V. limnense V. sinuatum Typification of the names Verbascum limnense and Celsia tomentosa (Scrophulariaceae) and a new nothospecies, V. × sipiadense, with the hybrid formula V. limnense × V. sinuatum Zografidis, Aris Esser, Hans-Joachim Dimopoulos, Panayotis Raus, Thomas Phytotaxa 2022 2022-04-05 542 2 214 220 Zograf. Zograf. [189,562,1486,1513] Magnoliopsida Scrophulariaceae Verbascum true Plantae Lamiales 2 216 Tracheophyta species sipiadense  (  V. limnense×  V. sinuatum), nothosp. nov.   — Type: GREECE. Nomos Evvias, Eparchia Chalkidos: Island Evvia, just outside the settlement of Sipiada(near Limni) on roadsides, 38°46’N, 23°16’E,  30 m,  5 May 2018,  Zografidis360( holotype UPA!;  isotype ATH!). Fig. 3.   Diagnosis: Intermediate between the parental species  V. limnenseand  V. sinuatumbut sterile. Differing from the former by having the lamina of basal leaves coarsely crenate to lobed (vs. crenate to subentire), leafy flowering stems (vs. subaphyllous), up to 4-flowered glomerules (vs. solitary flowers), bracteoles present (vs. bracteoles absent) and bicoloured, white-violet stamen hairs (vs. white). Differing from the latter by the presence of long eglandular hairs at the base of the petioles of rosette leaves, 2- to 7-flowered glomerules (vs. 1- to 4-flowered), glabrous outer surface of corolla (vs. stellate-tomentose) and bicoloured stamen hairs (vs. violet).   Description: Sterile biennial herb producing a basal leaf rosette in the first year of growth and an erect, terete, leafy flowering stem 1–1.8 mof height in the second year. Indumentum covering the aerial part of the plant eglandular, grayish-yellowish, persistent. Rosette-leaves petiolate with petioles 1–8 cmlong, base of petioles with longer hairs; lamina obovate to obovate-elliptic, coarsely crenate to lobed, attenuate to petiole, obtuse at apex, 7–30 × 2.5–10 cm. Lower cauline leaves similar but ±sessile. Middle and upper cauline leaves sessile, progressively smaller, ovate-oblong to ovate-triangular, crenulate, subacute to acute, the upper ones ca. 2 × 1 cm. Inflorescence freely branched from below, lax. Flowers solitary or in groups (glomerules) of 2–4; bracts c. 2 × 1 mm, ovate-lanceolate, acute; bracteoles present, lanceolate; pedicels 1–3 mm. Calyx 2–3 mm, segments lanceolate-oblong, acute, divided almost to the base. Corolla yellow, 2–2.5 cmin diameter, with pellucid glands, outer surface glabrous, inner surface glabrous or with simple hairs on the proximal part of upper lobes (on the central vein of the lobe); stamens 4(–5) with medifixed anthers; filament hairs white and violet. Capsules not formed.   Notes: Natural hybrids between  Verbascumspp.are readily produced when species coexist, whereas all hitherto described hybrid taxa are invariably sterile (e.g., Murbeck 1933, Huber-Morath & Rechinger 1960). It is also noted that, unlike V. × sipiadense, the parental species  V. limnensealways exhibits white stamen hairs, not violet ones as cited in former literature ( Ferguson 1972, Trigas 2003, Strid 2016a). It is thus hypothesized that the various descriptions of  V. limnensein previous floras might have been partially based on heterologous material consisting of the new nothospecies. 3747628303 2018-05-05 UPA Zografidis Greece Sipiada 30 38.766666 Nomos Evvias 1173 23.266666 Island Evvia 2 216 1 holotype 3747628305 [721,961,1558,1584] 2018-05-05 ATH Greece Sipiada 30 38.766666 Type 1173 23.266666 Island Evvia 2 216 1 isotype