Myrtus goetheana Martius ex De Candolle (1828: 240) Taxonomic study of Myrtaceae in forest fragments in Amazonian Maranhão, Brazil Amorim, Gabriela Dos Santos Barbosa, Maria Regina De Vasconcellos Jr, Eduardo Bezerra De Almeida Thomas, William Wayt Phytotaxa 2022 2022-10-11 568 1 27 60 Q5S9 2054440 (Mart. ex DC.) O. Berg O. Berg DC. [165,705,785,812] Magnoliopsida Myrtaceae Calycolpus Plantae Myrtales 4 31 Tracheophyta species goetheanus  Basionym:—  Myrtus goetheanaMartius ex De Candolle (1828: 240).   Type:— BRAZIL. “Habitat in Brasiliae prov. Rio Negro”,  Martius2762(M; lectotypedesignated by Landrum 2010: 381). ( Fig. 2A–B.)  Trees 1– 12 m. Twigs glabrous or glabrescent, the indumentum whitish or reddish-brown. Leaves with petioles 1–7 mmlong, glabrous or glabrescent, unchanneled or shallowly channeled; blades elliptic to broadly so, lanceolate, ovate or obovate, 3.3–14.4 × 2.4–5.5 cm, glabrous, membranaceous or chartaceous, discolorous; glands conspicuous; base cuneate, acute or rounded; apex acuminate, rarely attenuate or cuneate; midvein adaxially sulcate or impressed; lateral veins moderately visible, not arched, 10 to 20 pairs; intra-marginal vein 1–2 mmfrom the margin, margin flat. Inflorescence axillary or terminal, fascicle, the axis ca. 3 mmlong, sometimes inconspicuous, glabrous; bracts deltate, 0.3–0.5 × 0.6 mm, puberulent adaxially, glabrous abaxially, persistent in the fruit; pedicels 9–40 mmlong, glabrous; bracteoles triangular to narrowly so, or deltate, 0.5–2 × 0.2–0.7 mm, glabrous, persistent in the fruit; flower buds globose, 7 × 12 mm, not constricted above ovary; calyx-lobes free, five, narrowly triangular, deltate to broadly so, 5–10 × 3–6 mm, glabrous, not reflexed, persistent in the fruit; ovary 3-5-locular. Immature fruits subglobose or globose, 0.4–0.7 × 0.5–1.1 cm, glabrous, surface smooth.   Note:—  Calycolpus goetheanusmay be recognized by its glabrous leaves, inflorescence in fascicles; buds with five free calyx-lobes (5–10 × 3–6 mm); ovary 3–5-locular; and glabrous, subglobose or globose fruits.   Distribution and habitat:—Known from Colombia, French Guiana, Guyana, Suriname, Trinidad-Tobago, Venezuela, northeastern and northwestern Brazil(Amazonas, Maranhão, Pará and Roraimastates) ( Landrum 2010, Flora do Brasil2020). In Amazonian Maranhão,  C. goetheanuswas found growing along a road, and in both “terra firme’’ and flooded forests.   Phenology:—Flowering in January, February, May and October to December, and fruiting in January, February and October.   Specimens examined:—  BRAZIL. Maranhão: Alcântara, Piriuaçú, 02°24 32” S, 44°24’53” W,  11 April 1954(st.),  R.L. Fróes30771( IAN!).  Cururupu, Maracassumé River Region, Campo de Boa Esperança, 02°02’29” S, 45°57’19” W,  22 October 1932(fl., fr.),  R. Fróes1977( NY!).  São José de Ribamar, Sítio Aguahy, 02°38’47” S, 44°09’05” W,  28 October 2017(fl.),  G.S. Amorim378( MAR!); idem,  20 February 2018(fr.), G.S. Amorim467( MAR!).  