Anachemmis Chamberlin, 1919: 12
A. sober Chamberlin
Anachemmis sober Chamberlin, 1919: 13
Silva, 2003: 41
Anachemmis scaber
Roewer, 1955: 607
A Revision of the North American Spider Genus Anachemmis Chamberlin (Araneae, Tengellidae)
PLATNICK, NORMAN I.
UBICK, DARRELL
American Museum Novitates
2005
2005-05-11
3477
1
20
62BWS
Chamberlin
Chamberlin. In
1919
[797,1129,1335,1359]
Arachnida
Zoropsidae
Anachemmis
Animalia
Araneae
1
2
Arthropoda
genus
Anachemmis Chamberlin, 1919: 12( typespecies by original designation A. soberChamberlin).
DIAGNOSIS: The unusual character combination of three claws plus claw tufts (fig. 1) readily separates members of Anachemmisand its closest relatives from most other North American spiders. Specimens of Zorocrates, which are found from Arizonato Texasand south to Central America, have a tiny remnant of the third claw that is equally obscured by the distal hairs of the tarsal scopula, but Zorocratesspecieslack true claw tufts (Griswold et al., in press) and have a single retrolateral tibial apophysis on the male palp. Males of Anachemmisdiffer from those of Titiotusin having two rather than three or four prongs on the retrolateral tibial apophysis, and from those of Socalchemmisand Liocranoidesby having a flattened, bladeshaped ventral prong of the apophysis and by having the dorsal prong of the apophysis reduced to a small point. The female epigynum differs from those of Liocranoidesspeciesin lacking an anterior hood, from those of Titiotusspeciesin having a much smaller septum, and from those of Socalchemmisspeciesin having Liocranoideslike anterior vulval elaborations.
DESCRIPTION: Medium to large spiders, total length of males 7–10, of females 8–13. Carapace oval, widest at rear of coxae II, abruptly narrowed at level of palpi to less than twothirds of maximum width; thoracic groove long, longitudinal, very deep; surface coated with short recumbent and fewer, longer, erect dark setae, erect setae most numerous along midline and in ocular area; eight eyes in two rows; from above, both eye rows slightly recurved; from front, anterior row recurved, posterior row slightly pro curved; anterior median eyes round, smallest; other eyes oval, subequal, with canoeshaped tapeta; anterior median eyes separated by roughly their diameter, slightly closer to anterior laterals; posterior medians separated by roughly their diameter, farther from posterior laterals; lateral eyes of each side separated by less than their diameter; median ocular quadrangle wider in back than in front, wider in back than long; clypeal height about twice diameter of anterior median eyes, corners of clypeus with incised margins that overlie cheliceral boss; chilum weakly sclerotized, divided, composed of two triangular sclerites. Chelicerae vertical, anterior surface with few, erect setae; promargin with three teeth situated at proximal end of fang furrow, most proximal tooth smallest, retromargin with three larger, more distally situated teeth; very short, narrow, Ishaped posterior sclerite present, separating chelicerae at base. Labium short, distally invaginated at middle, reflexed at almost 908 angle relative to sternum. Endites rectangular, distally slightly convergent, with anteromedian scopula and anterolateral serrula consisting of single row of teeth. Sternum rounded, without extensions to or between coxae, with erect setae; posterior margin only slightly extended between coxae IV. Figs. 13–16. Anachemmis linsdalei, new species, male. 13.Spinnerets of left side, distal view. 14.Left anterior lateral spinneret, distal view. 15.Left posterior median spinneret, distal view. 16.Left posterior lateral spinneret, distal view. Scale bar 5 43 mm (14–16), 150 mm (13). Figs. 17–20. Anachemmis linsdalei, new species, male. 17.Left palp, prolateral view, showing interlocking lobes on tegulum and subtegulum. 18.Left palp, retrolateral view. 19.Left palp, ventral view. 20.Left embolus, conductor, and median apophysis, ventral view. Scale bar 5 200 mm (20), 600 mm (17–19). Figs. 21–24. Anachemmis soberChamberlin. 21.Left male palp, ventral view. 22.Same, retrolateral view. 23.Epigynum, ventral view. 24.Same, dorsal view. Leg formula 4123. Typical leg spination pattern (only surfaces bearing spines listed): femora: I d111, p011, r121; II d111, p021, r121; III d111, p211, r121; IV d111, p211, r001; patellae III, IV p01 0, r010; tibiae: I, II p010, v442; III, IV d101, p011, v222, r011; metatarsi: I, II v222; III p112, v221r, r112; IV p1 12, v221r, r122; tarsi with three claws and claw tufts (figs. 1–4), superior claws with several weak teeth, most distal teeth largest, inferior claws unarmed; all tarsi with strong ventral scopulae, scopular hairs distinct from those of claw tufts (fig. 1); distal segments with trichobothria in two rows, bases ridged (figs. 5, 6); tarsal organ capsulate (fig. 8); all trochanters strongly notched; males without tibial crack; metatarsi without preening combs. Abdomen without anterior or dorsal scutum; anterior lateral spinnerets large, composed of two articles (figs. 9, 13), distal article with two major ampullate gland spigots and about 10 small piriform gland spigots (figs. 10, 14); posterior median spinnerets composed of one article, small, triangular (figs. 11, 15), those of female expanded posteriorly, where they bear single large and three or four smaller cylindrical gland spigots, preceded anteriorly by smaller aciniform spigots and at least two larger minor ampullate gland spigots; posterior lateral spinnerets composed of two articles, distal article about onefifth as long as proximal article, that of female apparently without cylindrical gland spigots at base, in both sexes with smaller spigots around base and on tip (figs. 12, 16); colulus represented by setae on small lobe. Figs. 25–28. Anachemmis linsdalei, new species. 25.Left male palp, ventral view. 26.Same, retrolateral view. 27.Epigynum, ventral view. 28.Same, dorsal view. Male palp with patella not widened, retrolateral tibial apophysis bipartite, ventral prong large, bladeshaped (fig. 18), dorsal prong reduced to tiny point; subtegulum and tegulum with interlocking lobes (fig. 17), median apophysis heavily sclerotized at least distally, embolus short, with shallow division between prolateral prong and copulatory prong, accompanied by hyaline conductor (figs. 19, 20). Female palp with extremely long, dentate claw. Epigynum with large median septum wider anteriorly than posteriorly; spermathecal bulbs situated posterolaterally. MISPLACED SPECIES: Anachemmis dolichopus Chamberlin (1919)is now placed in Socalchemmis Platnick and Ubick (2001). For unknown reasons, Roewer (1955)listed Chemmis unicolor Banks (1901)as a member of Anachemmis, but that transfer is incorrect; the placement of Banks’ species will be discussed in detail in a subsequent paper in this series.
KEY TO SPECIES OF ANACHEMMIS 1. Males............................. 2 — Females........................... 6 2. Tegulum much wider opposite median apophysis than elsewhere (figs. 21, 25)..... 3 — Tegulum not much wider opposite median apophysis than elsewhere (figs. 29, 33, 37) ................................ 4 3. Tegular edge opposite median apophysis base rounded (fig. 21).............. sober — Tegular edge opposite median apophysis base pointed (fig. 25)............. linsdalei 4. Ventral prong of retrolateral tibial apophysis relatively wide at base (fig. 34).. aalbui — Ventral prong of retrolateral tibial apophysis relatively narrow at base (figs. 30, 38)... ................................. 5 Figs. 29–32. Anachemmis jungi, new species. 29.Left male palp, ventral view. 30.Same, retrolateral view. 31.Epigynum, ventral view. 32.Same, dorsal view. 5. Dorsal prong of retrolateral tibial apophysis relatively large (fig. 30).......... jungi — Dorsal prong of retrolateral tibial apophysis a tiny spur (fig. 38)............. beattyi 6. Widened anterior portion of median epigynal septum relatively short (figs. 31, 35).. 7 — Widened anterior portion of median epigynal septum relatively long (figs. 23, 27, 39).. ................................. 8 7. Epigynal surface opposite sides of median septum with distinct ridge (fig. 35)................................... aalbui — Epigynal surface opposite sides of median septum without distinct ridge (fig. 31)................................. jungi 8. Widened portion of epigynal septum with paramedian invaginations (fig. 39)................................. beattyi — Widened portion of epigynal septum without paramedian invaginations (figs. 23, 27).. ................................. 9 9. Epigynal septum relatively wide both anteriorly and posteriorly (fig. 23)..... sober — Epigynal septum relatively narrow both anteriorlyand posteriorly (fig. 27)................................. linsdalei Anachemmis soberChamberlin
Figures 21–24; Map 1
Anachemmis sober Chamberlin, 1919: 13, pl. 5, fig. 5 (female holotypefrom Claremont, Los Angeles Co., California, in MCZ, examined).— Silva, 2003: 41, fig. 13d (scanning electron micrograph of male palp). Anachemmis scaber(lapsus): Roewer, 1955: 607.
DIAGNOSIS: Males closely resemble those of A. linsdaleibut differ in having a rounded tegular edge opposite the median apophysis (fig. 21); females also closely resemble those of A. linsdaleibut have a wider epigynal septum and more anteriorly restricted lateral epigynal margins (fig. 23).
MALE: Total length 8.0. Carapace light brown, with darker triangular markings radiating from thoracic groove to intercoxal areas; abdomen mottled brownish gray; femora light brown, more distal leg segments grading to dark reddish brown on metatarsi and tarsi. Leg spination: femora: II p121; IV r0 02; tibiae: I d101, r010; II d001, r01 0; tibiae: I, II p110, r110; III p122; IV v222. Tegulum widened opposite median apophysis, with rounded edge (fig. 21); dorsal prong of retrolateral tibial apophysis reduced to tiny, sharp point tip (fig. 22). Figs. 33–36. Anachemmis aalbui, new species. 33.Left male palp, ventral view. 34.Same, retrolateral view. 35.Epigynum, ventral view. 36.Same, dorsal view. FEMALE: Total length 10.3. Coloration as in male. Leg spination: femora: III r112; IV r001; tibiae I p000; III p122; IV v22 2. Epigynum with median septum widened anteriorly and posteriorly, lateral margins restricted to anterior portion of epigynum (fig. 23); spermathecal bulbs oval (fig. 24). VARIATION: One male taken at Shortcut on the Angeles Crest Highway in the San Gabriel Mountains of Los Angeles County differs from other specimens in having a longer dorsal prong of the retrolateral tibial apophysis. One female from the nearby locality of Switzers Camp differs from most others in having the lateral epigynal bulges relatively ‘‘deflated’’. The geographic congruence of these specimens suggested that a separate species might be present in the San Gabriels, but thanks to the help of Wendell Icenogle, a good series of specimens from these and nearby localities has now been available for study. No other males show the abnormally long dorsal prong of the tibial apophysis, and no consistent differences have been detected among the palpal bulbs of any of these specimens. Another female shows the relatively ‘‘deflated’’ lateral epigynal bulges, but only on one side of the epigynum. We conclude that there is evidence for the existence of only a single species at these sites. Another male from Lake Fulmor in Riverside County has a relatively truncated left embolus tip, but is presumably either broken or represents a developmental deformity, as the right embolus of that specimen appears normal.
