Macrurus altipinnis
Macrourus nasutus
Macrourus rudis
Macrourus proximus
Lionurus proximus : Gilbert & Hubbs 1916:201
Nezumia spinosa
Lionurus abei Matsubara, 1943:146
Nezumia proximus : Okamura 1970a:94
Nezumia proxima : Okamura 1970b
Lionurus spinosus
Grenadiers (Teleostei: Gadiformes: Macrouridae) of Japan and adjacent waters, a taxonomic monograph
Nakayama, Naohide
Megataxa
2020
2020-11-05
3
1
1
383
47C4L
: Okamura
1970
[136,355,887,913]
Actinopterygii
Macrouridae
Nezumia
Animalia
Gadiformes
249
250
Chordata
species
proxima
[Japanese name: Higo-sokodara]
( Figs. 168–170; Table 12; Appendix 3-10C)
Macrurus altipinnis(not G̹nther 1877): G̹nther 1887:138 (brief description; in part, BMNH 1887.12.7.92; Enshu-nada). Macrourus nasutus(not G̹nther 1877): Jordan& Gilbert in Jordan& Starks 1904:614(in part, 1 spec.from off Izu; see Gilbert & Hubbs 1916:201). [?] Macrourus rudis(not G̹nther 1887): Franz 1910:16( 1 spec.from Aburatsubo); Jordan et al.1913:417(listed; Japan; new Japanese name: “Aka-sokodara”). Macrourus proximus Smith& Radcliffe in Radcliffe, 1912:119, pl. 26, fig. 2 [original description; holotype: USNM 72936, from Sogod Bay, Philippines, Albatrosssta. 5202, 10º12ʹ00ʺN, 125º04ʹ10ʺE, in 502 ftm ( 918 m)]. Lionurus proximus: Gilbert & Hubbs 1916:201[new combination; brief description; in part, 10 spec.from East ChinaSea, Hyuganada, off Cape Shionomisaki, and Enshu-nada; 1 spec.from Albatrosssta. 4918 (CAS-SU 22941) is Nezumia spinosa]; Okada & Matsubara 1938:453(in key; Japan); Matsubara 1955:1316(in key; Japan); Matsubara 1965:509(compiled; Japan; new Japanese name: “Higo-sokodara”). Lionurus abeiMatsubara, 1943:146, figs. 7–8 (original description; holotype: FAKU 4909, from Kumano-nada; new Japanese name: “Kasumi-hige”); Matsubara 1955:1317, pl. 134, fig. 458 (in key; Japan); Okada et al.1959:83(listed; Kumanonada); Matsubara 1965:509(compiled; Japan); Kuroda 1966:29(brief description; spec. from Suruga Bay). Nezumia proximus: Okamura 1970a:94, pl. XXI, text-fig. 41 (description; biological notes; 17 spec.from Pacific off southern Japanfrom Choshi to Kumano-nada and East ChinaSea); Kuroda 1971:56[listed from Suruga Bay after Okamura (1970a)]. Nezumia proxima: Okamura 1970b: table 1 (listed; Japan); Kataoka & Tomida 1981:78(listed; Mie Pref.); Tominaga & Uyeno 1981:489(listed; Japan); Okamura 1982:163, 351, fig. 162 (brief description; 6 spec.from Tosa Bay; photo based on BSKU 29500); Sawada 1983:107, 192, fig. 58 [brief description; 4 spec.from Pacific off Tohoku; photo based on HUMZ spec. (catalog no. unknown)]; Ohta 1983: table A (listed; Suruga Bay); Okamura 1984a:217, 362, fig. 152 (brief description; 8 spec.from OkinawaTrough; photo based on BSKU 29792); Okamura 1984b:94, pl. 81, fig. I (compiled); Okamura 1988:94, pl.81, fig.I (compiled); Iwamoto 1990:281, fig. 645 (synopsis); Nakabo 1993:360(in key; Japan); Hori 1996:27(listed; Ibaraki Pref.); Shinohara et al.1996:170( 2 spec.listed from Pacific off Tohoku); Okamura 1997:128, fig. 17 (compiled); Shinohara & Matsuura 1997:292(listed; Suruga Bay); Funabashi 1998:85(listed; Ibaraki Pref.); Nakabo 2000:424(in key; Japan); Shinohara et al.2001:306( 3 spec.listed from Tosa Bay); Nakabo 2002:424(in key; Japan); Yoda et al.2002:11(listed; East Chinaand Yellow Seas); Shinohara et al.2005:418(listed; Ryukyu Islands); Suetsugu & Ohta 2005: table 3 (listed; Enshu-nada); Senou et al.2006:421(listed; Sagami Sea); Shao et al. 2008b: table 2 ( 2 spec.listed from eastern Taiwanand South ChinaSea; first Taiwanese record); Kitagawa et al.2008:40, unnumbered fig. (brief description; spec. from Pacific off Tohoku); Shinohara et al.2009:709( 1 spec.listed from Pacific off Tohoku); Nakabo & Kai 2013:500(in key; Japan); Iwamoto et al.2015:93, fig. 19 (brief description; 3 spec.from northeastern and southwestern Taiwan, and South ChinaSea); Amaoka et al.2020:159, fig. 213 (erroneously listed from Hokkaido); Motomura 2020:39(listed; Japan). [?] Lionurus spinosus(not Gilbert & Hubbs 1916): Maruyama 1971:33(listed with a question mark; Iwate Pref.).
