Anachemmis Chamberlin, 1919: 12 A. sober Chamberlin Anachemmis sober Chamberlin, 1919: 13 Silva, 2003: 41 Anachemmis scaber Roewer, 1955: 607 A Revision of the North American Spider Genus Anachemmis Chamberlin (Araneae, Tengellidae) PLATNICK, NORMAN I. UBICK, DARRELL American Museum Novitates 2005 2005-05-11 3477 1 20 62BWS Chamberlin Chamberlin. In 1919 [797,1129,1335,1359] Arachnida Zoropsidae Anachemmis Animalia Araneae 1 2 Arthropoda genus      Anachemmis Chamberlin, 1919: 12( typespecies by original designation  A. soberChamberlin).  DIAGNOSIS: The unusual character combination of three claws plus claw tufts (fig. 1) readily separates members of  Anachemmisand its closest relatives from most other North American spiders. Specimens of  Zorocrates, which are found from Arizonato Texasand south to Central America, have a tiny remnant of the third claw that is equally obscured by the distal hairs of the tarsal scopula, but  Zorocratesspecieslack true claw tufts (Griswold et al., in press) and have a single retrolateral tibial apophysis on the male palp. Males of  Anachemmisdiffer from those of  Titiotusin having two rather than three or four prongs on the retrolateral tibial apophysis, and from those of  Socalchemmisand  Liocranoidesby having a flattened, blade­shaped ventral prong of the apophysis and by having the dorsal prong of the apophysis reduced to a small point. The female epigynum differs from those of  Liocranoidesspeciesin lacking an anterior hood, from those of  Titiotusspeciesin having a much smaller septum, and from those of  Socalchemmisspeciesin having  Liocranoides­like anterior vulval elaborations.  DESCRIPTION: Medium to large spiders, total length of males 7–10, of females 8–13. Carapace oval, widest at rear of coxae II, abruptly narrowed at level of palpi to less than two­thirds of maximum width; thoracic groove long, longitudinal, very deep; surface coated with short recumbent and fewer, longer, erect dark setae, erect setae most numerous along midline and in ocular area; eight eyes in two rows; from above, both eye rows slightly recurved; from front, anterior row recurved, posterior row slightly pro­ curved; anterior median eyes round, smallest; other eyes oval, subequal, with canoe­shaped tapeta; anterior median eyes separated by roughly their diameter, slightly closer to anterior laterals; posterior medians separated by roughly their diameter, farther from posterior laterals; lateral eyes of each side separated by less than their diameter; median ocular quadrangle wider in back than in front, wider in back than long; clypeal height about twice diameter of anterior median eyes, corners of clypeus with incised margins that overlie cheliceral boss; chilum weakly sclerotized, divided, composed of two triangular sclerites. Chelicerae vertical, anterior surface with few, erect setae; promargin with three teeth situated at proximal end of fang furrow, most proximal tooth smallest, retromargin with three larger, more distally situated teeth; very short, narrow, I­shaped posterior sclerite present, separating chelicerae at base. Labium short, distally invaginated at middle, reflexed at almost 908 angle relative to sternum. Endites rectangular, distally slightly convergent, with anteromedian scopula and anterolateral serrula consisting of single row of teeth. Sternum rounded, without extensions to or between coxae, with erect setae; posterior margin only slightly extended between coxae IV.  Figs. 13–16.  Anachemmis linsdalei, new species, male. 13.Spinnerets of left side, distal view. 14.Left anterior lateral spinneret, distal view. 15.Left posterior median spinneret, distal view. 16.Left posterior lateral spinneret, distal view. Scale bar 5 43 mm (14–16), 150 mm (13).  Figs. 17–20.  Anachemmis linsdalei, new species, male. 17.Left palp, prolateral view, showing interlocking lobes on tegulum and subtegulum. 18.Left palp, retrolateral view. 19.Left palp, ventral view. 20.Left embolus, conductor, and median apophysis, ventral view. Scale bar 5 200 mm (20), 600 mm (17–19).  Figs. 21–24.  Anachemmis soberChamberlin. 21.Left male palp, ventral view. 22.Same, retrolateral view. 23.Epigynum, ventral view. 24.Same, dorsal view. Leg formula 4123. Typical leg spination pattern (only surfaces bearing spines listed): femora: I d1­1­1, p01­1, r1­2­1; II d1­1­1, p0­2­1, r1­2­1; III d1­1­1, p2­1­1, r1­2­1; IV d1­1­1, p2­1­1, r0­0­1; patellae III, IV p0­1­ 0, r0­1­0; tibiae: I, II p0­1­0, v4­4­2; III, IV d1­0­1, p0­1­1, v2­2­2, r0­1­1; metatarsi: I, II v2­2­2; III p1­1­2, v2­2­1r, r1­1­2; IV p1­ 1­2, v2­2­1r, r1­2­2; tarsi with three claws and claw tufts (figs. 1–4), superior claws with several weak teeth, most distal teeth largest, inferior claws unarmed; all tarsi with strong ventral scopulae, scopular hairs distinct from those of claw tufts (fig. 1); distal segments with trichobothria in two rows, bases ridged (figs. 5, 6); tarsal organ capsulate (fig. 8); all trochanters strongly notched; males without tibial crack; metatarsi without preening combs. Abdomen without anterior or dorsal scutum; anterior lateral spinnerets large, composed of two articles (figs. 9, 13), distal article with two major ampullate gland spigots and about 10 small piriform gland spigots (figs. 10, 14); posterior median spinnerets composed of one article, small, triangular (figs. 11, 15), those of female expanded posteriorly, where they bear single large and three or four smaller cylindrical gland spigots, preceded anteriorly by smaller aciniform spigots and at least two larger minor ampullate gland spigots; posterior lateral spinnerets composed of two articles, distal article about one­fifth as long as proximal article, that of female apparently without cylindrical gland spigots at base, in both sexes with smaller spigots around base and on tip (figs. 12, 16); colulus represented by setae on small lobe.  Figs. 25–28.  Anachemmis linsdalei, new species. 25.Left male palp, ventral view. 26.Same, retrolateral view. 27.Epigynum, ventral view. 28.Same, dorsal view. Male palp with patella not widened, retrolateral tibial apophysis bipartite, ventral prong large, blade­shaped (fig. 18), dorsal prong reduced to tiny point; subtegulum and tegulum with interlocking lobes (fig. 17), median apophysis heavily sclerotized at least distally, embolus short, with shallow division between prolateral prong and copulatory prong, accompanied by hyaline conductor (figs. 19, 20). Female palp with extremely long, dentate claw. Epigynum with large median septum wider anteriorly than posteriorly; spermathecal bulbs situated posterolaterally. MISPLACED SPECIES:  Anachemmis dolichopus Chamberlin (1919)is now placed in  Socalchemmis Platnick and Ubick (2001). For unknown reasons, Roewer (1955)listed  Chemmis unicolor Banks (1901)as a member of  Anachemmis, but that transfer is incorrect; the placement of Banks’ species will be discussed in detail in a subsequent paper in this series.  