Duplicaria mozambiquensis— Aubry, 1992 Aubry etal. , 2006 Bratcher & Cernohorsky, 1982 Phylogenetic classification of the family Terebridae (Neogastropoda: Conoidea) Fedosov, Alexander E Malcolm, Gavin Terryn, Yves Gorson, Juliette Modica, Maria Vittoria Holford, Mandë Puillandre, Nicolas Journal of Molluscan Studies The Malacological Society of London 2019 2020-01-09 85 4 359 388 7R5S5 Malcolm, Terryn & Fedosov Malcolm, Terryn & Fedosov 2019 [861,1418,408,430] Gastropoda Terebridae Duplicaria Animalia Neogastropoda 22 23 Mollusca species herberti sp. nov.  ( Fig. 11A–C)      Duplicaria mozambiquensis— Aubry, 1992: Fig. 9. Aubry etal., 2006: pl. 16. (Both not Bratcher & Cernohorsky, 1982).    Typematerial:  Holotype: MNHN IM-2013-52381, lv, 29.7 mm; InhacaI., Mozambique, 26°00.0'S, 32°54.4'E,  4 m( INHACA 2011Stn MR15).  Paratypes1–3: Inhaca I., Mozambique, 26°03.1'S, 33°01.0'E,  50–53 m(INHACA 2011, Stn MD13).  Paratype1: MNHN IM-2013-52373, lv, 17.4 mm.  Paratype2: MNHN IM-2013-52379, lv, 23.0 mm.  Paratype3: MNHN IM- 2013-52383, lv, 28.2 mm.  Paratype4: MNHN IM-2013-52385, 1 lv, 30.8 mm; typelocality.  Paratype5: MNHN IM-2013-52405, lv, 24.0 mm; 26°05.0'S, 32°59.0'E,  0–35 m(INHACA 2011Stn MA15).  Paratypes6–19: YT, 14 lv, 22.4–32.5 mm; Inhaca I., Mozambique,  6–8 m.  Paratype20: SG, lv, 24.0 mm; off Inhaca I., Mozambique, 25°59'59.3''S, 32°54'43.2''E, 4 m.  Paratypes21–25: JR, 4 lv, 30.0– 39.5 mm; off Inhaca I., Mozambique,  4–8 m.  Paratype26: GM, lv, 26 mm; Inhambane, Mozambique.  Paratype27: YT, dd, 32.0 mm; off Durban Bluff, Natal, South Africa, in sand dredging at bay-head dump.   Zoobank registration: urn:lsid:zoobank.org:act: C26159E5-4A5A- 4218-AC7F-A76103809749   Diagnostic nucleotide positions: Table 11.   Description:Shell of medium size. Protoconch paucispiral, about 1.0–1.5 broad whorls. Teleconch of holotype11 whorls. Outline of teleoconch whorls straight. Spiral sculpture absent. Subsutural band with axially elongated nodes corresponding to axial ribs on remainder of whorl. Subsutural band bordered by deep and wide depression lacking punctations. Straight axial ribs stretching across whorl, sharply angular adapically and becoming more rounded abapically; ribs half as wide as interspaces; 17 ribs on penultimate whorl of holotype. Axial growth lines pronounced throughout whorl height. Aperture wide, somewhat quadrate, brown with white band; columella straight with visible fold. Shell colour dark, later whorls blackish brown with spiral whitish line at the periphery.   Habitat: Depth 0– 55 m.   Distribution: S Mozambique to off Durban, South Africa.   Etymology: The species honours Dr David G. Herbert (formerly Chief Curator of Mollusca, NMSA), who contributed with detailed observations on the  D. mozambiquensis typeseries at NMSA and provided additional historical information.   Remarks: Shell colour varies: juveniles are beige to fawn, but light brown specimens are known. This species has been confused with  D. mozambiquensis. The holotypeof  D. mozambiquensis(NMSA H7843/T2541) is a small, slender shell of 22.3 mm, but the typeseries shows a large discrepancy in many features between this holotypeand all the paratypes—which are specimens of  D. herberti n. sp.Subsequent authors have to our knowledge always featured specimens of  D. herberti n. sp.as ‘  D. mozambiquensis’, which should be considered a rarely encountered species.   