MATERIAL EXAMINED: CALIFORNIA: Inyo Figs. 37–40. Anachemmis beattyi, new species. 37.Left male palp, ventral view. 38.Same, retrolateral view. 39.Epigynum, ventral view. 40.Same, dorsal view. Co.: Ninemile Canyon, June 11, 1993, elev. ca. 7000 ft( R. Robertson, CDU), 1/. Kern Co.: Bakersfield, June 28, 1985( S. Stockwell, S. Taber, O. Francke, AMNH), 1?, 1/. Los Angeles Co.:Angeles Crest Highway, N La Canada, first canyon in Angeles National Forestc. 0.25 mi from ranger station, June 27, 1982, running on leaf litter, shaded canyon slope ( G. Lowe, AMNH), 1?; Beverly Glen Canyon, Santa Monica Mountains, May 9, 1953(W. McDonald, AMNH), 1/; Big Pines Highwayat Grand View Canyon, Angeles National Forest, Jan. 27, 1982, under log, elev. 5900 ft( B. Hebert, CBH), 1/; Big Rock Creek, Valyermo, June 12, 1943, elev. 4200 ft( K. Cowles, AMNH), 1/; CharltonChilao P. C., Angeles National Forest, Apr. 28, 1969( M. Thompson, MET), 1/; Charmlee CountyPark, May 13, 1975( M. Thompson, MET), 1?, May 3, 1997, elev. 396 m(J. Kempf, UCR), 1?; Claremont( R. Chamberlin, MCZ), 1/ ( paratype), Apr. 1913( R. Chamberlin, MCZ), 1/ ( holotype); Eaton Canyon Park, Aug. 5, 1965( M. Thompson, MET), 1?, Aug. 16, 1965(M. Thompson, MET), 1/, Aug. 4, 1967(M. Thompson, MET), 1/, Jan. 18, 1968(M. Thompson, MET), 2?, Feb. 19, 1968(M. Thompson, MET), 1/, May 5, 1968(M. Thompson, MET), 5?, 1/, May 7, 1968(M. Thompson, MET), 1?; Glendale, Nov. 25, 1950( E. Schlinger, AMNH), 1?, May 25, 1958(E. Schlinger, AMNH), 1?, May 10, 2002, in home (K. Schwellenbach, UCR), 1?; Gold Canyon, San Gabriel Mountains, Mar. 1955, matured Apr.17 ( R. Schick, AMNH), 1?, Apr. 17, 1955( R. Schick, AMNH), 1/, May 6–8, 1955( R. Schick, AMNH), 1?, 2/; Grandview Canyon, Big Pines Highway, c. 1 miW Lake Jackson, June 27, 1982, under log, elev. 5900 ft( G. Lowe, CGL), 1/; Henninger Flats, Angeles National Forest, Nov. 11–15, 1968( D. Marqua, MET), 2/; Hollywood, Oct. 1994( R. Vetter, UCR), 1/; Lady Bug Canyon, May 29–June 8, 1969–1970 ( M. Thompson, MET), 4?; Los Angeles, May 1981( G. Bakker, CDFA), 1?; Malibu, midApril 2002 ( M. Balducci, UCR), 1?; Malibu Lagoon, near Serra Road, Dec. 11, 1982, under rock ( G. Lowe, CGL), 1?; Marker Flats, Mount Baldy Road, Apr. 15, 1987, under rock on slope, elev. 6100 ft( G. Lowe, CGL), 1/; Millard Canyon, Altadena, Dec. 4, 1983, under stone on leaf litter ( G. Lowe, CGL), 1/, July 5, 1984, on rocks and tree trunks at night (C. Lowe, CGL), 2?, 2/; Millard Canyon, at junction with Canyon Crest Road, San Gabriel Mountains, Altadena, Jan. 18, 2002(matured Feb. 28, 2002), elev. 1600 ft, under trash on oak duff near stream ( W. Icenogle, J. Bond, WRI), 1?; Old Topanga Canyon Road, c. 1 mifrom Topanga Canyon Boulevardand Route27, Nov. 13, 1982, under rock, under cardboard debris on ground in oak forest ( G. Lowe, CGL), 2/, Sept. 9, 1989, found dead in spider web (B. Hebert, CBH), 1/; 4723 Palmer Canyon Road, Claremont, July 16, 1998, in kitchen sink ( A. Moore, UCR), 1?; Pasadena, May 2, 1966( C. Goodpasture, UCR), 1?; Placerita Canyon Park, 3 miE Newhall, July 26, 1972, elev. 1550 ft( F. Hovore, WRI), 1/; May 27, 1976(P. Sullivan, MET), 1/; Puente Hills, S Roland Heights, 33.969018N, 117.926888W, Aug. 25–Sept. 4, 1998, dry pitfall, elev. 294 m( R. Fisher, AMNH), 2/, Nov. 4–8, 2002, same, 1/; San Antonio Canyon, near Claremont, July 1, 1956( W. Gertsch, V. Roth, AMNH), 6?, 6/; San Antonio Canyon, Mount Baldy Road, 1.5 mi S Lower San Antonio Forest Station, San Gabriel Mountains, May 4, 1996, dead and dried in black widow web in big pipe under road ( W. Icenogle, WRI), 1?; San Antonio Canyon, 0.5 mi E Sunset Peak, 0.25 mi W Mount Baldy Road, July 9, 2001, elev. 3800 ft, under rock and log in leaf litter beside streamin shaded canyon bottom ( W. Icenogle, WRI), 1?, 1/; San Dimas Canyon, near Claremont, Sept. 2, 1956( V. Roth, AMNH), 1/; San Dimas Experimental Forest, Big Dalton Canyon Road, 0.75 mi NE Big Dalton Reservoirat junction of Volfe Canyon, 348109560N, 1178479460W, July 9, 2001(matured May 25–31, 2002), elev. 1920 ft, oak leaf litter ( M. Oxford, UCR), 1?; SantaAnita Canyon, San Gabriel Mountains, collected as juvenile Nov. 12, 1989, matured in captivity ( B. Hebert, CBH), 1?, collected Apr. 23, 1991, matured May 24, 1991(B. Hebert, CBH), 1?; 15 miW Santa Monica, Mar. 20, 1941(W. Ivie, AMNH), 1?; Shortcut, Angeles Crest Highway, San Gabriel Mountains, Apr. 26, 1969( M. Thompson, MET), 1?; Sierra Madre, June–July 1977( G. Gorelick, LACM), 3?; Spruce Canyon(W branch, San Antonio Canyon), 0.25 mi W Mount Baldy Road, San Gabriel Mountains, July 5, 2001, elev. 3600 ft, under log beside stream below oaks and alders ( W. Icenogle, WRI), 1?, 1/; Sunland, May 19, 1951( T. Tice, AMNH), 1?; Switzers Camp, Angeles Crest Highway, Angeles National Forest, June 27, 1985, elev. 3600–4000 ft( L. Sorkin, AMNH), 3/; Switzer Picnic Area, upper end of Arroyo Seco Canyon, 0.25 mi S Highway2, 0.5 mi E junction with HighwayN3, San Gabriel Mountains, Apr. 15, 2002(matured Apr. 27–29, 2002), elev. 3300 ft, under rock in duff below oaks and alders near stream at base of steep, rocky, northfacing canyon side ( W. Icenogle, WRI), 2?, 1/; Tanbark Flats, San Gabriel Mountains, June 20, 1952(M. Cazier, W. Gertsch, R. Schrammel, AMNH, JCC), 95?, 130/, Mar. 30, 1957( AMNH), 2/; Tapia Park, Santa Monica Mountains, Mar. 20, 1955, matured in Apr.( R. Schick, AMNH), 1/; Temescal Canyon, Pacific Palisades, Santa Monica Mountains, Apr. 10, 1995( B. Olodort, CBH), 1/; Topanga, off Highway27, 34.094838N, 118.601498W, Mar. 13, 2004, elev. 470 m( R. Keith, D. Palmer, J. Starrett, CDB), 1/; Upper Big Tujunga Canyon, just E Shortcut Ranger Station, San Gabriel Mountains, Aug. 27, 2001, elev. 4200 ft, dead and dried in web of Steatoda atascadera(W. Icenogle, WRI), 6?, 1/, June 11, 2002, elev. 4200 ft, on wall of 109 diameter pipe under road above small stream, conifer/oak forest ( W. Icenogle, WRI), 2?. Orange Co.:N Aliso Wood Canyon, E Laguna Beach, 33.558458N, 117.749608W, June 10–Aug. 31, 2001–2002, dry pitfalls, elev. 71 m( R. Fisher, AMNH), 2?, 4/; N Aliso Wood Canyon, E Laguna Beach, 33.567578N, 117.748258W, June 1–Aug. 9, 2000– 2002, wet and dry pitfalls, elev. 80 m( R. Fisher, AMNH), 7?, 2/; Chino Hills, Telegraph Canyon, 33.914488N, 117.785558W, Oct. 29–Nov. 2, 2001, dry pitfall, elev. 238 m( R. Fisher, AMNH), 1/; Chino Hills, ridge S Telegraph Canyon, 33.911968N, 117.789108W, Sept. 7–17, 1999, dry pitfall, elev. 358 m( R. Fisher, AMNH), 1/; Crow Canyon, 33.602898N, 117.544278W, Nov. 28– Dec. 8, 2000, dry pitfall, elev. 338 m( R. Fisher, AMNH), 1/; Laguna Beach, June 24, 1931( W. Ivie, AMNH), 1?, July 24, 1931(W. Ivie, AMNH), 1/, Dec. 28, 1932, under rock (W. Ivie, AMNH), 1/, Aug. 17, 2000( UCR), 1/; Limestone Canyon Regional Park, 33.736068N, 117.670488W, June 18–22, 2001, dry pitfall, elev. 341 m( R. Fisher, AMNH), 1?; Limestone Canyon Regional Park, E The Sinks, 33.716218N, 117.658928W, July 20–30, 1999, dry pitfall, elev. 453 m( R. Fisher, AMNH), 1?; Limestone Canyon Regional Park, ENE The Sinks, 33.722008N, 117.665798W, June 1–Sept. 3, 1998–2002, pitfall, dry pitfalls, elev. 378 m( R. Fisher, AMNH), 2?, 2/; N Niguel Hill, NW Laguna Niguel, 33.536208N, 117.730158W, Aug. 27–31, 2001, Jan. 28–Feb. 1, 2002, dry pitfalls, elev. 195 m( R. Fisher, AMNH), 2/; N Niguel Hill, NW Laguna Niguel, 33.538828N, 117.725448W, July 10–Aug. 31, 2000–2002, dry pitfalls, elev. 132 m( R. Fish er, AMNH), 3?, 1/; San Joaquin Hills, E branch Laguna Canyon, 33.594688N, 117.775298W, Sept. 9–13, 2002, dry pitfalls, elev. 175 m( R. Fisher, AMNH), 2/; Highway74, 2.5 mi E San Juan Fire Station, Hot Spring Canyonturnoff, Santa Ana Mountains, May 11–22, 2002, elev. 1200 ft, under trash on oak duff beside highway, 60 mfrom stream ( W. Icenogle, WRI), 5?, 5/; Silverado Canyon, Santa Ana Mountains, Nov. 20, 1955( R., A. Schick, AMNH), 1?; Starr Ranch, 33.630258N, 117.553088W, Sept. 5–Oct. 14, 1998–2000, elev. 297 m, dry pitfalls( R. Fisher, AMNH), 4/; N Temple Hill, W Aliso Wood Canyon, 33.569908N, 117.756338W, July 10–20, 2000, dry pitfall, elev. 215 m( R. Fisher, AMNH), 1?; N Temple Hill, W Aliso Wood