KEY TO SPECIES OF  ANACHEMMIS    1. Males............................. 2  — Females........................... 6   2. Tegulum much wider opposite median apophysis than elsewhere (figs. 21, 25)..... 3  — Tegulum not much wider opposite median apophysis than elsewhere (figs. 29, 33, 37) ................................ 4   3. Tegular edge opposite median apophysis base rounded (fig. 21)..............  sober  — Tegular edge opposite median apophysis base pointed (fig. 25).............  linsdalei   4. Ventral prong of retrolateral tibial apophysis relatively wide at base (fig. 34)..  aalbui  — Ventral prong of retrolateral tibial apophysis relatively narrow at base (figs. 30, 38)... ................................. 5  Figs. 29–32.  Anachemmis jungi, new species. 29.Left male palp, ventral view. 30.Same, retrolateral view. 31.Epigynum, ventral view. 32.Same, dorsal view.   5. Dorsal prong of retrolateral tibial apophysis relatively large (fig. 30)..........  jungi  — Dorsal prong of retrolateral tibial apophysis a tiny spur (fig. 38).............  beattyi   6. Widened anterior portion of median epigynal septum relatively short (figs. 31, 35).. 7  — Widened anterior portion of median epigynal septum relatively long (figs. 23, 27, 39).. ................................. 8   7. Epigynal surface opposite sides of median septum with distinct ridge (fig. 35)...................................  aalbui  — Epigynal surface opposite sides of median septum without distinct ridge (fig. 31).................................  jungi   8. Widened portion of epigynal septum with paramedian invaginations (fig. 39).................................  beattyi  — Widened portion of epigynal septum without paramedian invaginations (figs. 23, 27).. ................................. 9   9. Epigynal septum relatively wide both anteriorly and posteriorly (fig. 23).....  sober  — Epigynal septum relatively narrow both anteriorlyand posteriorly (fig. 27).................................  linsdalei   Anachemmis soberChamberlin    Figures 21–24; Map 1      Anachemmis sober Chamberlin, 1919: 13, pl. 5, fig. 5 (female holotypefrom Claremont, Los Angeles Co., California, in MCZ, examined).—  Silva, 2003: 41, fig. 13d (scanning electron micrograph of male palp).    Anachemmis scaber(lapsus):  Roewer, 1955: 607.  DIAGNOSIS: Males closely resemble those of  A. linsdaleibut differ in having a rounded tegular edge opposite the median apophysis (fig. 21); females also closely resemble those of  A. linsdaleibut have a wider epigynal septum and more anteriorly restricted lateral epigynal margins (fig. 23).  MALE: Total length 8.0. Carapace light brown, with darker triangular markings radiating from thoracic groove to intercoxal areas; abdomen mottled brownish gray; femora light brown, more distal leg segments grading to dark reddish brown on metatarsi and tarsi. Leg spination: femora: II p1­2­1; IV r0­ 0­2; tibiae: I d1­0­1, r0­1­0; II d0­0­1, r0­1­ 0; tibiae: I, II p1­1­0, r1­1­0; III p1­2­2; IV v2­2­2. Tegulum widened opposite median apophysis, with rounded edge (fig. 21); dorsal prong of retrolateral tibial apophysis reduced to tiny, sharp point tip (fig. 22).  Figs. 33–36.  Anachemmis aalbui, new species. 33.Left male palp, ventral view. 34.Same, retrolateral view. 35.Epigynum, ventral view. 36.Same, dorsal view. FEMALE: Total length 10.3. Coloration as in male. Leg spination: femora: III r1­1­2; IV r0­0­1; tibiae I p0­0­0; III p1­2­2; IV v2­2­ 2. Epigynum with median septum widened anteriorly and posteriorly, lateral margins restricted to anterior portion of epigynum (fig. 23); spermathecal bulbs oval (fig. 24). VARIATION: One male taken at Shortcut on the Angeles Crest Highway in the San Gabriel Mountains of Los Angeles County differs from other specimens in having a longer dorsal prong of the retrolateral tibial apophysis. One female from the nearby locality of Switzers Camp differs from most others in having the lateral epigynal bulges relatively ‘‘deflated’’. The geographic congruence of these specimens suggested that a separate species might be present in the San Gabriels, but thanks to the help of Wendell Icenogle, a good series of specimens from these and nearby localities has now been available for study. No other males show the abnormally long dorsal prong of the tibial apophysis, and no consistent differences have been detected among the palpal bulbs of any of these specimens. Another female shows the relatively ‘‘deflated’’ lateral epigynal bulges, but only on one side of the epigynum. We conclude that there is evidence for the existence of only a single species at these sites. Another male from Lake Fulmor in Riverside County has a relatively truncated left embolus tip, but is presumably either broken or represents a developmental deformity, as the right embolus of that specimen appears normal.  MATERIAL EXAMINED: CALIFORNIA: Inyo  Figs. 37–40.  Anachemmis beattyi, new species. 37.Left male palp, ventral view. 38.Same, retrolateral view. 39.Epigynum, ventral view. 40.Same, dorsal view.  Co.:  Ninemile Canyon,  June 11, 1993, elev. ca.  7000 ft( R. Robertson, CDU), 1/.  Kern Co.: Bakersfield,  June 28, 1985( S. Stockwell, S. Taber, O. Francke, AMNH), 1?, 1/.  Los Angeles Co.:Angeles Crest Highway, N La Canada, first canyon in Angeles National Forestc. 0.25 mi from ranger station,  June 27, 1982, running on leaf litter, shaded canyon slope ( G. Lowe, AMNH), 1?; Beverly Glen Canyon, Santa Monica Mountains,  May 9, 1953(W. McDonald, AMNH), 1/; Big Pines Highwayat Grand View Canyon, Angeles National Forest,  Jan. 27, 1982, under log, elev.  5900 ft( B. Hebert, CBH), 1/; Big Rock Creek, Valyermo,  June 12, 1943, elev.  4200 ft( K. Cowles, AMNH), 1/; Charlton­Chilao P. C., Angeles National Forest,  Apr. 28, 1969( M. Thompson, MET), 1/; Charmlee CountyPark,  May 13, 1975( M. Thompson, MET), 1?,  May 3, 1997, elev.  396 m(J. Kempf, UCR), 1?; Claremont( R. Chamberlin, MCZ), 1/ ( paratype),  Apr. 1913( R. Chamberlin, MCZ), 1/ ( holotype); Eaton Canyon Park,  Aug. 5, 1965( M. Thompson, MET), 1?,  Aug. 16, 1965(M. Thompson, MET), 1/,  Aug. 4, 1967(M. Thompson, MET), 1/,  Jan. 18, 1968(M. Thompson, MET), 2?,  Feb. 19, 1968(M. Thompson, MET), 1/,  May 5, 1968(M. Thompson, MET), 5?, 1/,  May 7, 1968(M. Thompson, MET), 1?; Glendale,  Nov. 25, 1950( E. Schlinger, AMNH), 