Diagnosis.A species of Nezumiawith 8–10 pelvic-fin rays. Snout moderately long, conical, protruding well beyond upper jaw, length 29–33% HL, ventral contour oblique in lateral view; underside of snout broadly naked; ventral surfaces of head scaled posterior to vertical through midorbit; mandibular rami mostly scaled. Second spinous ray of first dorsal fin not extremely prolonged (height of fin 92–118% HL). Body scales covered with short, greatly reclined, lanceolate spinules in tightly packed convergent rows or densely scattered over exposed portion; tip of last spinule in each row extending well beyond posterior scale margin; middle row of spinules not enlarged; scales below second dorsal-fin origin 6–8.5. Anterior dermal window of light organ (ADW) situated between inner pelvic-fin bases. Cephalic sensory pores present, but barely discernible. Broad dark band encircling trunk (often faint or absent in preserved specimens); first dorsal fin uniformly dark.
Material examined. 34 specimens. Japan: BSKU 99112(1, 50.4 mmHL, 314+ mm TL), BSKU 99117(1, 59.5 mmHL, 357+ mm TL), off Pacificcoast of Tohoku, FRV Wakataka-maru, sta. 32 (F-800), coll. H. Endo, date unknown; BSKU 26667(1, 61.7 mmHL, 336+ mm TL), west of Yaku-shima Island, Okinawa Trough, 30.0300ºN, 128.3650ºE, 900 m, F/ V Yuryo-maru, No.8, tr. 16-1, bottom trawl, coll. Y. Kinoshitaand S. Hagino, 3 Feb. 1978; BSKU 86874(1, 31.4 mmHL, 189+ mm TL), off Nobeoka, Hyuga-nada, 32.5163ºN, 132.2175ºE, 997–1030 m, R/ V Tansei-maru, cr. KT-99-18, sta. BT-2, beam trawl, coll. H. Endoand S. Nagatomo, 16 Dec. 1999; BSKU 44837(1, 54.2 mmHL, 319+ mm TL), Tosa Bay, 32.9886ºN, 133.5978ºE, 700–720 m, FRV Kotakamaru, otter trawl, 21 Jun. 1988; BSKU 98937(1, 33.3 mmHL, 197+ mm TL), BSKU 98938(1, 47.6 mmHL, 218+ mm TL), BSKU 98939(1, 28.4 mmHL, 147+ mm TL), Tosa Bay, 33.0452ºN, 133.6448ºE, 733–740 m, FRV Kotaka-maru, otter trawl, 25 Jan. 1989; BSKU 101591(1, 42.1 mmHL, 254+ mm TL), BSKU 101592(1, 49.2 mmHL, 303+ mm TL), BSKU 101593(1, 48.0 mm HL, 280+ mm TL), BSKU 101594(1, 27.9 mmHL, 176+ mm TL), BSKU 101595(1, 33.3 mmHL, 203+ mm TL), BSKU 101596(1, 39.9 mmHL, 229+ mm TL), BSKU 101597(1, 40.9 mmHL, 253+ mm TL), BSKU 101598(1, 35.2 mmHL, 