Canyon, 33.583848N, 117.750308W, May 13–17, 2002, dry pitfall, elev. 206 m( R. Fisher, AMNH), 1/; Weir Canyon, 33.809948N, 117.755648W, June 15–25, 1999, dry pitfall, elev. 236 m( R. Fisher, AMNH), 1?; Weir Canyon, 33.815558N, 117.747138W, Sept. 19–29, 2000, dry pitfall. elev. 232 m( R. Fihser, AMNH), 1/. Riverside Co.: Bay Tree Springs, San Jacinto Mountains, Nov. 13, 1976( M. Long, MET), 1/; Chino Canyon, Apr. 20, 1960( J. Powell, UCB), 2?; 0.25 mi S Dark Canyon Campground, 2 road mi N junction Hwy243, San Jacinto Mountains, June 4, 2002, under wood pile on ground near stream, conifer/oak forest ( W. Icenogle, WRI), 1/, June 5, 2002, on side of boulder near stream, conifer/oak forest (W. Icenogle, WRI), 1/, June 4, 2003, elev. 5765 ft, under wood piece beside recently molted skin, on ground near stream, conifer/oak forest (W. Icenogle, WRI), 1/; De Luz Road, 3 miS Murrieta, May 10, 1977, under rock in ravine below live oaks, elev. ca. 2000 ft( W. Icenogle, WRI), 1/; Idyllwild, San Jacinto Mountains, June 17, 1952(W. Gertsch, AMNH), 6?, 5/, July 7, 1953( W. Gertsch, V. Roth, AMNH), 4?, 4/, 1996, elev. prob. 6000 ft( UCR), 1/; James Reserve, San Jacinto Mountains, 9 mi N Idyllwildon Highway243, 33848.49N, 116846.49W, June 29, 2002, elev. 550 ft, coniferous forest, under granite exfoliations ( D., S. Ubick, M. Harvey, CDU), 3/; Lake Fulmor, James Reserve, San Jacinto Mountains, off Highway243, elev. 5380 ft( J. Kempf, UCR), 1/, Aug. 1968( D. Bixler, CDB), 2?, Sept. 1968(D. Bixler, CDB), 1/; Santa Margarita Ecological Reserve, W Temecula, 33.445238N, 117.164458W, Sept. 7–17, 1999, dry pitfall, elev. 393 m( R. Fisher, AMNH), 1/; Santa Margarita Ecological Reserve, W Temecula, 33.455048N, 117.171238W, May 11–21, 1998, dry pitfall, elev. 786 m( R. Fisher, AMNH), 1/; Santa Margarita Ecological Reserve, W Temecula, 33.456898N, 117.168458W, Jan. 28–Feb. 7, 2000, dry pitfall, elev. 788 m( R. Fisher, AMNH), 1/; Strawberry Creek, 0.25 mi N Highway74, San Jacinto Mountains, Aug. 12, 2001, elev. 3300 ft, under moist, rotting log beside stream ( W. Icenogle, WRI), 1/. San Bernardino Co.: Crestline, San Bernardino Mountains, matured Feb. 1997, elev. 5000 ft( J. Klotz, UCR), 1?; Crestline area, upper end of Waterman Canyon, Highway18, 0.5 mi NW junction with Highway138, San Bernardino Mountains, Aug. 30, 2001, elev. 4200 ft, on wall, or dead and dried in spider webs on wall, of long pipe under highway ( W. Icenogle, WRI), 2?, 2/; East Fork Mountain Home Creek, near Highway38, San Bernardino Mountains, July 6, 1999, under large rock in conifer forest near stream, elev. 5000 ft( W. Icenogle, WRI), 1/; Fontana, July 16, 1942( E. Schlinger, AMNH), 1?, 1/; Forest Falls, Aug. 10, 1985, elev. 7000 ft( D. Bixler, CDB), 2/; Forest Home, Hill Creek, June 11, 1971, in cabin, elev. 6000 ft( D. Lowrie, MCZ), 1?; Glass Road, San Bernardino Mountains, 1 miN Highway 38, 348109290N, 1168549000W, May 6, 2001(matured Apr. 1, 2003), elev. 1820 m, in deciduous duff from Quercus kelloggi(R. Vetter, UCR), 1/; off Highway38, East Fork Mountain, Eside Home Creek, San Bernardino Mountains, 348079300N, 1168589540W, Feb. 3, 2002(matured Oct. 26–Nov. 2, 2002), elev. 5120 ft, from dead fern leaves, ferns growing on westfacing rock outcropping on steep canyon wall ( T. Prentice, UCR), 1?; Highway173 near Lake Arrowhead Village, 348159020N, 1178119590W, July 10, 1999, elev. 5200 ft, along roadside and on hills at night ( B. Hebert, CBH), 3?, 2/; Lake Arrowhead, May 6, 1936( S. Bishop, AMNH), 1/; Lytle CreekCanyon, Aug. 20, 1970( D. Bixler, CDB), 1/, May 13, 1972, elev. 3000 ft( D. Bixler, CDB), 1?, May 16, 1972, in house ( D. Bixler, CDB), 1?, June 1, 1973( D. Bixler, CDB), 1?; Manker Flats Campground, San Gabriel Mountains, May 14, 1992, elev. 6200 ft( B. Hebert, CBH), 1?, 1/; Mill CreekCanyon, Falls Picnic Area, San Bernardino Mountains, May 15, 1969, molted May 28, 1969, under rock in conifer forest, elev. 6000 ft( W. Icenogle, WRI), 1?, June 18, 1969, elev. 6000 ft, under wood pieces and rocks (W. Icenogle, WRI), 1 penultimate /; Mountain Home Creek, Aug. 13, 1959( W. Gertsch, V. Roth, AMNH), 9/; 209 Pine Oak, Fawnskin, July 21, 1995, in lower level of house in tire chains ( V. Summer, UCR), 1?; Silverwood Lake State Recreation Area, SEend Silverwood Lake, 34.277768N, 117.316948W, Aug. 12–16, 2002, dry pitfall, elev. 1035 m( R. Fisher, AMNH), 1/; Skinner Creek, just E Highway38, 1.5 road mi NW junction with road to Forest Falls, San Bernardino Mountains, June 10, 2001, elev. 4200 ft, under rocks and logs beside and above stream in bottom of heavily wooded canyon ( W. Icenogle, WRI), 2?, 1/; Skinner Ridge, between Skinner Creek and Mountain Home Creek, just SW Highway 38, 348069480N, 1168589530W, Dec. 28, 1983– Jan. 26, 1984, elev. 4900 ft, pitfall traps, canyon live oak forest, northern exposure ( M. Narog, UCR), 5/; Smiley Park, Running Springs, July 12, 1995, in shower ( UCR), 1?; Smiley Park, 2 miS Running Springs on Highway330 off Fredalba Road, July 10, 1995( J. Wambaugh, UCR), 1/; Twin Peaks, San Bernardino Mountains, ( K. Wormald, UCR), 1?, 1/, late May– early June 1996, in house ( K. Wormald, UCR), 1?, 1/, midJuly 1996, male found in house, female outside (K. Wormald, UCR), 1?, 1/, Aug.– Sept. 1996, elev. 5500 ft(K. Wormald, UCR), 2/, mid May–mid June 1997( R. Vetter, UCR), 11?, June 11, 1997( K. Wormald, UCR), 1/. San Diego Co.: De Luz Road, 1 miN Fallbrook, May 19, 1984, in drain under road ( G. Lowe, CGL), 1?, Aug. 25, 1984(G. Lowe, CGL), 1/; Fry Creek, Mount Palomar, Aug.4, 1984 ( D. Boe, UCR), 1/; near Henshaw, Cleveland National Forest, July 30, 1956( W. Gertsch, V. Roth, AMNH), 1?, 2/; W side Lake Henshaw, just E route S6, 33.244388N, 116.764118W, May 16, 2004, elev. 308 m(D. Palmer, P. Paquin, T. Prentice, C. Vink, AMNH), 1?; SW end Mendenhall Valley, Palomar Mountain, June 3–July 2, 1997, pitfall, forest ( J. George, UCR), 3?, July 2–Aug. 7, 1997, pitfall, elev. 6000 ft(J. George, UCR), 2?; Mount Palomar, July 26, 1931( R. Chamberlin, AMNH), 1/, June 30, 1956, elev. 3000–5000 ft( W. Gertsch, V. Roth, AMNH), 4?, 7/; Palomar Mountain, July 4, 1967( B. Kaston, WRIK), 1?; Palomar Mountain, N Fry Creek, Wside route S6, 33.34588N, 116.87948W, May 16, 2004, elev. 1500 m(D. Palmer, P. Paquin, T. Prentice, C. Vink, AMNH), 3?, 1/; Palomar Mountain State Park, July 13, 1953( W., J. Gertsch, AMNH), 1?, 3/; Palomar Mountain State Park, Doone Trail, 338209290N, 116854950W, Feb. 20, 2003, in duff on creek bed by road, elev. 4639 ft( R. Vetter, UCR), 1? (matured midMarch 2003). Santa Barbara Co.: Cold Spring Trail, Santa Barbara, Aug. 14, 1959( W. Gertsch, V. Roth, AMNH), 31/; 5 miE Gaviota, July 17, 1953( W., J. Gertsch, AMNH), 13?, 17/ (one with apparently teratological epigynum); Refugio Rd., 23 miW Santa Barbara, Apr.6– May, 1978–1979 ( D. Boe, UCR), 4/; Santa Barbara, Nov.–Dec. 1950( H. Shantz, AMNH), 2?, July 17, 1953(H. Shantz, AMNH), 1?, July 27, 1970(M. Cravens, CDFA), 1?; 10 miW Santa Barbara, July 12, 1934(W. Ivie, AMNH), 1?, 2/; Sedgwick Ranch Reserve, 348439N, 1208019W, May–June 11, 1997, malaise traps, elev. 1000–1800 ft( E., M. Schlinger, CAS), 1?, 1/, June 24–July 21, 1997, malaise traps, elev. 800–1500 ft(E., M. Schlinger, CAS), 3?, Aug.27– Nov.1997, same, 3/, May 28, 1997, malaise trap, elev. 1500–2500 ft( E. M. Schlinger, CAS), 1?; Upper Oso Campground, near Santa Ynez River, Apr. 26–27, 1975, molted before May 18, 1975, under rock ( F. Hovore, WRI), 1?. Tulare Co.: Ash Mountain, Kaweah Power Station# 3, 40 miNE Visalia, July 3, 1982( D. Burdick, CAS), 1?, Mar. 12, 1983(D. Burdick, CAS), 1/, June 19–21, 1983(D. Haines, CAS), 2?, July 6–10, 1983, in flume (D. Burdick, CDU), 7?, 3/, July 17, 1983(D. Burdick, CAS), 2/, July 31, 1983( R. Haines, CDU), 1?, 3/, June 8, 1985( R. Haines, CDFA), 2?, 1/, June 7, 1986( R. Haines, CDFA), 1?, June 9, 1986( R. Haines, CDFA), 2/, June 28, 1986( R. Haines, CDFA), 1?, July 12, 1986( R. Haines, CDFA), 1/, Aug. 5–Sept. 3, 1995( D. Burdick, CAS), 3/, July 24, 1996(D. Burdick, CAS), 1?; near Ash Mountainentrance, Sequoia National Park, July 5, 1956, elev. 3700 ft( W. Gertsch, V. Roth, AMNH), 2?, 1/, July 