1?,  May 25, 1958(E. Schlinger, AMNH), 1?,  May 10, 2002, in home (K. Schwellenbach, UCR), 1?; Gold Canyon, San Gabriel Mountains,  Mar. 1955, matured Apr.17 ( R. Schick, AMNH), 1?,  Apr. 17, 1955( R. Schick, AMNH), 1/,  May 6–8, 1955( R. Schick, AMNH), 1?, 2/; Grandview Canyon, Big Pines Highway, c.  1 miW Lake Jackson,  June 27, 1982, under log, elev.  5900 ft( G. Lowe, CGL), 1/; Henninger Flats, Angeles National Forest,  Nov. 11–15, 1968( D. Marqua, MET), 2/; Hollywood,  Oct. 1994( R. Vetter, UCR), 1/; Lady Bug Canyon,  May 29–June 8, 1969–1970 ( M. Thompson, MET), 4?; Los Angeles,  May 1981( G. Bakker, CDFA), 1?; Malibu, mid­April 2002 ( M. Balducci, UCR), 1?; Malibu Lagoon, near Serra Road,  Dec. 11, 1982, under rock ( G. Lowe, CGL), 1?; Marker Flats, Mount Baldy Road,  Apr. 15, 1987, under rock on slope, elev.  6100 ft( G. Lowe, CGL), 1/; Millard Canyon, Altadena,  Dec. 4, 1983, under stone on leaf litter ( G. Lowe, CGL), 1/,  July 5, 1984, on rocks and tree trunks at night (C. Lowe, CGL), 2?, 2/; Millard Canyon, at junction with Canyon Crest Road, San Gabriel Mountains, Altadena,  Jan. 18, 2002(matured  Feb. 28, 2002), elev.  1600 ft, under trash on oak duff near stream ( W. Icenogle, J. Bond, WRI), 1?; Old Topanga Canyon Road, c. 1 mifrom Topanga Canyon Boulevardand Route27,  Nov. 13, 1982, under rock, under cardboard debris on ground in oak forest ( G. Lowe, CGL), 2/,  Sept. 9, 1989, found dead in spider web (B. Hebert, CBH), 1/; 4723 Palmer Canyon Road, Claremont,  July 16, 1998, in kitchen sink ( A. Moore, UCR), 1?; Pasadena,  May 2, 1966( C. Goodpasture, UCR), 1?; Placerita Canyon Park,  3 miE Newhall,  July 26, 1972, elev.  1550 ft( F. Hovore, WRI), 1/;  May 27, 1976(P. Sullivan, MET), 1/; Puente Hills, S Roland Heights,  33.969018N, 117.926888W,  Aug. 25–Sept. 4, 1998, dry pitfall, elev.  294 m( R. Fisher, AMNH), 2/,  Nov. 4–8, 2002, same, 1/; San Antonio Canyon, near Claremont,  July 1, 1956( W. Gertsch, V. Roth, AMNH), 6?, 6/; San Antonio Canyon, Mount Baldy Road, 1.5 mi S Lower San Antonio Forest Station, San Gabriel Mountains,  May 4, 1996, dead and dried in black widow web in big pipe under road ( W. Icenogle, WRI), 1?; San Antonio Canyon, 0.5 mi E Sunset Peak, 0.25 mi W Mount Baldy Road,  July 9, 2001, elev.  3800 ft, under rock and log in leaf litter beside streamin shaded canyon bottom ( W. Icenogle, WRI), 1?, 1/; San Dimas Canyon, near Claremont,  Sept. 2, 1956( V. Roth, AMNH), 1/; San Dimas Experimental Forest, Big Dalton Canyon Road, 0.75 mi NE Big Dalton Reservoirat junction of Volfe Canyon, 348109560N, 1178479460W,  July 9, 2001(matured  May 25–31, 2002), elev.  1920 ft, oak leaf litter ( M. Oxford, UCR), 1?; SantaAnita Canyon, San Gabriel Mountains, collected as juvenile  Nov. 12, 1989, matured in captivity ( B. Hebert, CBH), 1?, collected  Apr. 23, 1991, matured  May 24, 1991(B. Hebert, CBH), 1?;  15 miW Santa Monica,  Mar. 20, 1941(W. Ivie, AMNH), 1?; Shortcut, Angeles Crest Highway, San Gabriel Mountains,  Apr. 26, 1969( M. Thompson, MET), 1?; Sierra Madre,  June–July 1977( G. Gorelick, LACM), 3?; Spruce Canyon(W branch, San Antonio Canyon), 0.25 mi W Mount Baldy Road, San Gabriel Mountains,  July 5, 2001, elev.  3600 ft, under log beside stream below oaks and alders ( W. Icenogle, WRI), 1?, 1/; Sunland,  May 19, 1951( T. Tice, AMNH), 1?; Switzers Camp, Angeles Crest Highway, Angeles National Forest,  June 27, 1985, elev.  3600–4000 ft( L. Sorkin, AMNH), 3/; Switzer Picnic Area, upper end of Arroyo Seco Canyon, 0.25 mi S Highway2, 0.5 mi E junction with HighwayN3, San Gabriel Mountains,  Apr. 15, 2002(matured  Apr. 27–29, 2002), elev.  3300 ft, under rock in duff below oaks and alders near stream at base of steep, rocky, north­facing canyon side ( W. Icenogle, WRI), 2?, 1/; Tanbark Flats, San Gabriel Mountains,  June 20, 1952(M. Cazier, W. Gertsch, R. Schrammel, AMNH, JCC), 95?, 130/,  Mar. 30, 1957( AMNH), 2/; Tapia Park, Santa Monica Mountains,  Mar. 20, 1955, matured in Apr.( R. Schick, AMNH), 1/; Temescal Canyon, Pacific Palisades, Santa Monica Mountains,  Apr. 10, 1995( B. Olodort, CBH), 1/; Topanga, off Highway27,  34.094838N, 118.601498W,  Mar. 13, 2004, elev.  470 m( R. Keith, D. Palmer, J. Starrett, CDB), 1/; Upper Big Tujunga Canyon, just E Shortcut Ranger Station, San Gabriel Mountains,  Aug. 27, 2001, elev.  4200 ft, dead and dried in web of  Steatoda atascadera(W. Icenogle, WRI), 6?, 1/,  June 11, 2002, elev.  4200 ft, on wall of 109 diameter pipe under road above small stream, conifer/oak forest ( W. Icenogle, WRI), 2?.  Orange Co.:N Aliso Wood Canyon, E Laguna Beach,  33.558458N, 117.749608W,  June 10–Aug. 31, 2001–2002, dry pitfalls, elev.  71 m( R. Fisher, AMNH), 2?, 4/; N Aliso Wood Canyon, E Laguna Beach,  33.567578N, 117.748258W,  June 1–Aug. 9, 2000– 2002, wet and dry pitfalls, elev.  80 m( R. Fisher, AMNH), 7?, 2/; Chino Hills, Telegraph Canyon,  33.914488N, 117.785558W,  Oct. 29–Nov. 2, 2001, dry pitfall, elev.  238 m( R. Fisher, AMNH), 1/; Chino Hills, ridge S Telegraph Canyon,  33.911968N, 117.789108W,  Sept. 7–17, 1999, dry pitfall, elev.  358 m( R. Fisher, AMNH), 1/; Crow Canyon,  33.602898N, 117.544278W,  Nov. 28– Dec. 8, 2000, dry pitfall, elev.  338 m( R. Fisher, AMNH), 1/; Laguna Beach,  June 24, 1931( W. Ivie, AMNH), 1?,  July 24, 1931(W. Ivie, AMNH), 1/,  Dec. 28, 1932, under rock (W. Ivie, AMNH), 1/,  Aug. 17, 2000( UCR), 1/; Limestone Canyon Regional Park,  33.736068N, 117.670488W,  June 18–22, 2001, dry pitfall, elev.  341 m( R. Fisher, AMNH), 1?; Limestone Canyon Regional Park, E The Sinks,  33.716218N, 117.658928W,  July 20–30, 1999, dry pitfall, elev.  453 m( R. Fisher, AMNH), 1?; Limestone Canyon Regional Park, ENE The Sinks,  33.722008N, 117.665798W,  June 1–Sept. 3, 1998–2002, pitfall, dry pitfalls, elev.  378 m( R. Fisher, AMNH), 2?, 2/; N Niguel Hill, NW Laguna Niguel,  33.536208N, 117.730158W,  Aug. 27–31, 2001, Jan. 28–Feb. 1, 2002, dry pitfalls, elev.  195 m( R. Fisher, AMNH), 2/; N Niguel Hill, NW Laguna Niguel,  33.538828N, 117.725448W,  July 10–Aug. 31, 2000–2002, dry pitfalls, elev.  132 m( R. Fish­ er, AMNH), 3?, 1/; San Joaquin Hills, E branch Laguna Canyon,  33.594688N, 117.775298W,  Sept. 9–13, 2002, dry pitfalls, elev.  175 m( R. Fisher, AMNH), 2/; Highway74, 2.5 mi E San Juan Fire Station, Hot Spring Canyonturnoff, Santa Ana Mountains,  May 11–22, 2002, elev.  1200 ft, under trash on oak duff beside highway,  60 mfrom stream ( W. Icenogle, WRI), 5?, 5/; Silverado Canyon, Santa Ana Mountains,  Nov. 20, 1955( R., A. Schick, AMNH), 1?; Starr Ranch,  33.630258N, 117.553088W,  Sept. 5–Oct. 14, 1998–2000, elev.  297 m, dry pitfalls( R. Fisher, AMNH), 4/; N Temple Hill, W Aliso Wood Canyon,  33.569908N, 117.756338W,  July 10–20, 2000, dry pitfall, elev.  215 m( R. Fisher, AMNH), 1?; N Temple Hill, W Aliso Wood Canyon,  33.583848N, 117.750308W,  May 13–17, 2002, dry pitfall, elev.  206 m( R. Fisher, AMNH), 1/; Weir Canyon,  33.809948N, 117.755648W,  June 15–25, 1999, dry pitfall, elev.  236 m( R. Fisher, AMNH), 1?; Weir Canyon,  33.815558N, 117.747138W,  Sept. 19–29, 2000, dry pitfall. elev.  232 m( R. Fihser, AMNH), 1/.  