196+ mm TL), BSKU 101599(1, 49.9 mmHL, 302+ mm TL), Tosa Bay, 700 m, FRV Kotaka-maru, otter trawl, 22 May 1989; BSKU 47166(1, 37.6 mmHL, 233+ mm TL), off Tosa Bay, 32.8953ºN, 133.6777ºE, 996– 1010 m, R/ V Tansei-maru, cr. KT-89-06, sta. T1, beam trawl, 8 Nov. 1989; BSKU 23147(1, 45.1 mmHL, 274+ mm TL), Tosa Bay, 33.0350ºN, 133.7500ºE, 945 m, FRV Kaiyo-maru, cr. SK-75-2, sta. 18, tr. 3, bottom trawl, 26 Jan. 1975; BSKU 23076(1, 46.7 mmHL, 225+ mm TL), BSKU 23181(1, 48.1 mmHL, 291 mmTL), off Kochi, 32.9800ºN, 133.8550ºE, 1043 m, FRV Kaiyo-maru, cr. SK-75-2, sta. 19, tr. 2, bottom trawl, 26 Jan. 1975; * KPMNI28856(1, 44.8 mmHL, 236+ mm TL), ca. 24 kmwest-southwest of Shirahama, Wakayama, 33.5795ºN, 135.0995ºE, 532 m, R/ V Tansei-maru, cr. KT-10-16, sta. YK-2, box corer, coll. T. Sato, 15 Aug. 2010; BMNH 1887.12.7.92 (1, 24.0 mm HL, 149+ mm TL), off Cape Omaezaki, Enshu-nada, 34.1167ºN, 138.0000ºE, 565 ftm ( 1034 m), Challengersta. 235, trawl, 4 Jun. 1875; BSKU 19762(1, 41.6 mmHL, 211+ mm TL), BSKU 19764(1, 57.9 mmHL), Sagami Bay, 35.1167ºN, 139.4500ºE, 730– 790 m, FRV Soyo-maru, sta. T7, 23 Nov. 1966; BSKU 20250(1, 39.2 mmHL, 254+ mm TL), west of Misaki, Sagami Bay, 35.1283ºN, 139.4983ºE, 450 m, FRV Soyomaru, sta. 77, 30 Jun. 1972; BSKU 14184(1, 35.7 mmHL, 221+ mm TL), BSKU 14186(1, 29.2 mmHL, 190+ mm TL), off Hiratsuka, Sagami Bay, 35.2522ºN, 139.5178ºE, 510–600 m, R/ V Tansei-maru, coll. O. Okamura, 12 Apr. 1969; BSKU 45547(1, 43.8 mmHL, 264+ mm TL), east of Katsuura, 35.1400ºN, 140.8125ºE, 507–578 m, R/ V Tansei-maru, cr. KT-89-03, sta. TB5, 13 Mar. 1989; BSKU 101193(1, 60.3 mmHL, 374+ mm TL), off Hitachi, 900 m, FRV Wakataka-maru, sta. H, coll. H. Endo, 10 Oct. 1997; BSKU 97917(1, 56.7 mmHL, 345+ mm TL), off Oshika Peninsula, 38.5318ºN, 142.3370ºE, 772–783 m, FRV Wakataka-maru, sta. 32, 28 Oct. 1995; BSKU 97906(1, 58.7 mmHL, 375+ mm TL), BSKU 97910(1, 51.4 mmHL, 341+ mm TL), off Otsuchi, 39.1444ºN, 142.3748ºE, 847–935 m, R/ V Tansei-maru, cr. KT-89-03, sta. SR89, 10 Mar. 1989. Uncertainvariant: BSKU 19891( 50.8 mmHL, 274+ mm TL), east-southeast of Miyake-jima Island, Shichito-Iojima Ridge, Japan, 33.9183ºN, 140.0083ºE, 860–870 m, FRV Soyo-maru, beam trawl, 1 Dec. 1968.