9, 1958( W. Gertsch, V. Roth, AMNH), 9?, 8/; NE Case MountainPeak, 36.417908N, 118.793298W, July 14–18, 2003, dry pitfall, elev. 1691 m( R. Fisher, AMNH), 1?; NE Case Mountain Peak, 36.421838N, 118.793708W, July 14–18, 2003, dry pitfall, elev. 1646 m( R. Fisher, AMNH), 1?; NE Case Mountain Peak, N Case Mountain Grove, 36.413178N, 118.791908W, July 14–22, 2002–2003, dry pitfall, elev. 1810 m( R. Fisher, AMNH), 3?; NNE Case Mountain Peak, Eof E end of Salt Creek, 36.420358N, 118.793918W, July 19–22, 2002, dry pitfall, elev. 1644 m( R. Fisher, AMNH), 1?, 36.421088N, 118.799768W, July 19–22, 2002, dry pitfall, elev. 1819 m( R. Fisher, AMNH), 2?; NW Case Mountain Peak, S Salt Creek, 36.414428N, 118.813818W, July 14–22, 2002–2003, dry pitfalls, elev. 1855 m( R. Fisher, AMNH), 3?; NW Case Mountain Peak, S Salt Creek, 36.417818N, 118.819308W, July 14–18, 2003, dry pitfall, elev. 1709 m( R. Fisher, AMNH), 3?; SE Case MountainPeak, Case Mountain Grove, 36.403728N, 118.781908W, July 14–18, 2003, dry pitfall, elev. 1678 m( R. Fisher, AMNH), 1?; SE Case MountainPeak, Case Mountain Grove, 36.405118N, 118.189828W, July 19–22, 2002, dry pitfall, elev. 1805 m( R. Fisher, AMNH), 1/; SE Case Mountain Peak, Salt CreekRidge, 36.397408N, 118.775978W, July 19–22, 2002, dry pitfall, elev. 1801 m( R. Fisher, AMNH), 1?, 36.398878N, 118.787788W, July 19–22, 2002, dry pitfall, elev. 1848 m( R. Fisher, AMNH), 1?; Kaweah River, 5 miE Three Rivers, July 17, 1952, elev. 1258 ft( W. Gertsch, AMNH), 1/; 5–10 mion Mineral King Roadfrom Three Rivers, July 4, 1956, elev. 2000–4000 ft( W. Gertsch, V. Roth, AMNH), 5?, 5/; Moorehouse Creek, near Camp Nelson, July 11, 1958( W. Gertsch, V. Roth, AMNH), 1?, 5/; 1 miW Camp Nelson, July 11, 1958( W. Gertsch, V. Roth, AMNH), 2?; Roads End, Kern River, July 3, 1956( W. Gertsch, V. Roth, AMNH), 3?, 2/; Sequoia National Park, Kaweah Cave, 36829926.10N, 118849917.40W, Apr. 28, 2004, in cave entrance, elev. 490 m( D. Ubick, CDU), 1/ (matured May 15, 2004); Soda Creek, W Camp Nelson, July 11, 1958( W. Gertsch, V. Roth, AMNH), 3/, Sept. 14, 1959( W. Gertsch, V. Roth, AMNH), 1/. Ventura Co.: Ojai, Mar. 1978( D. Boe, UCR), 2/; Reyes Creek, 25 miSW Lakeof the Woods, May 6, 1980, elev. 4000 ft( C. Griswold, UCB), 1/; Stecker Park, Highway150 N Santa Paula, 348259N, 119859W, July 4, 1986, elev. 650 ft, from black widow web in restroom ( B. Hebert, CBH), 1?, 1/; Wheeler Springs, July 2, 1958( W. Gertsch, V. Roth, AMNH, CAS), 17?, 13/; Yerba Buena Road, 2–3 miN Route1, June 26, 1983, under rock by stream in canyon ( G. Lowe, CGL), 1?.
DISTRIBUTION: Southern California(map 1). NATURAL HISTORY: Collection records indicate that Anachemmis soberoccurs in forests, primarily oak but also coniferous, at elevations from 70 to 2000 m. Specimens have Map 1. Western North America, showing records of Anachemmis sober(open circles), A. linsdalei(filled circles), A. jungi(filled squares), A. aalbui(open square), and A. beattyi(triangles). been found beneath rocks, logs, and other ground litter, in and on leaf litter, on tree trunks, in a cave entrance, and in human structures, such as houses, culverts, and a flume. Several collections are from pitfall and malaise traps and of dead individuals in spider webs, including those of Latrodectusand Steatoda. The 700 specimensexamined indicate that both sexes occur year round, with females being more commonly collect ed, and a peak abundance from May to August (chart 1). That this species may be very common is documented by a single June day’s collection, at Tanbark Flats, Los Angeles County, of 95 malesand 130 females. Anachemmis linsdalei, new species Figures 1–20, 25–28; Map 1 TYPES: Male holotypeand female allotypetaken in house at Hastings Natural History State Reserve, Monterey Co., California( Jan. 24–25, 1946; J. Linsdale), deposited in AMNH.
ETYMOLOGY: The specific name is a patronym in honor of the collector of the typesand many other specimens.
DIAGNOSIS: Males closely resemble those of A. soberbut differ in having a sharply pointed tegular edge opposite the median apophysis (fig. 25); females also closely resemble those of A. soberbut have a narrower epigynal septum and less anteriorly restricted lateral epigynal margins (fig. 27).
MALE: Total length 7.4. Coloration as in A. sober. Leg spination: femora: II p121; IV r002; tibiae: I d001, r010; II d101, r010; III, IV d111; metatarsi: I, II p11 0, r110; III p122, r122; IV p122, r1 12. Tegulum widened opposite median apophysis, with pointed edge (fig. 25); dorsal prong of retrolateral tibial apophysis reduced to tiny, sharp point tip (fig. 26). FEMALE: Total length 9.7. Coloration as in A. sober. Leg spination: femora I r111; tibiae: II r0–10; III, IV d111; metatarsi III, IV p122. Epigynum with median septum widened at about half its length, lateral margins longer than widened portion of septum (fig. 27); spermathecal bulbs rounded (fig. 28). OTHER MATERIAL EXAMINED: CALIFORNIA: Alameda Co.:Berkeley, (AMNH), 4?, 2/. Humboldt Co.: Humboldt RedwoodsState Park, Albee Creek Campground, Sept. 10, 1992, elev. 360 ft(D. Ubick, J. Boutin, CDU), 1/. Monterey Co.: Arroyo Seco, Indians Road, Los Padres National Forest, 368149N, 121829.59W, May 7, 1995, dead yucca, elev. 1100 ft(D. Ubick, W. Savary, CDU), 1/, July 8, 1995, at night (D. Ubick, S. Fend, W. Savary, CDU), 1?, 1/, Aug. 24, 1996, oakchaparral, at night along road (D. Ubick, CDU), 4/, July 3, 1999, on road at night (S. Fend, J. Ledford, D. Ubick, CDU), 2?, Apr. 15, 2000, oak woodland, under rocks, in litter (D., S. Ubick, CDU), 2?, 2/, Apr. 28, 2001, oak woodland, under rocks and in litter (D., S. Ubick, CDU), 2/, Apr. 27–Aug. 19, 2001, pitfall traps, elev. 1100 ft(D., S. Ubick, CDU), 94?, 12/, Aug. 19–Nov. 11, 2001, same, 2/, Feb. 22–May 19, 2002, same, 7?, 2?; May 19–Oct. 13, 2002, same, 100?, 24/, Oct. 13, 2002– Jan. 25, 2003, same, 1/, Jan. 25–June 11, 2003, same, 8?, 3/, June 11–Nov. 28, 2003, same, 63?, 18/, Nov. 28, 2003– June 18, 2004, same, 44?, 4/, June 18–Oct. 24, 2004, same, 24?, 14/; Arroyo Seco Campground, Los Padres National Forest, Apr. 26, 1980, under rock (D. Ubick, CDU), 1/, May 13, 1994, at outhouse, elev. 950 ft(D. Ubick, J. Boutin, CDU), 1?, May 13–15, 1994, oak forest, ground litter, elev. 900 ft(J. Boutin, CDU), 3?, 3/, Apr. 24, 1999, oak riparian (D. Ubick, W. Savary, CDU), 1/; Arroyo Seco Campground, S The Lakes, Los Padres National Forest, 368149N, 1218299W, May 13–15, 1994, under granite, leaf litter, elev. 900 ft(D. Ubick, J. Boutin, CDU), 3?, 6/, May 6, 1995, under granite (D. Ubick, W. Savary, CDU), 1/, Apr. 27–May 18, 2001–2002, at granite talus, under rocks (D., S. Ubick, DUC), 3?, Apr. 27–Aug. 19, 2001, same, pitfall traps, 3?, 1?, Aug. 19, 2001– May 18, 2002, same, pitfall traps, 3/, June 11–Nov. 28, 2003, same, 6?, 5/, Nov. 28, 2003– June 18, 2004, same, 9?, June 18–Oct. 24, 2004. Same, 2?; Big Sur, July 1937(W. Ivie, AMNH), 1/, June 25, 1952( M. Cazier, W. Gertsch, R. Schrammel, AMNH), 1?, 4/, July 18, 1953(W., J. Gertsch, AMNH), 1?, 6/, Apr. 2, 1960( W. Gertsch, W. Ivie, R. Schrammel, AMNH), 4?, 6/; 14 miS Big Sur, Dec. 22, 1953( V. Roth, AMNH), 1?; Carmel, Nov. 12, 1953( AMNH), 1/, Apr. 5–May 23, 1954( AMNH), 4?, 5/; 1 miN Carmel, Dec. 21, 1953( V. Roth, AMNH), 1/; Carmel Valley, Apr. 9, 1979(D. Boe, UCR), 1?, 1/, May 1996(P. Stadille, CDU), 2?; Coast Ridge Trail, 0.8 mi SE Nacimiento Road, Jun. 1, 1991, oakpine forest, under bark litter, elev. 3000 ft(D. Ubick, CDU), 2?, 2/; Hastings Natural History State Reserve, 36822.89N, 121833.29W, May 19–29, 1939(J. Linsdale, CAS), 2?, May 30, 1940(J. Linsdale, CAS), 1/, Feb. 22, 1941(J. Linsdale, CAS), 1/, May 27, 1942(J. Linsdale, CAS), 1/, July 18, 1944(J. Linsdale, CAS), 1?, Jan. 24–25, 1946, house (J. Linsdale, AMNH), 2/, Apr. 10, 1946, among sticks (J. Linsdale, CAS), 1/, May 17, 1946, soil under house, passage in soil (J. Linsdale, AMNH), 1?, 1/, Nov. 18, 1946, Neotomanest (J. Linsdale, AMNH), 2?, Dec. 8, 1946, house wall (J. Linsdale, AMNH), 2/, Dec. 9, Chart 1. Seasonal abundance of Anachemmis soberbased upon the total specimens examined (728) from all localities. Yaxis 5 number of specimens; Xaxis 5 time in months; solid lines 5 females; dashed lines 5 males. 