Riverside Co.: Bay Tree Springs, San Jacinto Mountains,  Nov. 13, 1976( M. Long, MET), 1/; Chino Canyon,  Apr. 20, 1960( J. Powell, UCB), 2?; 0.25 mi S Dark Canyon Campground, 2 road mi N junction Hwy243, San Jacinto Mountains,  June 4, 2002, under wood pile on ground near stream, conifer/oak forest ( W. Icenogle, WRI), 1/,  June 5, 2002, on side of boulder near stream, conifer/oak forest (W. Icenogle, WRI), 1/,  June 4, 2003, elev.  5765 ft, under wood piece beside recently molted skin, on ground near stream, conifer/oak forest (W. Icenogle, WRI), 1/; De Luz Road,  3 miS Murrieta,  May 10, 1977, under rock in ravine below live oaks, elev. ca.  2000 ft( W. Icenogle, WRI), 1/; Idyllwild, San Jacinto Mountains,  June 17, 1952(W. Gertsch, AMNH), 6?, 5/,  July 7, 1953( W. Gertsch, V. Roth, AMNH), 4?, 4/, 1996, elev. prob.  6000 ft( UCR), 1/; James Reserve, San Jacinto Mountains, 9 mi N Idyllwildon Highway243, 33848.49N, 116846.49W,  June 29, 2002, elev.  550 ft, coniferous forest, under granite exfoliations ( D., S. Ubick, M. Harvey, CDU), 3/; Lake Fulmor, James Reserve, San Jacinto Mountains, off Highway243, elev.  5380 ft( J. Kempf, UCR), 1/,  Aug. 1968( D. Bixler, CDB), 2?,  Sept. 1968(D. Bixler, CDB), 1/; Santa Margarita Ecological Reserve, W Temecula,  33.445238N, 117.164458W,  Sept. 7–17, 1999, dry pitfall, elev.  393 m( R. Fisher, AMNH), 1/; Santa Margarita Ecological Reserve, W Temecula,  33.455048N, 117.171238W,  May 11–21, 1998, dry pitfall, elev.  786 m( R. Fisher, AMNH), 1/; Santa Margarita Ecological Reserve, W Temecula,  33.456898N, 117.168458W,  Jan. 28–Feb. 7, 2000, dry pitfall, elev.  788 m( R. Fisher, AMNH), 1/; Strawberry Creek, 0.25 mi N Highway74, San Jacinto Mountains,  Aug. 12, 2001, elev.  3300 ft, under moist, rotting log beside stream ( W. Icenogle, WRI), 1/.  San Bernardino Co.: Crestline, San Bernardino Mountains, matured  Feb. 1997, elev.  5000 ft( J. Klotz, UCR), 1?; Crestline area, upper end of Waterman Canyon, Highway18, 0.5 mi NW junction with Highway138, San Bernardino Mountains,  Aug. 30, 2001, elev.  4200 ft, on wall, or dead and dried in spider webs on wall, of long pipe under highway ( W. Icenogle, WRI), 2?, 2/; East Fork Mountain Home Creek, near Highway38, San Bernardino Mountains,  July 6, 1999, under large rock in conifer forest near stream, elev.  5000 ft( W. Icenogle, WRI), 1/; Fontana,  July 16, 1942( E. Schlinger, AMNH), 1?, 1/; Forest Falls,  Aug. 10, 1985, elev.  7000 ft( D. Bixler, CDB), 2/; Forest Home, Hill Creek,  June 11, 1971, in cabin, elev.  6000 ft( D. Lowrie, MCZ), 1?; Glass Road, San Bernardino Mountains,  1 miN Highway  38, 348109290N, 1168549000W,  May 6, 2001(matured  Apr. 1, 2003), elev.  1820 m, in deciduous duff from  Quercus kelloggi(R. Vetter, UCR), 1/; off Highway38,  East Fork Mountain, Eside Home Creek, San Bernardino Mountains, 348079300N, 1168589540W,  Feb. 3, 2002(matured  Oct. 26–Nov. 2, 2002), elev.  5120 ft, from dead fern leaves, ferns growing on west­facing rock outcropping on steep canyon wall ( T. Prentice, UCR), 1?; Highway173 near Lake Arrowhead Village, 348159020N, 1178119590W,  July 10, 1999, elev.  5200 ft, along roadside and on hills at night ( B. Hebert, CBH), 3?, 2/; Lake Arrowhead,  May 6, 1936( S. Bishop, AMNH), 1/;  Lytle CreekCanyon,  Aug. 20, 1970( D. Bixler, CDB), 1/,  May 13, 1972, elev.  3000 ft( D. Bixler, CDB), 1?,  May 16, 1972, in house ( D. Bixler, CDB), 1?,  June 1, 1973( D. Bixler, CDB), 1?; Manker Flats Campground, San Gabriel Mountains,  May 14, 1992, elev.  6200 ft( B. Hebert, CBH), 1?, 1/; Mill CreekCanyon, Falls Picnic Area, San Bernardino Mountains,  May 15, 1969, molted  May 28, 1969, under rock in conifer forest, elev.  6000 ft( W. Icenogle, WRI), 1?,  June 18, 1969, elev.  6000 ft, under wood pieces and rocks (W. Icenogle, WRI), 1 penultimate /; Mountain Home Creek,  Aug. 13, 1959( W. Gertsch, V. Roth, AMNH), 9/; 209 Pine Oak, Fawnskin,  July 21, 1995, in lower level of house in tire chains ( V. Summer, UCR), 1?;  Silverwood Lake State Recreation Area, SEend Silverwood Lake,  34.277768N, 117.316948W,  Aug. 12–16, 2002, dry pitfall, elev.  1035 m( R. Fisher, AMNH), 1/; Skinner Creek, just E Highway38, 1.5 road mi NW junction with road to Forest Falls, San Bernardino Mountains,  June 10, 2001, elev.  4200 ft, under rocks and logs beside and above stream in bottom of heavily wooded canyon ( W. Icenogle, WRI), 2?, 1/; Skinner Ridge, between Skinner Creek and Mountain Home Creek, just SW Highway 38, 348069480N, 1168589530W,  Dec. 28, 1983–  Jan. 26, 1984, elev.  4900 ft, pitfall traps, canyon live oak forest, northern exposure ( M. Narog, UCR), 5/; Smiley Park, Running Springs,  July 12, 1995, in shower ( UCR), 1?; Smiley Park,  2 miS Running Springs on Highway330 off Fredalba Road,  July 10, 1995( J. Wambaugh, UCR), 1/; Twin Peaks, San Bernardino Mountains, ( K. Wormald, UCR), 1?, 1/, late May– early  June 1996, in house ( K. Wormald, UCR), 1?, 1/, mid­July 1996, male found in house, female outside (K. Wormald, UCR), 1?, 1/,  Aug.– Sept. 1996, elev.  5500 ft(K. Wormald, UCR), 2/, mid May–mid  June 1997( R. Vetter, UCR), 11?,  June 11, 1997( K. Wormald, UCR), 1/.  San Diego Co.: De Luz Road,  1 miN Fallbrook,  May 19, 1984, in drain under road ( G. Lowe, CGL), 1?,  Aug. 25, 1984(G. Lowe, CGL), 1/; Fry Creek, Mount Palomar, Aug.4, 1984 ( D. Boe, UCR), 1/; near Henshaw, Cleveland National Forest,  July 30, 1956( W. Gertsch, V. Roth, AMNH), 1?, 2/; W side Lake Henshaw, just E route S­6,  33.244388N, 116.764118W,  May 16, 2004, elev.  308 m(D. Palmer, P. Paquin, T. Prentice, C. Vink, AMNH), 1?; SW end Mendenhall Valley, Palomar Mountain,  June 3–July 2, 1997, pitfall, forest ( J. George, UCR), 3?,  July 2–Aug. 7, 1997, pitfall, elev.  6000 ft(J. George, UCR), 2?; Mount Palomar,  July 26, 1931( R. Chamberlin, AMNH), 1/,  June 30, 1956, elev.  3000–5000 ft( W. Gertsch, V. Roth, AMNH), 4?, 7/; Palomar Mountain,  July 4, 1967( B. Kaston, WRIK), 1?; Palomar Mountain, N Fry Creek, Wside route S­6, 33.34588N, 116.87948W,  May 16, 2004, elev.  1500 m(D. Palmer, P. Paquin, T. Prentice, C. Vink, AMNH), 3?, 1/; Palomar Mountain State Park,  July 13, 1953( W., J. Gertsch, AMNH), 1?, 3/; Palomar Mountain State Park, Doone Trail, 338209290N, 116854950W,  Feb. 20, 2003, in duff on creek bed by road, elev.  4639 ft( R. Vetter, UCR), 1? (matured mid­March 2003).  