Description of Japanese specimens.General features are shown in Fig. 168. Counts and measurements are given in Table 12. Body long, slender, gradually tapering from first dorsal-fin origin to end of tail. Trunk short, laterally compressed, width over pectoral-fin bases 1.3–1.7 indepth below first dorsal-fin origin. Head short, HL about 5.2–6.6 inTL, its dorsal profile almost straight from tip of snout to nape. Snout moderately long, protruding well beyond upper jaw, conical in lateral view, bluntly pointed in dorsal view; snout length 0.9–1.1 times as long as orbit diameter; ventral contour oblique, forming angle of about 45º to horizontal axis of head and body; tip of snout situated on horizontal through midorbit. Orbit moderately large, circular, greatest diameter 1.1–1.9 inpostorbital length. Interorbital space almost flat, width 1.3–1.7 inorbit diameter. Mouth small, inferior, upperjaw length 0.9–1.2 inorbit diameter; posterior margin of maxilla extending beyond vertical through midorbit; lateral corner of mouth slightly restricted by lip folds; lips thick, fleshy, papillose near tooth bands. Suborbital region narrow, divided into upper and lower halves by infraorbital ridge. Preopercle large, broadly rounded ventrally, posterior margin slightly inclined from vertical; preopercular ridge low, poorly marked, forming slight backward extension at angle. Interopercle narrowly exposed beyond preopercle. Gill membranes broadly connected across, and attached mesially to isthmus, with broad posterior free fold. Anteroventral end of gill opening not reaching below posterior margin of lower jaw. Outermost gill slit moderately restricted by skin folds, length 2.0– 2.8 inorbit diameter. Gill rakers small, tubercular, armed with short, fine spines distally; those on outer side of first arch and inner side of fourth arch much smaller than others. Gill filaments moderately long. Barbel well developed, length 1.4–2.1 inorbit diameter. FIGURE 170.Uncertain variant of Nezumia proxima. BSKU 19891, 50.8 mm HL, 274+ mm TL, east-southeast of Miyake-jima Island, Shichito-Iojima Ridge, Pacific, 860–870 m depth. (A) Lateral view; (B) dorsal and (C) ventral views of head. Preserved condition. [Photos: N. Nakayama] Anus closer to pelvic-fin bases than to anal-fin origin. Periproct broad, oval in shape. Anterior dermal window of light organ small, circular, situated between inner bases of pelvic fins. Teeth short, slender, sharp, in broad tapered bands in both jaws. Premaxillary teeth arranged in about 5 tooth rows near symphysis, with outermost series distinctly enlarged. Mandibular band similar to that of premaxillary, with about 6 tooth rows at widest point near symphysis, but none significantly enlarged. Posterior ends of tooth bands falling short of lateral corner of mouth in both jaws. All teeth deeply embedded in thick layer of gum papillae. Body scales small, thin, moderately deciduous, covered with short, greatly reclined, lanceolate spinules in tightly packed convergent rows or densely scattered over exposed portion ( Fig. 169A–B); tip of last spinule in each row extending well beyond posterior scale margin; middle row of spinules not enlarged; buttresses of spinules absent; reticulate structure narrowly developed on posterior parts of unexposed portion. Body fully scaled except for fins, periproct, and dermal window of light organ; pelvic fins mostly naked, but basal portions covered with tiny scales. Suborbital shelf formed of 2 rows of stout, coarsely modified scales. Snout tipped with 2 enlarged, thickened, closely adjoined tubercles; scales on lateral angles of snout modified into scute-like scales. Scales on head generally similar to those on body, except that spinules more erect and arranged in subparallel rows. Dorsal and lateral surfaces of head almost fully scaled; membrane around anterior nostril and paired Y-shaped areas dorsally posterior to leading edges of snout narrowly naked. Underside of snout almost completely naked; lower half of suborbital region mostly scaled, but narrowly naked above upper lip; anterior 1/3–1/2 of mandibular rami naked; gular and branchiostegal membranes lacking scales. TABLE 12.Morphometric measurements and meristic counts* for