1946, Neotomanest (J. Linsdale, CAS), 1/, June 1947(J. Linsdale, AMNH), 1?, May 24, 1948, museum (J. Linsdale, AMNH), 1?, May 17, 1950, under log (J. Linsdale, CAS), 1/, June 24, 1950, headquarters, at night ( J. Linsdale, AMNH, 1/, July 23, 1999, on trail at night ( S. Fend, J. Ledford, D. Ubick, CDU), 9?, 5/, Aug. 2–4, 1999, in riparian woodland along creek ( D. Ubick, S. Fend, J. Ledford, CAS), 1/; Hastings Natural History State Reserve, Big Creek, 36.388998N, 121.548618W, Jan. 9, 2004, elev. 578 m(D. Palm er, N. Stewart, AMNH), 1/; Highway G16 at G17, Apr. 10, 1979(D. Boe, UCR), 1/; Landels Hill Big Creek Reservoir, 5 miN Lucia, June 4– 6, 1982(C. Griswold, UCB), 1/; Monterey, Sept.6, 2002, in house (C. Edwards, UCR), 1/; Nacimiento Roadat Kirk Creek, May 31, 1991, redwood forest, in duff, under rocks, elev. 1000 ft(D. Ubick, CDU), 1?, 1/; Nacimiento Roadat N Fork Mill Creek, June 1, 1991, redwood forest, under bark and mapletan oak litter, elev. 1500– 1600 ft(D. Ubick, CDU), 1?; Pacific Grove, Aug. 15, 1931(W. Ivie, AMNH), 1/; Pfeifer Big Sur State Park, 27 miS Carmel, May 3, 1964, elev. ca. 300 ft(P. Craig, CAS), 2/, Aug. 19, 1987, riparian (D. Gross, CAS), 1/; Redwood Canyon, 4 miS Gorda, Apr. 2, 1960(W. Gertsch, W. Ivie, R. Schrammel, AMNH), 2/; 4 miW Salinas, Sept. 25, 1961( W. Gertsch, W. Ivie, AMNH), 1/; Tassajara Spring Buddhistretreat, Sept. 13, 1978( R. Doty, AMNH), 1/; UCNRS Big Creek Reserve, nr. Whale Point Cabin, 3684.349N, 121835.819W, elev. ca 200 m, May 24– 27, 2002(S. Lew, UCB), 2/, Oct. 24, 2002(S. Lew, UCB), 1? (molted in captivity), May 12, 2003(S. Lew, UCB), 1/. San Benito Co.: Pinnacles National Monument, July 3, 1958( W. Gertsch, V. Roth, AMNH), 1?, 2/; Pinnacles National Monument, Balconies Caves area, June8, Chart 2. Seasonal abundance of Anachemmis linsdaleibased upon the total of handcollected specimens examined (182) from all localities. Yaxis 5 number of specimens; Xaxis 5 time in months; solid lines 5 females; dashed lines 5 males. 1991, oakriparian, under granite (D. Ubick, CDU), 1/; Pinnacles National Monument, Bear Gulch Trail at Chalone Creek, May 8, 1992, oakbuckeye grove, under rocks in talus slope (T. Briggs, D. Ubick, W. Rauscher, CDU), 1/. San Luis Obispo Co.:Cambria, Nov. 16, 1937(O. Bryant, AMNH), 1?, 3/, June 23, 1952(M. Cazier, W. Gertsch, R. Schrammel, AMNH), 13?, 17/, Feb. 7, 1967, in pine duff (V. Roth, AMNH), 1?; Old Morro Road, 1 miE Hwy 41, Apr. 3, 1985, oakgrassland, under serpentine (W. Rauscher, CDU), 1/, Apr. 3, 1985, matured in May (D. Ubick, CDU), 1?, 1/; 6 miN Paso Robles, Sept. 26, 1961(W. Gertsch, V. Roth, AMNH), 1/; 26 miW Paso Robles, May 12, 1961(R. Schuster, F. Raney, AMNH), 1/; Reservoir Canyon, San Luis Obispo, Aug. 15, 1959(W. Gertsch, V. Roth, AMNH), 2/; San Luis Obispo, Aug. 8, 1931(R. Chamberlin, AMNH), 1/. Santa Cruz Co.:Santa Cruz, Apr.7, 1979 (D. Boe, UCR), 2?, 1/.
DISTRIBUTION: Northern and central California(map 1). NATURAL HISTORY: Collection records of Anachemmis linsdaleiare from elevations of 100 to 500 mand predominantly from various typesof oak woodlands, but also from pine and redwood forests. This species occurs on ground habitats and has been collected under rocks, logs, in and on litter, in Neotomanests, in houses, and on roads at night. Hand collected specimens (about 200) show a similar abundance to A. sober, being collected year round with females being slightly more common and a peak of activity from April to June (chart 2). The 450 specimensobtained from ethylene glycol pitfall traps (10 traps) at Arroyo Seco, Monterey County, indicate a much higher proportion of males (chart 3). During periods of peak activity, this species was, by far, the most abundant spider in the pitfall traps.
Anachemmis jungi, new species Figures 29–32; Map 1 TYPES: Male holotypeand female allotypefrom Titus Canyon Cave, Death ValleyNational Monument, Inyo Co., California( Jan. 22, 1972; A. Jung), deposited in AMNH.
ETYMOLOGY: The specific name is a patronym in honor of the collector of the types.
DIAGNOSIS: Males of this longlegged, cavernicole species resemble those of A. aalbuiin having a narrow, erect embolus (fig. 29), but have a much narrower ventral prong of the retrolateral tibial apophysis (fig. 30); females also resemble those of A. aalbuiin having very short anterolateral epigynal margins, but have the epigynal septum widened at the level of the anterolateral epigynal margins (fig. 31).
MALE: Total length 8.9. Color paler than in A. sober, with carapace brownish orange, legs with femora light yellow, grading to brownish orange on more distal segments, abdomen pale white. Leg spination: femora: I p021; II p221; IV r012; tibiae: I, II d1 01, v642, r100; IV r101; metatarsi: I, II p110, r110; III p122; IV v222. Tegulum not widened opposite median apophysis, embolus relatively narrow, erect (fig. 29); dorsal prong of retrolateral tibial apophysis relatively large for genus, ventral prong relatively narrow (fig. 30). FEMALE: Total length 13.6. Coloration as in male. Leg spination: femora: I p031; II p221; III r211; IV p212, r012; tibiae: I, II v462, r010; metatarsi: III p122; IV r112. Anterolateral epigynal margins and atrium relatively short, septum widened at that level (fig. 31), median epigynal ducts massive (fig. 32). terolateral epigynal margins, but have the epigynal septum widest anteriorly, rather than at the level of the anterolateral epigynal margins (fig. 35). MALE: Total length 9.2. Coloration as in A. jungi. Leg spination (both legs I missing): femora: II p221; IV r002; tibiae II d10 1, v462, r010; metatarsi: II p200, r10 0; III p122, r122; IV r112. Tegulum not widened opposite median apophysis, embolus relatively narrow, erect (fig. 33); dorsal prong of retrolateral tibial apophysis relatively large for genus, ventral prong relatively wide, especially at base (fig. 34). FEMALE: Total length 12.3. Coloration as in A. jungi. Leg spination: femora: I p031; II p221; III r211; IV r002; tibiae I, II v462, r010; metatarsi: I, II p200, r20 0; III p122; IV p122, r112. Anterolateral epigynal margins and atrium relatively short, septum widened anteriorly (fig. 35), median epigynal ducts massive (fig. 36). OTHER MATERIAL EXAMINED: CALIFORNIA: San Bernardino Co.: Medicine Cave, Providence OTHER MATERIAL EXAMINED: CALIFORNIA: Inyo Co.: Defense Cave, Stone Canyon, Argus Range, Sept. 7, 1988, elev. 4000 ft( R. Aalbu, CAS), 1/; Titus Canyon Cave, Death ValleyNational Monument, Jan. 22, 1972( A. Jung, AMNH), 1?.
DISTRIBUTION: Known only from caves in Inyo Co., California(map 1). Anachemmis aalbui, new species Figures 33–36; Map 1
TYPE: Male holotypetaken in pitfalltrap at an elevation of 4300 ftat the Park Headquarters, Mitchell Caverns State Park, San Bernardino Co., California( Mar. 17–June 16, 1979; R. Aalbu), deposited in CAS.
ETYMOLOGY: The specific name is a patronym in honor of the collector of the types.
DIAGNOSIS: Males of this longlegged species resemble those of A. jungiin having a narrow, erect embolus (fig. 33), but have a wider ventral prong of the retrolateral tibial apophysis (fig. 34); females also resemble those of A. jungiin having very short an Chart 3. Seasonal abundance of Anachemmis linsdaleibased upon the specimens collected (450) from 10 pitfall traps maintained at Arroyo Seco, Monterey County, California. Yaxis 5 number of specimens; Xaxis 5 time; solid lines 5 females; dashed lines 5 males. Mountains State Recreation Area, June 20–Sept. 27, 1988, pitfall( R. Aalbu, CAS), 6/.
DISTRIBUTION: Known only from San Bernardino Co., California(map 1). Anachemmis beattyi, new species Figures 37–40; Map 1
TYPE: Male holotypefrom Peppersauce CaveCanyon(north side of Santa Catalina Mountains), Pinal Co., Arizona( Apr. 21, 1961; J. Beatty), deposited in AMNH.
ETYMOLOGY: The specific name is a patronym in honor of the collector of the types.
DIAGNOSIS: Males of this distinctive species can be recognized by the distally long and narrowed embolus (fig. 37), females by the anteriorly very wide epigynal septum bearing deep, paramedian invaginations (fig. 39).