Santa Barbara Co.: Cold Spring Trail, Santa Barbara,  Aug. 14, 1959( W. Gertsch, V. Roth, AMNH), 31/;  5 miE Gaviota,  July 17, 1953( W., J. Gertsch, AMNH), 13?, 17/ (one with apparently teratological epigynum); Refugio Rd.,  23 miW Santa Barbara, Apr.6– May, 1978–1979 ( D. Boe, UCR), 4/; Santa Barbara,  Nov.–Dec. 1950( H. Shantz, AMNH), 2?,  July 17, 1953(H. Shantz, AMNH), 1?,  July 27, 1970(M. Cravens, CDFA), 1?;  10 miW Santa Barbara,  July 12, 1934(W. Ivie, AMNH), 1?, 2/; Sedgwick Ranch Reserve, 348439N, 1208019W,  May–June 11, 1997, malaise traps, elev.  1000–1800 ft( E., M. Schlinger, CAS), 1?, 1/,  June 24–July 21, 1997, malaise traps, elev.  800–1500 ft(E., M. Schlinger, CAS), 3?, Aug.27– Nov.1997, same, 3/,  May 28, 1997, malaise trap, elev.  1500–2500 ft( E. M. Schlinger, CAS), 1?; Upper Oso Campground, near Santa Ynez River,  Apr. 26–27, 1975, molted before  May 18, 1975, under rock ( F. Hovore, WRI), 1?.  Tulare Co.: Ash Mountain, Kaweah Power Station#  3, 40 miNE Visalia,  July 3, 1982( D. Burdick, CAS), 1?,  Mar. 12, 1983(D. Burdick, CAS), 1/,  June 19–21, 1983(D. Haines, CAS), 2?,  July 6–10, 1983, in flume (D. Burdick, CDU), 7?, 3/,  July 17, 1983(D. Burdick, CAS), 2/,  July 31, 1983( R. Haines, CDU), 1?, 3/,  June 8, 1985( R. Haines, CDFA), 2?, 1/,  June 7, 1986( R. Haines, CDFA), 1?,  June 9, 1986( R. Haines, CDFA), 2/,  June 28, 1986( R. Haines, CDFA), 1?,  July 12, 1986( R. Haines, CDFA), 1/,  Aug. 5–Sept. 3, 1995( D. Burdick, CAS), 3/,  July 24, 1996(D. Burdick, CAS), 1?; near Ash Mountainentrance, Sequoia National Park,  July 5, 1956, elev.  3700 ft( W. Gertsch, V. Roth, AMNH), 2?, 1/,  July 9, 1958( W. Gertsch, V. Roth, AMNH), 9?, 8/; NE Case MountainPeak,  36.417908N, 118.793298W,  July 14–18, 2003, dry pitfall, elev.  1691 m( R. Fisher, AMNH), 1?; NE Case Mountain Peak,  36.421838N, 118.793708W,  July 14–18, 2003, dry pitfall, elev.  1646 m( R. Fisher, AMNH), 1?; NE Case Mountain Peak, N Case Mountain Grove,  36.413178N, 118.791908W,  July 14–22, 2002–2003, dry pitfall, elev.  1810 m( R. Fisher, AMNH), 3?; NNE Case Mountain Peak, Eof E end of Salt Creek,  36.420358N, 118.793918W,  July 19–22, 2002, dry pitfall, elev.  1644 m( R. Fisher, AMNH), 1?,  36.421088N, 118.799768W,  July 19–22, 2002, dry pitfall, elev.  1819 m( R. Fisher, AMNH), 2?; NW Case Mountain Peak, S Salt Creek,  36.414428N, 118.813818W,  July 14–22, 2002–2003, dry pitfalls, elev.  1855 m( R. Fisher, AMNH), 3?; NW Case Mountain Peak, S Salt Creek,  36.417818N, 118.819308W,  July 14–18, 2003, dry pitfall, elev.  1709 m( R. Fisher, AMNH), 3?; SE Case MountainPeak, Case Mountain Grove,  36.403728N, 118.781908W,  July 14–18, 2003, dry pitfall, elev.  1678 m( R. Fisher, AMNH), 1?; SE Case MountainPeak, Case Mountain Grove,  36.405118N, 118.189828W,  July 19–22, 2002, dry pitfall, elev.  1805 m( R. Fisher, AMNH), 1/; SE Case Mountain Peak, Salt CreekRidge,  36.397408N, 118.775978W,  July 19–22, 2002, dry pitfall, elev.  1801 m( R. Fisher, AMNH), 1?,  36.398878N, 118.787788W,  July 19–22, 2002, dry pitfall, elev.  1848 m( R. Fisher, AMNH), 1?; Kaweah River,  5 miE Three Rivers,  July 17, 1952, elev.  1258 ft( W. Gertsch, AMNH), 1/; 5–10 mion Mineral King Roadfrom Three Rivers,  July 4, 1956, elev.  2000–4000 ft( W. Gertsch, V. Roth, AMNH), 5?, 5/; Moorehouse Creek, near Camp Nelson,  July 11, 1958( W. Gertsch, V. Roth, AMNH), 1?, 5/;  1 miW Camp Nelson,  July 11, 1958( W. Gertsch, V. Roth, AMNH), 2?; Roads End, Kern River,  July 3, 1956( W. Gertsch, V. Roth, AMNH), 3?, 2/; Sequoia National Park, Kaweah Cave, 36829926.10N, 118849917.40W,  Apr. 28, 2004, in cave entrance, elev.  490 m( D. Ubick, CDU), 1/ (matured  May 15, 2004);  Soda Creek, W Camp Nelson,  July 11, 1958( W. Gertsch, V. Roth, AMNH), 3/,  Sept. 14, 1959( W. Gertsch, V. Roth, AMNH), 1/.  Ventura Co.: Ojai,  Mar. 1978( D. Boe, UCR), 2/; Reyes Creek,  25 miSW Lakeof the Woods,  May 6, 1980, elev.  4000 ft( C. Griswold, UCB), 1/; Stecker Park, Highway150 N Santa Paula, 348259N, 119859W,  July 4, 1986, elev.  650 ft, from black widow web in restroom ( B. Hebert, CBH), 1?, 1/; Wheeler Springs,  July 2, 1958( W. Gertsch, V. Roth, AMNH, CAS), 17?, 13/; Yerba Buena Road,  2–3 miN Route1,  June 26, 1983, under rock by stream in canyon ( G. Lowe, CGL), 1?.  DISTRIBUTION: Southern California(map 1). NATURAL HISTORY: Collection records indicate that  Anachemmis soberoccurs in forests, primarily oak but also coniferous, at elevations from 70 to 2000 m. Specimens have Map 1. Western North America, showing records of  Anachemmis sober(open circles),  A. linsdalei(filled circles),  A. jungi(filled squares),  A. aalbui(open square), and  A. beattyi(triangles). been found beneath rocks, logs, and other ground litter, in and on leaf litter, on tree trunks, in a cave entrance, and in human structures, such as houses, culverts, and a flume. Several collections are from pitfall and malaise traps and of dead individuals in spider webs, including those of  Latrodectusand  Steatoda. The 700 specimensexamined indicate that both sexes occur year round, with females being more commonly collect­ ed, and a peak abundance from May to August (chart 1). That this species may be very common is documented by a single June day’s collection, at Tanbark Flats, Los Angeles County, of 95 malesand 130 females.    Anachemmis linsdalei, new species   Figures 1–20, 25–28; Map 1  TYPES: Male holotypeand female allotypetaken in house at Hastings Natural History State Reserve, Monterey Co., California(  Jan. 24–25, 1946; J. Linsdale), deposited in AMNH.  ETYMOLOGY: The specific name is a patronym in honor of the collector of the typesand many other specimens.  DIAGNOSIS: Males closely resemble those of  A. soberbut differ in having a sharply pointed tegular edge opposite the median apophysis (fig. 25); females also closely resemble those of  A. soberbut have a narrower epigynal septum and less anteriorly restricted lateral epigynal margins (fig. 27).  MALE: Total length 7.4. Coloration as in  A. sober. Leg spination: femora: II p1­2­1; IV r0­0­2; tibiae: I d0­0­1, r0­1­0; II d1­0­1, r0­1­0; III, IV d1­1­1; metatarsi: I, II p1­1­ 0, r1­1­0; III p1­2­2, r1­2­2; IV p1­2­2, r1­ 1­2. Tegulum widened opposite median apophysis, with pointed edge (fig. 25); dorsal prong of retrolateral tibial apophysis reduced to tiny, sharp point tip (fig. 26). FEMALE: Total length 9.7. Coloration as in  A. sober. Leg spination: femora I r1­1­1; tibiae: II r0–1­0; III, IV d1­1­1; metatarsi III, IV p1­2­2. Epigynum with median septum widened at about half its length, lateral margins longer than widened portion of septum (fig. 27); spermathecal bulbs rounded (fig. 28). OTHER MATERIAL EXAMINED: CALIFORNIA:  Alameda Co.:Berkeley, (AMNH), 4?, 2/.   