Nezumia proxima. Type status NT Identification Typical specimens Uncertain variant Number of specimens n= 32 n= 1 Measurements (mm) TL 147+–375+ 274+ HL 24.0–61.7 50.8 % of HL (% of PRL) Snout length 29–33 (39–46) 31 (44) Orbit diameter 28–35 (40–48) 29 (40) Postorbital length 37–45 (52–61) 44 (62) Postrostral length 70–75 (–) 71 (–) Orbit–preopercle distance 32–40 (44–54) 35 (50) Suborbital width 13–16 (18–22) 14 (20) Upper-jaw length 28–34 (39–47) 31 (44) Length of rictus 23–28 (32–38) 27 (38) Length of PMX tooth band 16–20 (22–28) 20 (28) Preoral length 22–30 (30–42) 25 (35) Dist. tip and lat. angle of snout 16–20 (21–28) 16 (23) Snout width 25–31 (34–42) 26 (37) Internasal width 19–23 (27–32) 24 (34) Interorbital width 19–24 (27–32) 25 (35) Body width over P. bases 43–70 (61–94) 59 (83) Body depth at 1D. origin 70–92 (97–124) 79 (111) Body depth at A. origin 58–92 (80–124) 70 (98) Pre-V. length 103–124 (144–174) 114 (161) Pre-anus length 121–145 (169–200) 134 (188) Pre-A. length 146–176 (200–241) 157 (220) Isthmus–V. distance 32–45 (44–62) 33 (46) Isthmus–anus distance 46–69 (64–95) 51 (72) Isthmus–A. distance 72–107 (99–147) 76 (107) V.–A. distance 39–64 (55–89) 45 (63) Anus–A. distance 21–38 (30–53) 26 (36) V. length 44–71 (59–99) 69 (97) P. length 49–61 (69–84) 56 (79) Pre-1D. length 108–125 (147–173) 114 (160) Height of 1D. 92–118 (126–162) 97 (136) Length of 1D. base 23–33 (32–46) 24 (33) Interdorsal length 31–66 (44–92) 44 (62) Length of gill slit 11–16 (15–21) 15 (21) Length of posterior nostril 2–6 (3–8) 9 (12) Barbel length 15–21 (20–29) 15 (21) Counts 1D. rays II,8–10 II,9 P. rays i17–i21 i19 V. rays 8–10 9 Outer GR on first arch 5–9 7 Inner GR on first arch 8–10 10 Outer GR on second arch 8–9 9 Inner GR on second arch 8–10 9 Longitudinal scales 32–40 39 Scales below 1D. origin 7.5–10 9 Scales below 1D. midbase 5–8 5.5 Scales below 2D. origin 6–8 8.5 Scales above A. origin 19–26 21.5 ... See the next page for the table footer *Abbreviations: 1D.—first dorsal fin; 2D.—second dorsal fin; A.—anal fin; GR—gill rakers; NT—non-type; P.—pectoral fin; PMX—premaxillary; V.—pelvic fin Cephalic sensory pores present, arranged along sensory canals; those on underside of head very slightly enlarged, more or less tubular. Free neuromasts small, tubercular, densely scattered on underside of snout and mandibular rami; similar but much smaller neuromasts serially arrayed along leading edges of snout, supranarial ridges, suborbital shelf, and supraorbital canals. Grooved lateral line incomplete anteriorly, occurring as short interrupted segments, but complete posterior to vertical through midbase of first dorsal fin. Origin of first dorsal fin slightly posterior to vertical through pectoral- and pelvic-fin bases, the latter two on about same vertical; first dorsal fin high, its height 3.2–4.7 times as long as its base length; second spinous ray long, but not especially elongate, armed along its leading edge with short, reclined, conical denticles; its tip extending to bases of 4th–20th second dorsal-fin rays when laid back. Second dorsal fin well separated from first dorsal, originating above 5th–13th anal-fin rays; interdorsal space 1.1–2.4 times as long as first dorsal-fin base length. Outermost pelvic-fin ray slightly prolonged, hair-line distally, its tip usually extending beyond anal-fin origin when laid back. Color when fresh( Fig. 168). Uniformly dark brown; broad dark band encircling trunk (especially prominent when body scales mostly missing; Fig. 168C). Color in alcohol.Similar to that in fresh specimens; head and body dark brown overall, with broad dark band encircling trunk (band often indistinct in old specimens); abdomen and gill cover prominently darker in smaller specimens; lips dark; oral cavity blackish except for pale gums; gill cavity mostly black, but paler on ceratohyal, epihyal, and anterior to lower end of cleithrum; gill rakers and arches dark dusky, filaments pale; chin barbel varying from pale to dark; gular membrane blackish, but abruptly pallid laterally; branchiostegal membranes black, especially along distal margins; fins uniformly dark. Size. To about 40 cmTL ( NSMT-P 94347, 398+ mm TL, off Hitachi, Ibaraki Pref., Japan).