MALE: Total length 8.4. Coloration as in A. jungi. Leg spination: femora: I p031, r2 12; II p211; IV r012; tibiae: I d101, v4 62, r110; II d101, v462, r100; metatarsi: I r100; II p010, r110. Tegulum not widened opposite median apophysis, embolus narrowed distally (fig. 37); dorsal prong of retrolateral tibial apophysis reduced to tiny point, ventral prong relatively narrow (fig. 38). FEMALE: Total length 8.8. Coloration as in A. jungi. Leg spination: femora: I p021, r1 11; IV p201, r002; patellae: III p000, r000; IV p000; tibiae I, II v462, r010; metatarsi III p122. Anterolateral epigynal margins and atrium relatively short, septum much widened at that level, bearing deep paramedian invaginations (fig. 39), median epigynal ducts massive (fig. 40). OTHER MATERIAL EXAMINED: ARIZONA: Coconino Co.: Manzanita Camp, Oak CreekCanyon, July 25, 1952( M. Cazier, AMNH), 2/. Graham Co.: Lower Canyon, Mount Graham, July 14, 1956( W. Gertsch, V. Roth, AMNH), 1/. MEXICO: SONORA: Cajon Bonito, 318189N, 1088559W, Sept. 11, 1973( V. Roth, CAS), 3/.
DISTRIBUTION: Known only from Arizona and Sonora(map 1).
1985-06-28
1993-06-11
1985-06-28
R, CDU, AMNH
Robertson & S. Stockwell & S. Taber & O. Francke
Kern Co.
2134
Ninemile Canyon
Bakersfield
11
12
1
1982-06-27
AMNH
N La & G. Lowe
Canada
Los Angeles Co.
Highway
Angeles National Forest
11
12
1
1953-05-09
AMNH
Heard Island and McDonald Islands
Beverly Glen Canyon
Santa Monica Mountains
11
12
1
1982-01-27
CBH
B. Hebert
Heard Island and McDonald Islands
Angeles National Forest
1798
Highway
Grand View Canyon
11
12
1
1943-06-12
AMNH
K. Cowles
Heard Island and McDonald Islands
1280
Big Rock Creek
Valyermo
11
12
1
1969-04-28
MET
Charlton-Chilao P. C. & M. Thompson
Heard Island and McDonald Islands
Angeles National Forest
11
12
1
1975-05-13
MET
M. Thompson
Heard Island and McDonald Islands
Charmlee County
Charmlee County
11
12
1
1997-05-03
UCR
Kempf
Heard Island and McDonald Islands
396
11
12
1
1913-04
1965-08-05
1913-04
R, MCZ, MET
Claremont & Chamberlin & M. Thompson
Heard Island and McDonald Islands
Chamberlin
Eaton Canyon Park
11
12
2
paratype
1965-08-16
MET
Thompson
Heard Island and McDonald Islands
11
12
1
paratype
1967-08-04
MET
Thompson
Heard Island and McDonald Islands
11
12
1
paratype
[462,948,1107,1741]
1968-01-18
MET
Thompson
Heard Island and McDonald Islands
11
12
1
paratype
1968-02-19
MET
Thompson
Heard Island and McDonald Islands
11
12
1
paratype
1968-05-05
MET
Thompson
Heard Island and McDonald Islands
11
12
1
paratype
1968-05-07
MET
Thompson
Heard Island and McDonald Islands
11
12
1
paratype
1950-11-25
AMNH
E. Schlinger
Heard Island and McDonald Islands
Glendale
11
12
1
paratype
1958-05-25
AMNH
Schlinger
Heard Island and McDonald Islands
11
12
1
paratype
2002-05-10
UCR
Schwellenbach
Heard Island and McDonald Islands
11
12
1
paratype
1955-03
1982-06-27
1955-03
R, AMNH, CGL
G. Lowe
Heard Island and McDonald Islands
Highway
1798
Gold Canyon
Big Pines Highway
11
12
1
paratype
1968-11-11
1968-11-15
1968-11-11
MET
D. Marqua
Heard Island and McDonald Islands
Henninger Flats
Angeles National Forest
11
12
1
paratype
1969-05-29
1994-10
1969-05-29
R, UCR, MET
Vetter & M. Thompson
Heard Island and McDonald Islands
Hollywood
Lady Bug Canyon
11
12
1
paratype
1981-05
CDFA
G. Bakker
Heard Island and McDonald Islands
Los Angeles
11
12
1
paratype
UCR
Malibu & M. Balducci
Heard Island and McDonald Islands
11
12
1
paratype
1982-12-11
CGL
G. Lowe
Heard Island and McDonald Islands
Malibu Lagoon
Serra Road
11
12
1
paratype
1987-04-15
CGL
G. Lowe
Heard Island and McDonald Islands
1859
Marker Flats
Mount Baldy Road
11
12
1
paratype
1983-12-04
CGL
G. Lowe
Heard Island and McDonald Islands
12
13
Millard Canyon
Altadena
11
12
1
paratype
1984-07-05
CGL
Lowe
Heard Island and McDonald Islands
12
13
1
paratype
2002-01-18
2002-02-28
2002-01-18
WRI
W. Icenogle & J. Bond
Heard Island and McDonald Islands
Altadena
488
Millard Canyon
San Gabriel Mountains
12
13
1
paratype
1982-11-13
CGL
G. Lowe
Heard Island and McDonald Islands
Route
Old Topanga Canyon Road
Topanga Canyon Boulevard
12
13
1
paratype
1989-09-09
CBH
Hebert
Heard Island and McDonald Islands
12
13
1
paratype
1998-07-16
UCR
A. Moore
Heard Island and McDonald Islands
Palmer Canyon Road
Claremont
12
13
1
paratype
1966-05-02
UCR
C. Goodpasture
Heard Island and McDonald Islands
Pasadena
12
13
1
paratype
1972-07-26
WRI
F. Hovore
Heard Island and McDonald Islands
472
Placerita Canyon Park
12
13
1
paratype
1976-05-27
MET
Sullivan
Heard Island and McDonald Islands
12
13
1
paratype
S Roland Heights
Heard Island and McDonald Islands
Puente Hills
12
13
1
paratype
1956-07-01
2002-11-08
1956-07-01
R, AMNH, V, WRI
Fisher & San Antonio Canyon & Claremont & W. Gertsch & Roth & S Lower San Antonio Forest Station & W. Icenogle
Heard Island and McDonald Islands
San Gabriel Mountains
294
33.969017
Mount Baldy Road
1
-117.92689
Lower San Antonio Forest Station
12
13
1
paratype
2001-07-09
WRI
E Sunset Peak & W. Icenogle
Heard Island and McDonald Islands
1158
San Antonio Canyon
Mount Baldy Road
12
13
1
paratype
1956-09-02
2002-05-31
1956-09-02
V, AMNH, UCR
NE Big Dalton Reservoir & M. Oxford
Heard Island and McDonald Islands
Volfe Canyon
585
San Dimas Canyon
Big Dalton Canyon Road
12
13
1
paratype
1989-11-12
CBH
B. Hebert
Heard Island and McDonald Islands
Santa
San Gabriel Mountains
12
13
1
paratype
1991-04-23
1991-05-24
1991-04-23
CBH
Hebert
Heard Island and McDonald Islands
12
13
1
paratype
1941-03-20
AMNH
Ivie
Heard Island and McDonald Islands
15 mi W Santa Monica
12
13
1
paratype
1969-04-26
MET
M. Thompson
Heard Island and McDonald Islands
San Gabriel Mountains
Shortcut
Highway
12
13
1
paratype
1977-06
1977-07-31
1977-06
LACM
G. Gorelick
Heard Island and McDonald Islands
Sierra Madre
12
13
1
paratype
2001-07-05
WRI
W. Icenogle
Heard Island and McDonald Islands
San Gabriel Mountains
1097
Spruce Canyon
Mount Baldy Road
12
13
1
paratype
1951-05-19
1985-06-27
1951-05-19
T, AMNH
Tice & L. Sorkin
Heard Island and McDonald Islands
Angeles National Forest
1158
Sunland
Highway
12
13
1
paratype
2002-04-15
2002-04-29
2002-04-15
WRI
W. Icenogle
Heard Island and McDonald Islands
San Gabriel Mountains
1006
Switzer Picnic Area
Highway
12
13
1
paratype
1952-06-20
AMNH, JCC
Cazier & W. Gertsch & R. Schrammel
Heard Island and McDonald Islands
Tanbark Flats
San Gabriel Mountains
12
13
1
paratype
[700,1105,352,373]
1957-03-30
AMNH
Heard Island and McDonald Islands
12
13
1
paratype
1955-03-20
1995-04-10
1955-03-20
R, AMNH, CBH
Apr. & Schick & B. Olodort
Heard Island and McDonald Islands
Santa Monica Mountains
Tapia Park
Pacific Palisades
12
13
1
paratype
Heard Island and McDonald Islands
Topanga
Highway
12
13
1
paratype
2001-08-27
2004-03-13
2001-08-27
R, WRI
D. Palmer & J. Starrett & W. Icenogle
Heard Island and McDonald Islands
San Gabriel Mountains
470
34.094837
Keith
1
-118.6015
Shortcut Ranger Station
12
13
1
paratype
Aliso Wood Canyon & E Laguna Beach
Heard Island and McDonald Islands
Orange Co.
Orange Co.
12
13
1
paratype
2001-06-10
2001-08-31
2001-06-10
R, AMNH
Fisher & N Aliso Wood Canyon, E
Heard Island and McDonald Islands
71
33.558456
Laguna Beach
1
-117.74961
12
13
1
paratype
2000-06-01
2000-08-09
2000-06-01
R, AMNH
Fisher
Heard Island and McDonald Islands
80
33.567577
Chino Hills
1
-117.74826
Telegraph Canyon
12
13
1
paratype
2001-10-29
2001-11-02
2001-10-29
R, AMNH
Fisher
Heard Island and McDonald Islands
238
33.91449
Chino Hills
1
-117.78556
Telegraph Canyon
12
13
1
paratype
1999-09-07
1999-09-17
1999-09-07
R, AMNH
Fisher
Heard Island and McDonald Islands
358
33.91197
Crow Canyon
1
-117.78911
12
13
1
paratype
1931-06-24
2000-12-08
1931-06-24
R, AMNH
W. Ivie
Heard Island and McDonald Islands
338
33.602898
Fisher
1
-117.54428
Laguna Beach
12
13
1
paratype
1931-07-24
AMNH
Ivie
Heard Island and McDonald Islands
12
13
1
paratype
1932-12-28
AMNH
Ivie
Heard Island and McDonald Islands
12
13
1
paratype
[806,1098,1208,1229]
2000-08-17
UCR
Heard Island and McDonald Islands
12
13
1
paratype
2001-06-18
2001-06-22
2001-06-18
R, AMNH
Fisher & E The Sinks
Heard Island and McDonald Islands
341
33.73607
Limestone Canyon Regional Park
1
-117.67049
12
13
1
paratype
1999-07-20
1999-07-30
1999-07-20
R, AMNH
Fisher
Heard Island and McDonald Islands
453
33.716217
Limestone Canyon Regional Park
1
-117.65893
The Sinks
12
13
1
paratype
1998-06-01
1998-09-03
1998-06-01
R, AMNH
Fisher & N Niguel Hill, NW
Heard Island and McDonald Islands
378
33.722008
Laguna Niguel
1
-117.665794
12
13
1
paratype
Aug. 27 - 31, 2001, Jan. 28 - Feb. 1, 2002
R, AMNH
Fisher & N Niguel Hill, NW
Heard Island and McDonald Islands
195
33.53621
Laguna Niguel
1
-117.730156
12
13
1
paratype
2000-07-10
2000-08-31
2000-07-10
R, AMNH
Heard Island and McDonald Islands
132
33.53883
San Joaquin Hills
1
-117.72545
Laguna Canyon
12
13
1
paratype
2002-05-11
2002-09-13
2002-05-11
R, AMNH, WRI
Fisher & W. Icenogle
Heard Island and McDonald Islands
Santa Ana Mountains
175
13
14
33.59469
Highway
1
-117.7753
Hot Spring Canyon
12
13
1
paratype
1955-11-20
R, AMNH
A. Schick
Heard Island and McDonald Islands
Starr Ranch
Silverado Canyon
Santa Ana Mountains
13
14
1
paratype
1998-09-05
1998-10-14
1998-09-05
R, AMNH
Fisher & N Temple Hill & W Aliso Wood Canyon
Heard Island and McDonald Islands
297
33.630257
1
-117.553085
13
14
1
paratype
2000-07-10
2000-07-20
2000-07-10
R, AMNH
Fisher & N Temple Hill & W Aliso Wood Canyon
Heard Island and McDonald Islands
215
33.56991
1
-117.75634
13
14
1
paratype
2002-05-13
2002-05-17
2002-05-13
R, AMNH
Fisher
Heard Island and McDonald Islands
206
33.583847
Weir Canyon
1
-117.750305
13
14
1
paratype
1999-06-15
1999-06-25
1999-06-15
R, AMNH
Fisher
Heard Island and McDonald Islands
236
33.809948
Weir Canyon
1
-117.755646
13
14
1
paratype
1976-11-13
2000-09-29
1976-11-13
R, AMNH, MET
Fihser & M. Long
Heard Island and McDonald Islands
Riverside Co.