Humboldt Co.: Humboldt RedwoodsState Park, Albee Creek Campground,   Sept. 10, 1992, elev.  360 ft(D. Ubick, J. Boutin, CDU), 1/.  Monterey Co.: Arroyo Seco, Indians Road, Los Padres National Forest, 368149N, 121829.59W,   May 7, 1995, dead yucca, elev.  1100 ft(D. Ubick, W. Savary, CDU), 1/,   July 8, 1995, at night (D. Ubick, S. Fend, W. Savary, CDU), 1?, 1/,   Aug. 24, 1996, oak­chaparral, at night along road (D. Ubick, CDU), 4/,   July 3, 1999, on road at night (S. Fend, J. Ledford, D. Ubick, CDU), 2?,   Apr. 15, 2000, oak woodland, under rocks, in litter (D., S. Ubick, CDU), 2?, 2/,   Apr. 28, 2001, oak woodland, under rocks and in litter (D., S. Ubick, CDU), 2/,   Apr. 27–Aug. 19, 2001, pitfall traps, elev.  1100 ft(D., S. Ubick, CDU), 94?, 12/, Aug. 19–Nov. 11, 2001, same, 2/, Feb. 22–May 19, 2002, same, 7?, 2?; May 19–Oct. 13, 2002, same, 100?, 24/, Oct. 13, 2002– Jan. 25, 2003, same, 1/, Jan. 25–June 11, 2003, same, 8?, 3/, June 11–Nov. 28, 2003, same, 63?, 18/, Nov. 28, 2003– June 18, 2004, same, 44?, 4/,   June 18–Oct. 24, 2004, same, 24?, 14/; Arroyo Seco Campground, Los Padres National Forest,   Apr. 26, 1980, under rock (D. Ubick, CDU), 1/,   May 13, 1994, at outhouse, elev.  950 ft(D. Ubick, J. Boutin, CDU), 1?,   May 13–15, 1994, oak forest, ground litter, elev.  900 ft(J. Boutin, CDU), 3?, 3/,   Apr. 24, 1999, oak riparian (D. Ubick, W. Savary, CDU), 1/; Arroyo Seco Campground, S The Lakes, Los Padres National Forest, 368149N, 1218299W,   May 13–15, 1994, under granite, leaf litter, elev.  900 ft(D. Ubick, J. Boutin, CDU), 3?, 6/,   May 6, 1995, under granite (D. Ubick, W. Savary, CDU), 1/,   Apr. 27–May 18, 2001–2002, at granite talus, under rocks (D., S. Ubick, DUC), 3?, Apr. 27–Aug. 19, 2001, same, pitfall traps, 3?, 1?, Aug. 19, 2001– May 18, 2002, same, pitfall traps, 3/, June 11–Nov. 28, 2003, same, 6?, 5/, Nov. 28, 2003– June 18, 2004, same, 9?,   June 18–Oct. 24, 2004. Same, 2?; Big Sur,   July 1937(W. Ivie, AMNH), 1/,   June 25, 1952( M. Cazier, W. Gertsch, R. Schrammel, AMNH), 1?, 4/,   July 18, 1953(W., J. Gertsch, AMNH), 1?, 6/,   Apr. 2, 1960( W. Gertsch, W. Ivie, R. Schrammel, AMNH), 4?, 6/;  14 miS Big Sur,   Dec. 22, 1953( V. Roth, AMNH), 1?; Carmel,   Nov. 12, 1953( AMNH), 1/,   Apr. 5–May 23, 1954( AMNH), 4?, 5/;  1 miN Carmel,   Dec. 21, 1953( V. Roth, AMNH), 1/; Carmel Valley,   Apr. 9, 1979(D. Boe, UCR), 1?, 1/,   May 1996(P. Stadille, CDU), 2?; Coast Ridge Trail, 0.8 mi SE Nacimiento Road,   Jun. 1, 1991, oak­pine forest, under bark litter, elev.  3000 ft(D. Ubick, CDU), 2?, 2/; Hastings Natural History State Reserve, 36822.89N, 121833.29W,   May 19–29, 1939(J. Linsdale, CAS), 2?,   May 30, 1940(J. Linsdale, CAS), 1/,   Feb. 22, 1941(J. Linsdale, CAS), 1/,   May 27, 1942(J. Linsdale, CAS), 1/,   July 18, 1944(J. Linsdale, CAS), 1?,   Jan. 24–25, 1946, house (J. Linsdale, AMNH), 2/,   Apr. 10, 1946, among sticks (J. Linsdale, CAS), 1/,   May 17, 1946, soil under house, passage in soil (J. Linsdale, AMNH), 1?, 1/,   Nov. 18, 1946,  Neotomanest (J. Linsdale, AMNH), 2?,   Dec. 8, 1946, house wall (J. Linsdale, AMNH), 2/, Dec. 9, Chart 1. Seasonal abundance of  Anachemmis soberbased upon the total specimens examined (728) from all localities. Y­axis 5 number of specimens; X­axis 5 time in months; solid lines 5 females; dashed lines 5 males.  1946,  Neotomanest (J. Linsdale, CAS), 1/,  June 1947(J. Linsdale, AMNH), 1?,  May 24, 1948, museum (J. Linsdale, AMNH), 1?,  May 17, 1950, under log (J. Linsdale, CAS), 1/,  June 24, 1950, headquarters, at night ( J. Linsdale, AMNH, 1/,  July 23, 1999, on trail at night ( S. Fend, J. Ledford, D. Ubick, CDU), 9?, 5/,  Aug. 2–4, 1999, in riparian woodland along creek ( D. Ubick, S. Fend, J. Ledford, CAS), 1/; Hastings Natural History State Reserve, Big Creek, 36.388998N, 121.548618W,  Jan. 9, 2004, elev.  578 m(D. Palm­ er, N. Stewart, AMNH), 1/; Highway G16 at G17,  Apr. 10, 1979(D. Boe, UCR), 1/; Landels­ Hill Big Creek Reservoir,  5 miN Lucia,  June 4– 6, 1982(C. Griswold, UCB), 1/; Monterey, Sept.6, 2002, in house (C. Edwards, UCR), 1/; Nacimiento Roadat Kirk Creek,  May 31, 1991, redwood forest, in duff, under rocks, elev.  1000 ft(D. Ubick, CDU), 1?, 1/; Nacimiento Roadat N Fork Mill Creek,  June 1, 1991, redwood forest, under bark and maple­tan oak litter, elev.  1500– 1600 ft(D. Ubick, CDU), 1?; Pacific Grove,  Aug. 15, 1931(W. Ivie, AMNH), 1/; Pfeifer Big Sur State Park,  27 miS Carmel,  May 3, 1964, elev. ca.  300 ft(P. Craig, CAS), 2/,  Aug. 19, 1987, riparian (D. Gross, CAS), 1/; Redwood Canyon, 4 miS Gorda,  Apr. 2, 1960(W. Gertsch, W. Ivie, R. Schrammel, AMNH), 2/;  4 miW Salinas,  Sept. 25, 1961( W. Gertsch, W. Ivie, AMNH), 1/; Tassajara Spring Buddhistretreat,  Sept. 13, 1978( R. Doty, AMNH), 1/; UCNRS Big Creek Reserve, nr. Whale Point Cabin, 3684.349N, 121835.819W, elev. ca  200 m,  May 24– 27, 2002(S. Lew, UCB), 2/,  Oct. 24, 2002(S. Lew, UCB), 1? (molted in captivity),  May 12, 2003(S. Lew, UCB), 1/.  San Benito Co.: Pinnacles National Monument,  July 3, 1958( W. Gertsch, V. Roth, AMNH), 1?, 2/; Pinnacles National Monument, Balconies Caves area, June8, Chart 2. Seasonal abundance of  Anachemmis linsdaleibased upon the total of hand­collected specimens examined (182) from all localities. Yaxis 5 number of specimens; X­axis 5 time in months; solid lines 5 females; dashed lines 5 males. 1991, oak­riparian, under granite (D. Ubick, CDU), 1/; Pinnacles National Monument, Bear Gulch Trail at Chalone Creek, May 8, 1992, oakbuckeye grove, under rocks in talus slope (T. Briggs, D. Ubick, W. Rauscher, CDU), 1/. San Luis Obispo Co.:Cambria, Nov. 16, 1937(O. Bryant, AMNH), 1?, 3/, June 23, 1952(M. Cazier, W. Gertsch, R. Schrammel, AMNH), 13?, 17/, Feb. 7, 1967, in pine duff (V. Roth, AMNH), 1?; Old Morro Road, 1 miE Hwy 41, Apr. 3, 1985, oak­grassland, under serpentine (W. Rauscher, CDU), 1/, Apr. 3, 1985, matured in May (D. Ubick, CDU), 1?, 1/; 6 miN Paso Robles, Sept. 26, 1961(W. Gertsch, V. Roth, AMNH), 1/; 26 miW Paso Robles, May 12, 1961(R. Schuster, F. Raney, AMNH), 1/; Reservoir Canyon, San Luis Obispo, Aug. 15, 1959(W. Gertsch, V. Roth, AMNH), 2/; San Luis Obispo, Aug. 8, 1931(R. Chamberlin, AMNH), 1/. Santa Cruz Co.:Santa Cruz, Apr.7, 1979 (D. Boe, UCR), 2?, 1/.  