Figure 11.  Duplicaria herberti n. sp.,  Partecosta bozzettii n. sp.and morphologically similar species. A.  D. herberti, holotype, MNHN-IM-2013-52381, INHACA 2011 Stn MR15, 26 °00.0 'S, 32 °54.4 'E, 0–1 m, 29.7 mm. B.  D. herberti, MNHN-IM-2013-52366, INHACA 2011 Stn MM1, 26 °02.3 'S, 32 °54.1 'E, 0–1 m, 12.4 mm. C.  D. herberti(paratype of  D. mozambiquensis), NMSA 566, 27 mm. D.  D. mozambiquensis, YT, Quelimane Pebane, Mozambique, 35–45 m, 19.6 mm. E.  Partecosta bozzettiiholotype, MNHN-IM-2009-10163, ATIMO VATAE Stn TP29, 25 °03 '43.9 ''S, 46 °57 '42.9 ''E, 3–4 m, 12.5 mm. F.  Partecosta trilineataholotype, MNHN-IM-2000-21473, S Madagascar, Lavanono, 8.85 mm. G.  P. daniaeholotype, MMM, Farol das Lagostas, Luanda, Angola, 12 mm. The whorls of  D. mozambiquensis( Fig. 11D) have a narrower apical angle, its convex axial ribs giving an extremely rounded convex outline. The subsutural band comprises round nodes, compared with elongated nodes on  D. herberti n. sp.The axial ribs of  D. herberti n. sp.become straight, creating an angular projection posteriorly. The peripheral white band in  D. mozambiquensisis wider and more clearly defined. Both have a subsutural furrow with minute axial growth striae, appearing as a punctate groove in  D. mozambiquensis, while this feature is not present in  D. herberti n. sp.We have no confirmation that  D. mozambiquensishas ever been found in southern Mozambique, while specimens of  D. herberti n. sp.are found extensively in southern Mozambique and South Africa. Within  Duplicaria, several species change the sculpture of their whorls as they grow and do so at variable rates. However, a comparison of the two species highlights differences in the early whorls and protoconch. The protoconch of  D. mozambiquensisis about 30% narrower than that of  D. herberti n. sp.and the latter has a broader, inflated shape. The early whorls of  D. mozambiquensishave distinctive round nodes compared with convex ribs on  D. herberti n. sp. 3026465321 MNHN Mozambique 4 -26.0 Inhaca 124 32.906666 22 23 IM-2013-52381, INHACA 2011 1 Inhaca holotype 3026465318 2011 MNHN Mozambique 52 -26.051666 Inhaca I. 124 33.016666 22 23 IM-2013-52373 2 Inhaca I. paratype 3026465301 [837,1351,701,723] 2011 MNHN Mozambique 52 -26.051666 Inhaca I. 124 33.016666 22 23 IM-2013-52373 2 Inhaca I. paratype 3026465311 MNHN 22 23 IM-2013-52379 1 paratype 3026248327 MNHN 22 23 IM- 2013-52383 1 paratype 3026248317 MNHN 22 23 IM-2013-52385 2 paratype 3026465316 2011 MNHN 18 -26.083334 124 32.983334 22 23 IM-2013-52405 1 paratype 3026248340 Mozambique 7 Inhaca I. 22 23 1 Inhaca I. paratype 3026248337 Mozambique -25.999805 Inhaca I. 1 32.912003 22 23 1 Inhaca I. paratype 3026248313 Mozambique 6 Inhaca I. 22 23 1 Inhaca I. paratype 3026248314 [837,1384,941,963] Mozambique Inhambane 22 23 1 Inhambane paratype 3026465323 sand dredging at bay-head dump South Africa Durban Bluff Durban Bluff 22 23 1 Natal paratype