
Eacham National Park
7
145.6167
87
87
1
1
OON 25854
1975-03-21
ANIC, PBI
Taylor
760
-17.2833
Eacham National Park
7
145.6167
87
87
1
1
S31926, OON 22232
1989-12-09
QM, PBI
Monteith & G. Thompson & H. Janetzki
700
-17.26667
Gadgarra Road
1
145.6667
87
87
2
2
S95960
2009-04-09
QM, PBI
Edward & Waldock
1000
-17.06777
Mount Edith Road
1
145.57304
87
87
OON 5533
1
1
S95961, OON 26317
2009-04-09
QM, PBI
Edward & Waldock
1000
-17.06777
Mount Edith Road
1
145.57304
87
87
3
3
S95961, OON 26317
2009-04-09
WAM
Edward & Waldock
1000
-17.06777
Mount Edith Road
1
145.57304
87
87
T130762
2
2
S47016, OON 22066
1990-11-20
1990-11-22
1990-11-20
QM, PBI
Monteith & G. Thompson
-17.11667
North Bell Peak
1
145.8833
Malbon Thompson Range
87
87
1
1
OON 25806
1978-06-14
ANIC, PBI
Walford-Huggins
-17.17805
Tolga
1
145.45917
Atherton
87
87
1
S24209, OON 21747
1991-11-05
1992-07-23
1991-11-05
QM, PBI
Raven & P. Lawless & E. Lawless & M. Shaw
-17.32167
Wongabel
1
145.4908
87
87
2
2
S22607, OON 22098
1992-07-23
1992-11-26
1992-07-23
QM, PBI
Raven & Lawless & M. Shaw
-17.32167
Wongabel
1
145.4908
87
87
1
S95962
2009-04-22
QM, PBI
Edward & Cullen
-17.325
Wongabel
1
145.49583
87
87
OON 5561
1
1
S95963
2009-04-27
QM, PBI
Edward & P. Cullen
-17.61
Wooroonooran National Park
1
145.77722
Tchupala Falls
87
87
OON 5557
1
S95964, OON 26307
2009-04-27
QM, PBI
Edward & P. Cullen
-17.61
Wooroonooran National Park
1
145.77722
Tchupala Falls
87
87
1