DISTRIBUTION: Northern and central California(map 1). NATURAL HISTORY: Collection records of  Anachemmis linsdaleiare from elevations of 100 to 500 mand predominantly from various typesof oak woodlands, but also from pine and redwood forests. This species occurs on ground habitats and has been collected under rocks, logs, in and on litter, in  Neotomanests, in houses, and on roads at night. Hand collected specimens (about 200) show a similar abundance to  A. sober, being collected year round with females being slightly more common and a peak of activity from April to June (chart 2). The 450 specimensobtained from ethylene glycol pitfall traps (10 traps) at Arroyo Seco, Monterey County, indicate a much higher proportion of males (chart 3). During periods of peak activity, this species was, by far, the most abundant spider in the pitfall traps.     Anachemmis jungi, new species   Figures 29–32; Map 1  TYPES: Male holotypeand female allotypefrom Titus Canyon Cave,  Death ValleyNational Monument, Inyo Co., California(  Jan. 22, 1972; A. Jung), deposited in AMNH.  ETYMOLOGY: The specific name is a patronym in honor of the collector of the types.  DIAGNOSIS: Males of this long­legged, cavernicole species resemble those of  A. aalbuiin having a narrow, erect embolus (fig. 29), but have a much narrower ventral prong of the retrolateral tibial apophysis (fig. 30); females also resemble those of  A. aalbuiin having very short anterolateral epigynal margins, but have the epigynal septum widened at the level of the anterolateral epigynal margins (fig. 31).  MALE: Total length 8.9. Color paler than in  A. sober, with carapace brownish orange, legs with femora light yellow, grading to brownish orange on more distal segments, abdomen pale white. Leg spination: femora: I p0­2­1; II p2­2­1; IV r0­1­2; tibiae: I, II d1­ 0­1, v6­4­2, r1­0­0; IV r1­0­1; metatarsi: I, II p1­1­0, r1­1­0; III p1­2­2; IV v2­2­2. Tegulum not widened opposite median apophysis, embolus relatively narrow, erect (fig. 29); dorsal prong of retrolateral tibial apophysis relatively large for genus, ventral prong relatively narrow (fig. 30). FEMALE: Total length 13.6. Coloration as in male. Leg spination: femora: I p0­3­1; II p2­2­1; III r2­1­1; IV p2­1­2, r0­1­2; tibiae: I, II v4­6­2, r0­1­0; metatarsi: III p1­2­2; IV r1­1­2. Anterolateral epigynal margins and atrium relatively short, septum widened at that level (fig. 31), median epigynal ducts massive (fig. 32). terolateral epigynal margins, but have the epigynal septum widest anteriorly, rather than at the level of the anterolateral epigynal margins (fig. 35). MALE: Total length 9.2. Coloration as in  A. jungi. Leg spination (both legs I missing): femora: II p2­2­1; IV r0­0­2; tibiae II d1­0­ 1, v4­6­2, r0­1­0; metatarsi: II p2­0­0, r1­0­ 0; III p1­2­2, r1­2­2; IV r1­1­2. Tegulum not widened opposite median apophysis, embolus relatively narrow, erect (fig. 33); dorsal prong of retrolateral tibial apophysis relatively large for genus, ventral prong relatively wide, especially at base (fig. 34). FEMALE: Total length 12.3. Coloration as in  A. jungi. Leg spination: femora: I p0­3­1; II p2­2­1; III r2­1­1; IV r0­0­2; tibiae I, II v4­6­2, r0­1­0; metatarsi: I, II p2­0­0, r2­0­ 0; III p1­2­2; IV p1­2­2, r1­1­2. Anterolateral epigynal margins and atrium relatively short, septum widened anteriorly (fig. 35), median epigynal ducts massive (fig. 36).  OTHER MATERIAL EXAMINED: CALIFORNIA:  San Bernardino Co.: Medicine Cave, Providence  OTHER MATERIAL EXAMINED: CALIFORNIA:  Inyo Co.: Defense Cave, Stone Canyon, Argus Range,  Sept. 7, 1988, elev.  4000 ft( R. Aalbu, CAS), 1/; Titus Canyon Cave,  Death ValleyNational Monument,  Jan. 22, 1972( A. Jung, AMNH), 1?.  DISTRIBUTION: Known only from caves in   Inyo Co., California(map 1).    Anachemmis aalbui, new species   Figures 33–36; Map 1   TYPE: Male holotypetaken in pitfalltrap at an elevation of  4300 ftat the Park Headquarters, Mitchell Caverns State Park, San Bernardino Co., California(  Mar. 17–June 16, 1979; R. Aalbu), deposited in CAS.  ETYMOLOGY: The specific name is a patronym in honor of the collector of the types.  DIAGNOSIS: Males of this long­legged species resemble those of  A. jungiin having a narrow, erect embolus (fig. 33), but have a wider ventral prong of the retrolateral tibial apophysis (fig. 34); females also resemble those of  A. jungiin having very short an­ Chart 3. Seasonal abundance of  Anachemmis linsdaleibased upon the specimens collected (450) from 10 pitfall traps maintained at Arroyo Seco, Monterey County, California. Y­axis 5 number of specimens; X­axis 5 time; solid lines 5 females; dashed lines 5 males.   Mountains State Recreation Area,  June 20–Sept. 27, 1988, pitfall( R. Aalbu, CAS), 6/.   DISTRIBUTION: Known only from San Bernardino Co., California(map 1).    Anachemmis beattyi, new species   Figures 37–40; Map 1   TYPE: Male holotypefrom  Peppersauce CaveCanyon(north side of Santa Catalina Mountains), Pinal Co., Arizona(  Apr. 21, 1961; J. Beatty), deposited in AMNH.  ETYMOLOGY: The specific name is a patronym in honor of the collector of the types.  DIAGNOSIS: Males of this distinctive species can be recognized by the distally long and narrowed embolus (fig. 37), females by the anteriorly very wide epigynal septum bearing deep, paramedian invaginations (fig. 39).  MALE: Total length 8.4. Coloration as in  A. jungi. Leg spination: femora: I p0­3­1, r2­ 1­2; II p2­1­1; IV r0­1­2; tibiae: I d1­0­1, v4­ 6­2, r1­1­0; II d1­0­1, v4­6­2, r1­0­0; metatarsi: I r1­0­0; II p0­1­0, r1­1­0. Tegulum not widened opposite median apophysis, embolus narrowed distally (fig. 37); dorsal prong of retrolateral tibial apophysis reduced to tiny point, ventral prong relatively narrow (fig. 38). FEMALE: Total length 8.8. Coloration as in  A. jungi. Leg spination: femora: I p0­2­1, r1­ 1­1; IV p2­0­1, r0­0­2; patellae: III p0­0­0, r0­0­0; IV p0­0­0; tibiae I, II v4­6­2, r0­1­0; metatarsi III p1­2­2. Anterolateral epigynal margins and atrium relatively short, septum much widened at that level, bearing deep paramedian invaginations (fig. 39), median epigynal ducts massive (fig. 40).   OTHER MATERIAL EXAMINED: ARIZONA:  Coconino Co.: Manzanita Camp,  Oak CreekCanyon,  July 25, 1952( M. Cazier, AMNH), 2/.  Graham Co.: Lower Canyon, Mount Graham,  July 14, 1956( W. Gertsch, V. Roth, AMNH), 1/. MEXICO:  SONORA: Cajon Bonito, 318189N, 1088559W,  Sept. 11, 1973( V. Roth, CAS), 3/.  DISTRIBUTION: Known only from Arizona and Sonora(map 1). 