Distribution.Distributed in the western Pacific from the Philippinesto Japan( Radcliffe 1912; Gilbert & Hubbs 1920; Shao et al.2008b; this study). In Japanese waters, known from off the Pacific coasts northward to Kuji ( 40.19ºN; Iwate Pref.) and the OkinawaTrough at depths of 361‾ 1348 m(Appendix 3-10C). Common, especially in the Pacific off Honshu north of Choshi ( 35.74ºN; Chiba Pref.).
Remarks. Nezumia proximawas originally described from the Philippines, and first recorded from Japanby Gilbert & Hubbs (1916). Matsubara (1943) described N. abei(as a species of LionurusG̹nther, 1887) from a single specimen collected from the Kumano-nada, Japan, but this species has been regarded as a junior synonym of N. proxima(e.g., Okamura 1970a; Iwamoto 1990; Iwamoto et al.2015). Unfortunately, the holotypeof N. abeiis presumed to be lost (see the Comments on holotypeof Coelorinchus hige). Among the FAKU collection was a single specimen of N. proximacollected from off Minamidaito-jima Island (FAKU 13366, 1 spec., 59.4 mmHL, 366+ mm TL), but the register indicates its locality with a question mark. This record was not included in the distribution of this species (Appendix 3-10C). Amaoka et al.(2020)recently listed the species from Hokkaido, but it is obviously based on Sawada’s (1983)record from the Pacific off Tohoku. Relationships and comparisons. Nezumia proximais putatively most similar to N. namatahi McCann & McKnight, 1980, a species known only from New Zealandand Australia, and they share the combination of 8–10 pelvic-fin rays ( 9–10 in N. namatahi), a broad naked area on the underside of the snout, a long barbel (15–21% HL in N. proxima, and 13–30% in N. namatahi), body scales covered with lanceolate to shield-shaped spinules in parallel to convergent rows, and a broad, prominent, dark band completely encircling the trunk. However, N. proximareadily differs from that species in that the anterior dermal window of the light organ (ADW) is situated between the inner pelvic-fin bases (vs. ADW usually distinctly posterior to a line connecting inner pelvic-fin bases). It further differs from N. namatahiin its tiny pores along the infraorbital and mandibular canals (vs. large and prominent). [Data for N. namatahiare based on Iwamoto & Williams (1999:194–195), Iwamoto & Graham (2001:489), and this study.] Uncertain variant.A specimen collected from the Shichito-Iojima Ridge at a depth of 860–870 m( Fig. 170; Table 12) was initially regarded as a different species from N. proximaby having a wider interorbital space (25% HL vs. 19–24%). It further differs from the typical N. proximaspecimens examined in having a wider internasal space (24% HL vs. 19–23%), a larger posterior nostril (9% HL vs. 2–6%), a relatively shorter barbel (15% HL vs. 15– 21%), and slightly more scales below the second dorsal-fin origin (8.5 vs. 6–8). However, no significant differences were evident in squamation, spinulation of body scales ( Fig. 169), and the number and arrangement of cephalic sensory pores. Although the above differences may warrant their specific separation, the paucity of materials from the Shichito-Iojima Ridge did not allow a definitive conclusion. Therefore, the specimen was tentatively identified as an uncertain variant of N. proxima.
