Liogenys nigrofuscus Moser, 1918: 103
Frey 1969: 50
Krajčík 2012: 145
Cherman et al. 2017: 4
Liogenys nigrofusca : Blackwelder 1944: 227
Evans 2003: 211
Evans & Smith 2009: 179
Liogenys longicrus Moser, 1918: 100
Frey 1965: 145
Liogenys longicra : Blackwelder 1944: 227
Liogenys Guérin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) from the Chacoan Province and its boundaries: taxonomic overview with four new species
Cherman, Mariana Alejandra
Basílio, Daniel Silva
Mise, Kleber Makoto
Frisch, Johannes
Almeida, Lúcia Massutti De
Zootaxa
2021
2021-02-25
4938
1
1
59
Moser, 1918
Moser
1918
[151,556,1795,1822]
Insecta
Melolonthidae
Liogenys
Animalia
Coleoptera
30
31
Arthropoda
species
nigrofusca
Figs. 14; 24.
Liogenys nigrofuscus Moser, 1918: 103; Frey 1969: 50(key); Krajčík 2012: 145(checklist); Cherman et al.2017: 4(generic history). Liogenys nigrofusca: Blackwelder 1944: 227(checklist); Evans 2003: 211(checklist); Evans & Smith 2009: 179(checklist). Liogenys longicrus Moser, 1918: 100; Frey 1965: 145(synonymy). Liogenys longicra: Blackwelder 1944: 227(checklist).
Type material. Liogenys nigrofuscus male holotype( ZMHB): [white, typeset and handwritten] “ Argentinien/ Gran Chaco”, [white handwritten] “ Liogenys/ nigrofuscus / Typen Mos”, [red typeset] “ Typus”, [white typeset] “ Liogenys/ nigrofuscus / Mos.” Liogenys longicrus male holotype( ZMHB): [white typeset] “ Argentinien/ Santiago del Estero”, [white handwritten] “ Liogenys/ longicrus / Typen Mos”, [red typeset] “ Typus”, [white handwritten] “ Liogenys/ nigrofuscus / Mos. [typeset] det. G. Frey 1964” Non-type material(10). PARAGUAY. Boquerón: Colonia Menonitas, VII.1945, Podtiaguin 2 males( MZUC); Guachalla, A. Pilcomayo, 23.VIII.1945, P. Willim, 1 male( MZUC). ARGENTINA. Salta: Viñaco( 15 kmElCarril), 12.II.1982, H. Howden, 1 female( CMNC); Coronel Moldes, XII.1958, A. Martínez, 1 female( MZSP); Chaco: Pampa del Infierno[ 26º31’S, 61º10’W], IX.1982, A. Martínez, 1 female( CMNC); Estación San Francisco, Departamento San Fernando, 16–17.X.2013, 54 m, 27°30’35.90’’S, 59°4’47.60’’W, M. Ibarra Polesel, 1 specimen; Antequera, Departamento Primero de Mayo, 22.XII.2014, 52 m, 27°26.500’S, 58°51.459’W, M. Ibarra Polesel, 1 specimen(CARTROUNNE); Santiago del Estero: El Pinto, without date and collector, 2 females( NHMB).
Diagnosis. Length: 9.1–10.0 mm; width: 4.1–4.8 mm. Body and elytra dark brown, elongate, shiny, sides parallel; pronotum and elytra unicolored, head darker; elytra glabrous ( Fig. 14A); pronotum roughly punctate; frons somewhat swollen and depressed medially in females; clypeal emargination deep, rounded and narrow; outer sides of anterior teeth parallel; teeth sharp; clypeal lateral margin bearing a sharp tooth-like projection, angle between this projection and the anterior teeth approximately 90º; antennae with 10 antennomeres; pronotal posterior corners rounded ( Fig. 14B); hypomere, mesepisternum, sides of metaventrite, metacoxae and sides of ventrites with sparse long bristles ( Fig. 14C); protibial teeth, distance between medial and apical shorter than basal and medial; mesotibia cylindrical in cross section; two strong transverse carinae posteriorly, the basal sloped, the apical complete; pygidium flat; quadrate; pygidial disc very coarsely and densely punctate, sparser in females; with bristles throughout, bristles erect, long ( Fig. 14D); in males, inner margin of metatibia strongly carinate straight towards the apex, somewhat curved outwards ( Fig. 14C); protarsomere II elongate; parameres narrowed medially; parameral split at the midline; apex club shaped and strongly truncate on upper margin ( Fig. 14E); parameres in lateral view convex, not coplanar ( Fig. 14F).
Typelocality. Liogenys nigrofusca: ARGENTINA, “Gran Chaco”; L. longicra: ARGENTINA, Santiago del Estero. Geographical distribution. PARAGUAY( Boquerón); ARGENTINA ( Salta, Chaco, Santiago del Estero).
Remarks. Frey (1965)synonymized L. nigrofuscawith L. longicra, and compared this species with the preceding one in order of description, L. parallela Frey, 1965( Fig. 18). According to Frey (1965), both are almost undistinguishable but the parameres. Although L. nigrofuscaresembles L. parallelain the body size and in the shape of the clypeus, especially in females, they have external differences. This species differs from L. parallela(in parenthesis) mainly in the color darker, basal transverse carina on the posterior margin of mesotibia and metatibia strong and sloped (basal transverse carina less marked and straight on mesotibia, absent on metatibia); and in the pygidium quadrate, with punctures on disc coarser and uniformly disposed.
3047558467
ZMHB
Argentina
Gran
Typus
Typen Mos
32
33
1
Chaco
holotype
3047558428
ZMHB
Argentina
Typus
Typen Mos
32
33
1
Santiago del Estero
holotype
3047558432
[483,1432,366,393]
1945-07
MZUC
Podtiaguin
Paraguay
Colonia Menonitas
32
33
2
2
Boqueron
3047558316
[151,918,402,429]
1945-08-23
MZUC
Guachalla, A & P. Willim
Paraguay
Pilcomayo
32
33
1
1
Boqueron
3047558311
1982-02-12
CMNC
H. Howden
Argentina
Vinaco
32
33
1
1
Salta
3047558455
[690,1346,438,465]
1958-12
MZSP
A. Martinez
Argentina
Coronel Moldes
32
33
1
1
Salta
3047558479
1982-09
CMNC
A. Martinez
Argentina
-26.516666
Pampa del Infierno
1241
-61.166668
32
33
1
1
Chaco
3047558367
2013-10-16
2013-10-17
2013-10-16
M. Ibarra Polesel
Argentina
-27.509972
San Fernando
1
-59.079887
Estacion San Francisco
32
33
1
Chaco
3047558374
2014-12-22
M. Ibarra Polesel
Argentina
-27.441668
Departamento Primero de Mayo
1
-58.85765
Antequera
32
33
1
Chaco
3047558353
[487,1350,582,609]
NHMB
El Pinto
Argentina
Santiago del Estero
32
33
2
2
Santiago del Estero