Phylogenetic classification of the family Terebridae (Neogastropoda: Conoidea) Fedosov, Alexander E Malcolm, Gavin Terryn, Yves Gorson, Juliette Modica, Maria Vittoria Holford, Mandë Puillandre, Nicolas Journal of Molluscan Studies The Malacological Society of London 2019 2020-01-09 85 4 359 388 7QB73 Malcolm, Terryn & Fedosov Malcolm, Terryn & Fedosov 2020 [175,799,232,254] Gastropoda Terebridae Bathyterebra GBIF Animalia Neogastropoda 22 23 Mollusca species zhongshaensis sp. nov.  ( Fig. 2H)    Typematerial:  Holotype: MNHN IM-2013-61800, 17.4 mm; South China Sea, Helen Shoal, 19°13'N, 113°55'E,  470–494 m(ZhongSha 2015Stn DW4138).   Zoobank registration: urn:lsid:zoobank.org:act: C8461C7A-BF53- 48D4-BE1B-EF23F17413F8   Diagnostic nucleotide positions: Table 11.   Description:Shell small, with orthocline spire, wide apical angle. Protoconch wide, paucispiral, 2.5 whorls, with small nucleus. Teleoconch 9 whorls. Early whorls convex in outline with strong, sharply convex ribs; subsequent whorls with narrow constriction formed by indentation, with straight ribs below suture; suture distinct, ragged; mature whorls with less convex outline, weaker and more numerous ribs, 30 on penultimate whorl. Spiral sculpture of very weak threads. Last whorl bulbous at base, but elongate with wide aperture. Columella straight, with weak fold. Overall colour white, with extensive patches of light-tan axial flammules on mature whorls.   Habitat:Depths of 470– 494 m.   Distribution:Only known from typelocality.   Etymology:Named for the Zhongsha sand bars, which form a subsurface atoll on top of the Macclesfield Bank.   Remarks: Bathyterebra zhongshaensis n. sp.is similar to the deep-water species,  B. coriolisiand  B. benthalis.  Bathyterebra coriolisi( Fig. 2E) is similar in colour, in the sculpture of the early whorls and in the shape of its aperture. However,  B. coriolisidiffers in having a much narrower shape, with a strong subsutural groove and fewer ribs on the early whorls, creating a band with nodes. Its subsutural groove becomes weaker, with just a slight indentation on mature whorls.  Bathyterebra benthalis( Fig. 2I) is similar in colour and in overall sculpture of the shell, but differs from  B. zhongshaensis n. sp.in the outline of its mature whorls being flat and the ribs on the early whorls extremely convex, creating nodes that dominate the profile of the early whorls. 3026465310 2015 MNHN South China Sea 482 19.216667 Helen Shoal 1272 113.916664 22 23 IM-2013-61800 1 Helen Shoal holotype