A revision of male ants of the Malagasy region (Hymenoptera: Formicidae): Key to subfamilies and treatment of the genera of Ponerinae. Yoshimura, M. Fisher, B. L. Zootaxa 2007 1654 21 40 623T5 36668 Insecta Formicidae Animalia Hymenoptera 28 Arthropoda family  A few genera are divided into parts, reflecting distinct characters sets outlined in Table 1. 1. Two distinct, long, narrow spines or lobes present on apical portion of abdominal sternum IX (Fig. 3a) or, if absent, then mandibles extremely elongated, distinctly longer than head, and volsella massive, claw-shaped, directed dorsally. Pygostyles absent........................................................... Cerapachyinae - Spines or lobes absent on apical portion of abdominal sternum IX or the apical portion bilobed, with each lobe very wide (Fig. 3b). Mandibles not elongated, distinctly shorter than head. Volsella moderate, not claw-shaped, not directed dorsally. Pygostyles present or absent......................................................2 2. Abdominal segment III much smaller than segment IV in lateral view (Fig. 3c)....................................3 - Abdominal segment III nearly as large as segment IV in lateral view (Fig. 3d)......................................4 3. Hind tibia with two spurs (Fig. 3e)......................................................................... Pseudomyrmecinae - Hind tibia with one spur or without spurs (Fig. 3f) ............................................................. Myrmicinae 4. Anal region of hind wing vestigial. Oblique mesopleural furrow reaching pronotum close to its posteroventral corner (Fig. 4a)...................................................................................................... Proceratiinae - Anal region of hind wing well developed. Oblique mesopleural furrow not reaching pronotum, its anterior termination well separated from the pronotum (Fig. 1a)..................................................................5 5. Petiole (abdominal segment II) broadly and dorsally attached to abdominal segment III; dorsal constriction between petiole and abdominal segment III very shallow or indistinct in lateral view (Fig. 4b......... ........................................................................................................................................ Amblyoponinae - Petiole (abdominal segment II) narrowly and ventrally attached to abdominal segment III; dorsal constriction between petiole and abdominal segment III deep in lateral view (Fig. 6a)...6 6. Scuto-scutellar suture usually longitudinally sculptured. Forewing clearly with cross vein 2rs-m (Fig. 1b); if vein weak then at least with vestigial branches on Radial sector and Media. Scape short, not reaching posterior margin of head in full-face view (Fig. 5c). Constriction between abdominal segments III and IV present in some cases.............................................................................................. Ponerinae - Scuto-scutellar suture not longitudinally sculptured. Forewing usually without any trace of cross vein 2rs-m (Fig. 4c). Scape short (Fig. 4d) or long (Fig. 2a). Constriction between abdominal segments III and IV absent...........................................................................................................................................7 7. Many minute, serrate teeth present on masticatory margin of mandible (Fig. 4d), or, if teeth absent, then scape not reaching posterior margin of head in full-face view........................................ Dolichoderinae - Several larger teeth present on masticatory margin of mandible (Fig. 2a). Scape long, distinctly exceeding posterior margin of head in full-face view (Fig. 2a)........................................................ Formicinae