232
33.81556
Bay Tree Springs
1
-117.74714
San Jacinto Mountains
13
14
1
paratype
1960-04-20
UCB
J. Powell
Heard Island and McDonald Islands
Chino Canyon
13
14
1
paratype
2002-06-04
WRI
S Dark Canyon Campground & W. Icenogle
Heard Island and McDonald Islands
Hwy
San Jacinto Mountains
13
14
1
paratype
2002-06-05
WRI
Icenogle
Heard Island and McDonald Islands
13
14
1
paratype
2003-06-04
WRI
Icenogle
Heard Island and McDonald Islands
1757
13
14
1
paratype
1977-05-10
WRI
W. Icenogle
Heard Island and McDonald Islands
610
De Luz Road
13
14
1
paratype
1952-06-17
AMNH
Gertsch
Heard Island and McDonald Islands
13
14
1
paratype
1953-07-07
V, AMNH, UCR
W. Gertsch & Roth
Heard Island and McDonald Islands
1829
13
14
1
paratype
2002-06-29
CDU
D., S. Ubick & M. Harvey
Heard Island and McDonald Islands
Highway
168
James Reserve
San Jacinto Mountains
13
14
1
paratype
UCR
J. Kempf
Heard Island and McDonald Islands
Highway
1640
Lake Fulmor
San Jacinto Mountains
13
14
1
paratype
1968-08
1968-09
1968-08
D. Bixler & Bixler & W Temecula
Heard Island and McDonald Islands
Santa Margarita Ecological Reserve
13
14
1
paratype
1999-09-07
1999-09-17
1999-09-07
R, AMNH
Fisher & W Temecula
Heard Island and McDonald Islands
393
33.445236
Santa Margarita Ecological Reserve
1
-117.16446
13
14
1
paratype
1998-05-11
1998-05-21
1998-05-11
R, AMNH
Fisher & W Temecula
Heard Island and McDonald Islands
786
33.455048
Santa Margarita Ecological Reserve
1
-117.17124
13
14
1
paratype
2000-01-28
2001-08-12
2000-01-28
R, AMNH, WRI
Fisher & W. Icenogle
Heard Island and McDonald Islands
San Jacinto Mountains
788
33.456898
Strawberry Creek
1
-117.16846
Highway
13
14
1
paratype
1997-02
UCR
J. Klotz
Heard Island and McDonald Islands
San Bernardino Co.
1524
Crestline
San Bernardino Mountains
13
14
1
paratype
2001-08-30
WRI
W. Icenogle
Heard Island and McDonald Islands
San Bernardino Mountains
1280
Crestline area
Highway
13
14
1
paratype
1999-07-06
WRI
W. Icenogle
Heard Island and McDonald Islands
San Bernardino Mountains
1524
East Fork Mountain Home Creek
Highway
13
14
1
paratype
1942-07-16
AMNH
E. Schlinger
Heard Island and McDonald Islands
Fontana
13
14
1
paratype
1971-06-11
1985-08-10
1971-06-11
MCZ
D. Bixler & D. Lowrie
Heard Island and McDonald Islands
Hill Creek
2134
Forest Falls
Forest Home
13
14
1
paratype
Heard Island and McDonald Islands
Glass Road
San Bernardino Mountains
13
14
1
paratype
1999-07-10
2003-04-01
1999-07-10
T, UCR, CBH
East Fork Mountain, E & Prentice & B. Hebert
Heard Island and McDonald Islands
Lake Arrowhead Village
1820
38.34811
Highway
Highway
13
14
1
paratype
1936-05-06
AMNH
S. Bishop
Heard Island and McDonald Islands
Lake Arrowhead
13
14
1
paratype
1970-08-20
1992-05-14
1970-08-20
CBH
D. Bixler & Manker Flats Campground & B. Hebert
Heard Island and McDonald Islands
San Gabriel Mountains
914
Lytle Creek Canyon
Lytle Creek
13
14
1
paratype
1969-05-15
1969-05-28
1969-05-15
WRI
W. Icenogle
Heard Island and McDonald Islands
San Bernardino Mountains
1829
Mill Creek
Falls Picnic Area
13
14
1
paratype
1969-06-18
WRI
Icenogle
Heard Island and McDonald Islands
1829
13
14
1
paratype
1959-08-13
1995-07-21
1959-08-13
V, AMNH, UCR
W. Gertsch & Summer & Silverwood Lake State Recreation Area, SE
Heard Island and McDonald Islands
Silverwood Lake
Mountain Home Creek
Silverwood Lake State Recreation Area
13
14
1
paratype
2001-06-10
2002-08-16
2001-06-10
R, AMNH, WRI
Fisher & W. Icenogle
Heard Island and McDonald Islands
San Bernardino Mountains
1035
34.277767
Skinner Creek
1
-117.31695
Forest Falls
13
14
1
paratype
1983-12-28
1984-01-26
1983-12-28
UCR
M. Narog
Heard Island and McDonald Islands
1494
14
15
38.34807
Skinner Ridge
Highway
13
14
1
paratype
1995-07-12
UCR
Heard Island and McDonald Islands
Smiley Park
Running Springs
14
15
1
paratype
1995-07-10
UCR
J. Wambaugh
Heard Island and McDonald Islands
Smiley Park
Fredalba Road
14
15
1
paratype
UCR
Twin Peaks & K. Wormald
Heard Island and McDonald Islands
San Bernardino Mountains
14
15
1
paratype
1996-06
UCR
K. Wormald
Heard Island and McDonald Islands
May
14
15
1
paratype
UCR
Wormald
Heard Island and McDonald Islands
14
15
1
paratype
1996-08
1996-09-31
1996-08
UCR
Wormald
Heard Island and McDonald Islands
1676
14
15
1
paratype
1997-06
1997-06-11
1997-06
R, UCR
Vetter & K. Wormald
Heard Island and McDonald Islands
May
14
15
1
paratype
1984-05-19
CGL
De Luz Road & G. Lowe
Heard Island and McDonald Islands
San Diego Co.
1 mi N Fallbrook
14
15
1
paratype
[124,544,592,613]
1984-08-25
CGL
Lowe
Heard Island and McDonald Islands
14
15
1
paratype
1984-08-04
UCR
D. Boe
Heard Island and McDonald Islands
Fry Creek
Mount
14
15
1
paratype
1956-07-30
V, AMNH
W. Gertsch & Roth
Heard Island and McDonald Islands
Lake Henshaw
Henshaw
Cleveland National Forest
14
15
1
paratype
2004-05-16
AMNH
Palmer & P. Paquin & T. Prentice & C. Vink
Heard Island and McDonald Islands
308
33.24439
1
-116.764114
14
15
1
paratype
1997-06-03
1997-07-02
1997-06-03
UCR
J. George
Heard Island and McDonald Islands
Mendenhall Valley
14
15
1
paratype
1997-07-02
1997-08-07
1997-07-02
UCR
George
Heard Island and McDonald Islands
1829
14
15
1
paratype
1931-07-26
1967-07-04
1931-07-26
R, AMNH, V, WRIK
Chamberlin & W. Gertsch & B. Kaston
Heard Island and McDonald Islands
1219
Mount Palomar
Roth
14
15
1
paratype
2004-05-16
AMNH
Palmer & P. Paquin & T. Prentice & C. Vink
Heard Island and McDonald Islands
1500
33.34588
1
-116.87948
14
15
1
paratype
1953-07-13
AMNH
W., J. Gertsch
Heard Island and McDonald Islands
State Park
14
15
1
paratype
1953-07-17
2003-02-20
1953-07-17
R, UCR, V, AMNH
Vetter & Cold Spring Trail & W. Gertsch & Roth & W., J. Gertsch
Heard Island and McDonald Islands
Santa Barbara Co.
1414
State Park
Santa Barbara
14
15
1
paratype
UCR
May & D. Boe
Heard Island and McDonald Islands
Refugio Rd.
Apr.
14
15
1
paratype
1950-11
1950-12-31
1950-11
AMNH
H. Shantz
Heard Island and McDonald Islands
Santa Barbara
14
15
1
paratype
1953-07-17
AMNH
Shantz
Heard Island and McDonald Islands
14
15
1
paratype
1970-07-27
CDFA
Cravens
Heard Island and McDonald Islands
14
15
1
paratype
1934-07-12
AMNH
Ivie
Heard Island and McDonald Islands
10 mi W Santa Barbara
14
15
1
paratype
1997-05-11
1997-06-11
1997-05-11
CAS
E., M. Schlinger
Heard Island and McDonald Islands
427
Sedgwick Ranch Reserve
14
15
1
paratype
1997-06-24
1997-07-21
1997-06-24
CAS
Schlinger
Heard Island and McDonald Islands
351
14
15
1
paratype
1997-05-28
CAS
E. M. Schlinger
Heard Island and McDonald Islands
610
Aug.
Nov.