São Luís, Alumar, 02º41’30” S, 44°9’44” W, 29 November994 (fl.),  N. Figueiredo7 ( MAR!);  02°41’30” S, 44°19’44” W,  18 January 1995[fl], N. Figueiredo4( MAR!);  02°41’30” S, 44°19’44” W,  13 November 1995(fl.),  K.B. Ferreira& E.C. Gimoss.n.( MAR!);  Estradaque vai do Rio Anilpara Maioba,  10 January 1950(fl.), R.L. Fróes25652(U, image); Reserva Florestal do Sacavém, 02º34’04” S, 44º15’44” W,  03 December 1988(fl.),  R.C. Reis57( MAR!);   22 January 1992[fl],  F.H. Muniz17(RB);   26 February 1992[fl],  F.H. Muniz196(HUEFS, image); 02°35’59” S, 44°16’34” W,  27 May 1992(fl.),  F.H. Muniz256(HUEFS, image);  02°30’ S, 44°15’ W,  21 January 1993(fr.),  F.H. Muniz287(INPA, image);  Rio Anil, 02°28’59” S, 44°12’00” W,  13 January 1951(fl., fr.),  B.E. Franz26348(ASU, image);  Sede da Cemar, 02°31’47” S, 44°18’10” W,  27 November 1985(fl.),  G.T. Prance& R. Henriques29886( US, image);  Sítio Santa Eulália, 02°30’30” S, 44°16’39” W,  29 January 2018(fl.),  G.S. Amorim414, 419( MAR!);  02°30’52” S, 44°16’34” W,  26 February 2018(fr.), G. S. Amorim491( MAR!). 3923982565 Martius Brazil Rio Negro 4 31 1 Rio Negro lectotype 3923982422 1954-04-11 IAN R. L. Froes Brazil -2.408889 Piriuacu 21 -44.414722 Alcantara 4 31 1 Maranhao 3923982350 1932-10-22 NY R. Froes Brazil Cururupu -2.0413888 Campo de Boa Esperanca 21 -45.95528 Maracassume River Region 4 31 1 3923982551 2017-10-28 MAR G. S. Amorim Brazil -2.6463888 Sitio Aguahy 21 -44.15139 Sao Jose de Ribamar 4 31 1 3923982363 [700,1333,1701,1727] 2018-02-20 MAR Amorim Brazil -2.6463888 Sitio Aguahy 21 -44.15139 Sao Jose de Ribamar 4 31 1 3923982336 MAR November & N. Figueiredo Brazil -2.6916668 Alumar 21 -44.162224 Sao Luis 4 31 1 3923982393 1995-01-18 MAR Figueiredo Brazil -2.6916668 21 -44.328888 4 31 1 3923982321 1995-11-13 MAR K. B. Ferreira & Gimos Brazil -2.6916668 21 -44.328888 4 31 1 3923982377 1950-01-10 1988-12-03 1950-01-10 MAR R. C. Reis Brazil Estrada -2.5677776 Maioba 21 -44.262222 Rio Anil 4 31 1 3923982417 1951-01-13 1993-01-21 1951-01-13 F. H. Muniz United States of America -2.5297222 Sede da Cemar 21 -44.302776 Rio Anil 4 31 1 3923982420 1951-01-13 1993-01-21 1951-01-13 F. H. Muniz United States of America -2.5297222 Sede da Cemar 21 -44.302776 Rio Anil 4 31 1 3923982548 1951-01-13 1993-01-21 1951-01-13 F. H. Muniz United States of America -2.5297222 Sede da Cemar 21 -44.302776 Rio Anil 4 31 1 3923982411 [314,1390,1947,1974] 1951-01-13 1993-01-21 1951-01-13 B. E. Franz United States of America -2.5297222 Sede da Cemar 21 -44.302776 Rio Anil 4 31 1 3923982602 1951-01-13 1993-01-21 1951-01-13 G. T. Prance & R. Henriques United States of America -2.5297222 Sede da Cemar 21 -44.302776 Rio Anil 4 31 1 3923982537 [136,1290,2018,2044] 2018-01-29 MAR G. S. Amorim United States of America -2.5083334 Sitio Santa Eulalia 21 -44.2775 4 31 1 3923982347 2018-02-26 MAR Amorim United States of America -2.5144444 21 -44.27611 4 31 1