3750954320
USNM
Smith & Radcliffe
Philippines
918
10.2
Sogod Bay
21
125.06944
Albatross
249
250
USNM 72936
1
holotype
3750954962
BSKU
H. Endo
Japan
Japan
249
250
BSKU 99112, BSKU 99117
34
3750955409
1978-02-03
BSKU
Y. Kinoshita & S. Hagino
900
30.03
Trough
6
128.365
Yaku-shima Island
250
251
BSKU 26667
1
Okinawa
3750955405
1999-12-16
BSKU
H. Endo & S. Nagatomo
1014
32.5163
Hyuga-nada
6
132.2175
Nobeoka
250
251
BSKU 86874
1
Okinawa
3750955479
1988-06-21
BSKU
Kotakamaru
710
32.9886
Tosa Bay
6
133.5978
250
251
BSKU 44837
1
Okinawa
3750955347
BSKU
250
251
BSKU 98937
1
Okinawa
3750954478
BSKU
250
251
BSKU 98938
1
Okinawa
3750955530
1989-01-25
BSKU
Kotaka-maru
737
33.0452
Tosa Bay
6
133.6448
250
251
BSKU 98939
1
Okinawa
3750955075
BSKU
250
251
BSKU 101591
1
Okinawa
3750955270
BSKU
250
251
BSKU 101592
1
Okinawa
3750954872
[357,906,1500,2070]
BSKU
250
251
BSKU 101593
1
Okinawa
3750954373
[917,1447,1499,1526]
BSKU
250
251
BSKU 101594
1
Okinawa
3750954638
[810,1362,1533,1560]
BSKU
250
251
BSKU 101595
1
Okinawa
3750954454
BSKU
250
251
BSKU 101596
1
Okinawa
3750954325
BSKU
250
251
BSKU 101597
1
Okinawa
3750954737
BSKU
250
251
BSKU 101598
1
Okinawa
3750954940
1989-05-22
BSKU
Kotaka-maru
700
Tosa Bay
250
251
BSKU 101599
1
Okinawa
3750955384
1989-11-08
BSKU
V Tansei-maru
1003
32.8953
Tosa Bay
6
133.6777
250
251
BSKU 47166
1
Okinawa
3750955258
1975-01-26
BSKU
Kaiyo-maru
945
33.035
Tosa Bay
6
133.75
250
251
BSKU 23147
1
Okinawa
3750955267
[932,1447,1907,1934]
BSKU
250
251
BSKU 23076
1
Okinawa
3750954740
1975-01-26
BSKU
Kaiyo-maru
1043
32.98
Kochi
6
133.855
250
251
BSKU 23181
1
Kochi
3750954390
2010-08-15
KPM-
V Tansei-maru & T. Sato
532
251
252
33.5795
Shirahama
6
135.0995
250
251
NI 28856
1
Wakayama
3750955158
1875-06-04
BMNH
Challenger
1034
34.1167
Enshu-nada
6
138.0
Cape Omaezaki
251
252
BMNH 1887.12
1
Wakayama
3750955089
BSKU
251
252
BSKU 19762
1
Wakayama
3750955433
1966-11-23
BSKU
Soyo-maru
760
35.1167
Sagami Bay
6
139.45
251
252
BSKU 19764
1
Wakayama
3750955111
1972-06-30
BSKU
Soyomaru
450
35.1283
Sagami Bay
6
139.4983
Misaki
251
252
BSKU 20250
1
Wakayama
3750955361
BSKU
251
252
BSKU 14184
1
Wakayama
3750954662
1969-04-12
BSKU
V Tansei-maru & O. Okamura
555
35.2522
Sagami Bay
6
139.5178
Hiratsuka
251
252
BSKU 14186
1
Wakayama
3750954914
1989-03-13
BSKU
V Tansei-maru
543
35.14
Katsuura
6
140.8125
251
252
BSKU 45547
1
Wakayama
3750955247
1997-10-10
BSKU
Wakataka-maru & H. Endo
900
Hitachi
251
252
BSKU 101193
1
Wakayama
3750955403
1995-10-28
BSKU
Wakataka-maru
778
38.5318
Oshika Peninsula
6
142.337
251
252
BSKU 97917
1
Wakayama
3750954668
BSKU
251
252
BSKU 97906
1
Wakayama
3750955157
1989-03-10
BSKU
V Tansei-maru
891
39.1444
Otsuchi
6
142.3748
251
252
BSKU 97910
1
Wakayama
3750954452
[943,1163,1601,1627]
Uncertain
251
252
1
Wakayama
3750955136
1968-12-01
BSKU
Soyo-maru
Japan
865
33.9183
Shichito-Iojima Ridge
6
140.0083
Miyake-jima Island
251
252
BSKU 19891
1
3750954459
NSMT-P
Japan
Hitachi
254
255
NSMT-P 94347, 398
1
Ibaraki