14
15
1
paratype
1975-04-26
1975-05-18
1975-04-26
WRI
F. Hovore
Heard Island and McDonald Islands
Upper Oso Campground
Santa Ynez River
14
15
1
paratype
1982-07-03
CAS
D. Burdick
Heard Island and McDonald Islands
Ash Mountain
Kaweah Power Station
14
15
1
paratype
1983-03-12
CAS
Burdick
Heard Island and McDonald Islands
14
15
1
paratype
1983-06-19
1983-06-21
1983-06-19
CAS
Haines
Heard Island and McDonald Islands
14
15
1
paratype
1983-07-06
1983-07-10
1983-07-06
CDU
Burdick
Heard Island and McDonald Islands
14
15
1
paratype
[778,1225,352,373]
1983-07-17
CAS
Burdick
Heard Island and McDonald Islands
14
15
1
paratype
1983-07-31
1995-09-03
1983-07-31
R, CDU, CDFA, CAS
D. Burdick
Heard Island and McDonald Islands
14
15
1
paratype
1996-07-24
CAS
Burdick
Heard Island and McDonald Islands
14
15
1
paratype
1956-07-05
1958-07-09
1956-07-05
V, AMNH
W. Gertsch & Roth & NE Case Mountain Peak
Heard Island and McDonald Islands
Mountain Peak
1128
Ash Mountain
Case Mountain
14
15
1
paratype
2003-07-14
2003-07-18
2003-07-14
R, AMNH
Fisher & NE Case Mountain Peak
Heard Island and McDonald Islands
1691
36.417908
1
-118.7933
14
15
1
paratype
2003-07-14
2003-07-18
2003-07-14
R, AMNH
Fisher & NE Case Mountain Peak
Heard Island and McDonald Islands
1646
36.421837
Mountain Grove
1
-118.79371
14
15
1
paratype
2002-07-14
2002-07-22
2002-07-14
R, AMNH
Fisher & Case Mountain Peak, E
Heard Island and McDonald Islands
1810
36.413177
Salt Creek
1
-118.79191
14
15
1
paratype
2002-07-19
2002-07-22
2002-07-19
R, AMNH
Fisher
Heard Island and McDonald Islands
1644
36.420357
1
-118.793915
14
15
1
paratype
2002-07-19
2002-07-22
2002-07-19
R, AMNH
Fisher & NW Case Mountain Peak, S
Heard Island and McDonald Islands
1819
36.42109
Salt Creek
1
-118.79977
14
15
1
paratype
2002-07-14
2002-07-22
2002-07-14
R, AMNH
Fisher & NW Case Mountain Peak, S
Heard Island and McDonald Islands
1855
36.41443
Salt Creek
1
-118.81382
14
15
1
paratype
2003-07-14
2003-07-18
2003-07-14
R, AMNH
Fisher
Heard Island and McDonald Islands
1709
36.417816
Case Mountain
1
-118.819305
Mountain Grove
14
15
1
paratype
2003-07-14
2003-07-18
2003-07-14
R, AMNH
Fisher
Heard Island and McDonald Islands
1678
36.40373
Case Mountain
1
-118.781906
Mountain Grove
14
15
1
paratype
2002-07-19
2002-07-22
2002-07-19
R, AMNH
Fisher & SE Case Mountain Peak
Heard Island and McDonald Islands
1805
36.405117
Salt Creek
1
-118.18983
14
15
1
paratype
2002-07-19
2002-07-22
2002-07-19
R, AMNH
Fisher
Heard Island and McDonald Islands
1801
36.397408
1
-118.77598
14
15
1
paratype
1952-07-17
2002-07-22
1952-07-17
R, AMNH
Fisher & W. Gertsch
Heard Island and McDonald Islands
1848
36.398876
Kaweah River
1
-118.78779
14
15
1
paratype
1956-07-03
2004-04-28
1956-07-03
V, AMNH, CDU
W. Gertsch & D. Ubick
Heard Island and McDonald Islands
Kaweah Cave
914
15
16
Mineral King Road
Sequoia National Park
14
15
1
paratype
1958-07-11
2004-05-15
1958-07-11
V, AMNH, UCR
Soda Creek, W & W. Gertsch & Ojai & D. Boe
Heard Island and McDonald Islands
Ventura Co.
Soda Creek
Camp Nelson
15
16
1
paratype
1980-05-06
UCB
C. Griswold
Heard Island and McDonald Islands
1219
Reyes Creek
Woods
15
16
1
paratype
1986-07-04
CBH
N Santa Paula & B. Hebert
Heard Island and McDonald Islands
198
Stecker Park
Highway
15
16
1
paratype
1958-07-02
1983-06-26
1958-07-02
V, AMNH, CAS, CGL
W. Gertsch & Roth & G. Lowe
Heard Island and McDonald Islands
Wheeler Springs
Yerba Buena Road
15
16
1
paratype
1946-01-24
1946-01-25
1946-01-24
J. Linsdale
Hastings Natural History State Reserve
15
16
2
California
holotype
1992-09-10
CDU
Ubick & J. Boutin
110
16
17
1
Arroyo Seco & Indians Road
Los Padres National Forest
16
17
1
1995-05-07
CDU
Ubick & W. Savary
335
16
17
1
1995-07-08
CDU
Ubick & S. Fend & W. Savary
16
17
1
1996-08-24
CDU
Ubick
16
17
1
1999-07-03
CDU
Fend & J. Ledford & D. Ubick
16
17
1
2000-04-15
CDU
Ubick
16
17
1
2001-04-28
CDU
Ubick
16
17
1
2001-04-27
2001-08-19
2001-04-27
CDU
Ubick
335
16
17
1
2004-06-18
2004-10-24
2004-06-18
Arroyo Seco Campground
Los Padres National Forest
16
17
1
1980-04-26
CDU
Ubick
16
17
1
1994-05-13
CDU
Ubick & J. Boutin
290
16
17
1
1994-05-13
1994-05-15
1994-05-13
CDU
Boutin
274
16
17
1
1999-04-24
CDU
Ubick & W. Savary
16
17
1
Arroyo Seco Campground & S The Lakes
Los Padres National Forest
16
17
1
1994-05-13
1994-05-15
1994-05-13
CDU
Ubick & J. Boutin
274
16
17
1
1995-05-06
CDU
Ubick & W. Savary
16
17
1
2001-04-27
2001-05-18
2001-04-27
Ubick
16
17
1
2004-06-18
2004-10-24
2004-06-18
Same
Big Sur
16
17
1
1937-07
AMNH
Ivie
16
17
1
1952-06-25
R, AMNH
M. Cazier & W. Gertsch & Schrammel
16
17
1
1953-07-18
AMNH
Gertsch
16
17
1
1960-04-02
R, AMNH
W. Gertsch & W. Ivie & Schrammel
14 mi S Big Sur
16
17
1
1953-12-22
V, AMNH
Roth
Carmel
16
17
1
1953-11-12
AMNH
16
17
1
[293,648,1451,1472]
1954-04-05
1954-05-23
1954-04-05
AMNH
16
17
1
[124,386,1477,1499]
1 mi N Carmel
16
17
1
1953-12-21
V, AMNH
Roth
Carmel Valley
16
17
1
1979-04-09
UCR
Boe
16
17
1
1996-05
CDU
Stadille
16
17
1
1991-06-01
CDU
Ubick
914
16
17
1
Hastings Natural History State Reserve
16
17
1
1939-05-19
1939-05-29
1939-05-19
CAS
Linsdale
16
17
1
1940-05-30
CAS
Linsdale
16
17
1
[246,602,1720,1741]
1941-02-22
CAS
Linsdale
16
17
1
[700,1075,192,213]
1942-05-27
CAS
Linsdale
16
17
1
1944-07-18
CAS
Linsdale
16
17
1
1946-01-24
1946-01-25
1946-01-24
AMNH
Linsdale
16
17
1
1946-04-10
CAS
Linsdale
16
17
1
1946-05-17
AMNH
Linsdale
16
17
1
1946-11-18
AMNH
Linsdale
16
17
1
1946-12-08
AMNH
Linsdale
16
17
1
[123,544,192,213]
CAS
Linsdale
17
18
1
1947-06
AMNH
Linsdale
17
18
1
1948-05-24
AMNH
Linsdale
17
18
1
1950-05-17
CAS
Linsdale
17
18
1
1950-06-24
1999-07-23
1950-06-24
AMNH, CDU
J. Linsdale & S. Fend & Ledford & Ubick
17
18
1
1999-08-02
1999-08-04
1999-08-02
CAS
D. Ubick & S. Fend & Ledford
17
18
1
2004-01-09
AMNH
Stewart
578
36.388996
Hastings Natural History State Reserve
1
-121.548615
Big Creek
17
18
1
1979-04-10
UCR
Highway G & Boe
17
18
1
1982-06-04
1982-06-06
1982-06-04
UCB
Griswold
Hill Big Creek Reservoir
17
18
1
2002-09-06
UCR
Edwards
17
18
1
1991-05-31
CDU
Ubick
305
Nacimiento Road
Kirk Creek
17
18
1
1991-06-01
CDU
Ubick
472
Nacimiento Road
Fork Mill Creek
17
18
1
1931-08-15
AMNH
Ivie
Pacific Grove
17
18
1
1964-05-03
CAS
Craig
91
Pfeifer Big Sur State Park
17
18
1
1987-08-19
CAS
Gross
17
18
1
1960-04-02
1961-09-25
1960-04-02
R, AMNH
Gertsch & Ivie & Schrammel & W. Gertsch & W. Ivie
Redwood Canyon
Gorda
17
18
1
1978-09-13
2002-05-27
1978-09-13
R, AMNH, UCB
Doty & Lew
Whale Point Cabin
200
Tassajara Spring Buddhist
Big Creek Reserve
17
18
1
2002-10-24
UCB
Lew
17
18
1
2003-05-12
UCB
Lew
17
18
1
1958-07-03
V, AMNH
W. Gertsch & Roth
San Benito Co.
Pinnacles National Monument
June
17
18
1
1972-01-22
A. Jung
Inyo Co.
Titus Canyon Cave
Male
17
18
2
California
holotype
San Bernardino Co.
Medicine Cave
18
19
1
California
1972-01-22
1988-09-07
1972-01-22
R, CAS, AMNH
Aalbu & Death Valley National Monument & A. Jung
Inyo Co.
1219
Stone Canyon
Defense Cave
18
19
1
California
[124,469,1224,1247]
Inyo Co.
Inyo Co.
18
19
1
California
1979-03-17
1979-06-16
1979-03-17
R
San Bernardino Co.
1311
Mitchell Caverns State Park
Park Headquarters
18
19
1
California
holotype
1988-06-20
1988-09-27
1988-06-20
R, CAS
Mountains State Recreation Area
19
20
1
San Bernardino Co.
San Bernardino Co.
19
20
1
California
1961-04-21
J. Beatty
Pinal Co.
Peppersauce Cave Canyon
Male
19
20
1
Arizona
holotype
1952-07-25
AMNH
M. Cazier
Coconino Co.
Oak Creek Canyon
Manzanita Camp
19
20
1
OTHER MATERIAL EXAMINED
1956-07-14
V, AMNH
W. Gertsch & Roth
Mexico
Graham Co.
Mount Graham
Lower Canyon
19
20
1
ARIZONA
1973-09-11
V, CAS
Roth
Mexico
Cajon Bonito
19
20
1
Sonora