1985-06-28 1993-06-11 1985-06-28 R, CDU, AMNH Robertson & S. Stockwell & S. Taber & O. Francke Kern Co. 2134 Ninemile Canyon Bakersfield 11 12 1 1982-06-27 AMNH N La & G. Lowe Canada Los Angeles Co. Highway Angeles National Forest 11 12 1 1953-05-09 AMNH Heard Island and McDonald Islands Beverly Glen Canyon Santa Monica Mountains 11 12 1 1982-01-27 CBH B. Hebert Heard Island and McDonald Islands Angeles National Forest 1798 Highway Grand View Canyon 11 12 1 1943-06-12 AMNH K. Cowles Heard Island and McDonald Islands 1280 Big Rock Creek Valyermo 11 12 1 1969-04-28 MET Charlton-Chilao P. C. & M. Thompson Heard Island and McDonald Islands Angeles National Forest 11 12 1 1975-05-13 MET M. Thompson Heard Island and McDonald Islands Charmlee County Charmlee County 11 12 1 1997-05-03 UCR Kempf Heard Island and McDonald Islands 396 11 12 1 1913-04 1965-08-05 1913-04 R, MCZ, MET Claremont & Chamberlin & M. Thompson Heard Island and McDonald Islands Chamberlin Eaton Canyon Park 11 12 2 paratype 1965-08-16 MET Thompson Heard Island and McDonald Islands 11 12 1 paratype 1967-08-04 MET Thompson Heard Island and McDonald Islands 11 12 1 paratype [462,948,1107,1741] 1968-01-18 MET Thompson Heard Island and McDonald Islands 11 12 1 paratype 1968-02-19 MET Thompson Heard Island and McDonald Islands 11 12 1 paratype 1968-05-05 MET Thompson Heard Island and McDonald Islands 11 12 1 paratype 1968-05-07 MET Thompson Heard Island and McDonald Islands 11 12 1 paratype 1950-11-25 AMNH E. Schlinger Heard Island and McDonald Islands Glendale 11 12 1 paratype 1958-05-25 AMNH Schlinger Heard Island and McDonald Islands 11 12 1 paratype 2002-05-10 UCR Schwellenbach Heard Island and McDonald Islands 11 12 1 paratype 1955-03 1982-06-27 1955-03 R, AMNH, CGL G. Lowe Heard Island and McDonald Islands Highway 1798 Gold Canyon Big Pines Highway 11 12 1 paratype 1968-11-11 1968-11-15 1968-11-11 MET D. Marqua Heard Island and McDonald Islands Henninger Flats Angeles National Forest 11 12 1 paratype 1969-05-29 1994-10 1969-05-29 R, UCR, MET Vetter & M. Thompson Heard Island and McDonald Islands Hollywood Lady Bug Canyon 11 12 1 paratype 1981-05 CDFA G. Bakker Heard Island and McDonald Islands Los Angeles 11 12 1 paratype UCR Malibu & M. Balducci Heard Island and McDonald Islands 11 12 1 paratype 1982-12-11 CGL G. Lowe Heard Island and McDonald Islands Malibu Lagoon Serra Road 11 12 1 paratype 1987-04-15 CGL G. Lowe Heard Island and McDonald Islands 1859 Marker Flats Mount Baldy Road 11 12 1 paratype 1983-12-04 CGL G. Lowe Heard Island and McDonald Islands 12 13 Millard Canyon Altadena 11 12 1 paratype 1984-07-05 CGL Lowe Heard Island and McDonald Islands 12 13 1 paratype 2002-01-18 2002-02-28 2002-01-18 WRI W. Icenogle & J. Bond Heard Island and McDonald Islands Altadena 488 Millard Canyon San Gabriel Mountains 12 13 1 paratype 1982-11-13 CGL G. Lowe Heard Island and McDonald Islands Route Old Topanga Canyon Road Topanga Canyon Boulevard 12 13 1 paratype 1989-09-09 CBH Hebert Heard Island and McDonald Islands 12 13 1 paratype 1998-07-16 UCR A. Moore Heard Island and McDonald Islands Palmer Canyon Road Claremont 12 13 1 paratype 1966-05-02 UCR C. Goodpasture Heard Island and McDonald Islands Pasadena 12 13 1 paratype 1972-07-26 WRI F. Hovore Heard Island and McDonald Islands 472 Placerita Canyon Park 12 13 1 paratype 1976-05-27 MET Sullivan Heard Island and McDonald Islands 12 13 1 paratype S Roland Heights Heard Island and McDonald Islands Puente Hills 12 13 1 paratype 1956-07-01 2002-11-08 1956-07-01 R, AMNH, V, WRI Fisher & San Antonio Canyon & Claremont & W. Gertsch & Roth & S Lower San Antonio Forest Station & W. Icenogle Heard Island and McDonald Islands San Gabriel Mountains 294 33.969017 Mount Baldy Road 1 -117.92689 Lower San Antonio Forest Station 12 13 1 paratype 2001-07-09 WRI E Sunset Peak & W. Icenogle Heard Island and McDonald Islands 1158 San Antonio Canyon Mount Baldy Road 12 13 1 paratype 1956-09-02 2002-05-31 1956-09-02 V, AMNH, UCR NE Big Dalton Reservoir & M. Oxford Heard Island and McDonald Islands Volfe Canyon 585 San Dimas Canyon Big Dalton Canyon Road 12 13 1 paratype 1989-11-12 CBH B. Hebert Heard Island and McDonald Islands Santa San Gabriel Mountains 12 13 1 paratype 1991-04-23 1991-05-24 1991-04-23 CBH Hebert Heard Island and McDonald Islands 12 13 1 paratype 1941-03-20 AMNH Ivie Heard Island and McDonald Islands 15 mi W Santa Monica 12 13 1 paratype 1969-04-26 MET M. Thompson Heard Island and McDonald Islands San Gabriel Mountains Shortcut Highway 12 13 1 paratype 1977-06 1977-07-31 1977-06 LACM G. Gorelick Heard Island and McDonald Islands Sierra Madre 12 13 1 paratype 2001-07-05 WRI W. Icenogle Heard Island and McDonald Islands San Gabriel Mountains 1097 Spruce Canyon Mount Baldy Road 12 13 1 paratype 1951-05-19 1985-06-27 1951-05-19 T, AMNH Tice & L. Sorkin Heard Island and McDonald Islands Angeles National Forest 1158 Sunland Highway 12 13 1 paratype 2002-04-15 2002-04-29 2002-04-15 WRI W. Icenogle Heard Island and McDonald Islands San Gabriel Mountains 1006 Switzer Picnic Area Highway 12 13 1 paratype 1952-06-20 AMNH, JCC Cazier & W. Gertsch & R. Schrammel Heard Island and McDonald Islands Tanbark Flats San Gabriel Mountains 12 13 1 paratype [700,1105,352,373] 1957-03-30 AMNH Heard Island and McDonald Islands 12 13 1 paratype 1955-03-20 1995-04-10 1955-03-20 R, AMNH, CBH Apr. & Schick & B. Olodort Heard Island and McDonald Islands Santa Monica Mountains Tapia Park Pacific Palisades 12 13 1 paratype Heard Island and McDonald Islands Topanga Highway 12 13 1 paratype 2001-08-27 2004-03-13 2001-08-27 R, WRI D. Palmer & J. Starrett & W. Icenogle Heard Island and McDonald Islands San Gabriel Mountains 470 34.094837 Keith 1 -118.6015 Shortcut Ranger Station 12 13 1 paratype Aliso Wood Canyon & E Laguna Beach Heard Island and McDonald Islands Orange Co. Orange Co. 12 13 1 paratype 2001-06-10 2001-08-31 2001-06-10 R, AMNH Fisher & N Aliso Wood Canyon, E Heard Island and McDonald Islands 71 33.558456 Laguna Beach 1 -117.74961 12 13 1 paratype 2000-06-01 2000-08-09 2000-06-01 R, AMNH Fisher Heard Island and McDonald Islands 80 33.567577 Chino Hills 1 -117.74826 Telegraph Canyon 12 13 1 paratype 2001-10-29 2001-11-02 2001-10-29 R, AMNH Fisher Heard Island and McDonald Islands 238 33.91449 Chino Hills 1 -117.78556 Telegraph Canyon 12 13 1 paratype 1999-09-07 1999-09-17 1999-09-07 R, AMNH Fisher Heard Island and McDonald Islands 358 33.91197 Crow Canyon 1 -117.78911 12 13 1 paratype 1931-06-24 2000-12-08 1931-06-24 R, AMNH W. Ivie Heard Island and McDonald Islands 338 33.602898 Fisher 1 -117.54428 Laguna Beach 12 13 1 paratype 1931-07-24 AMNH Ivie Heard Island and McDonald Islands 12 13 1 paratype 1932-12-28 AMNH Ivie Heard Island and McDonald Islands 12 13 1 paratype [806,1098,1208,1229] 2000-08-17 UCR Heard Island and McDonald Islands 12 13 1 paratype 2001-06-18 2001-06-22 2001-06-18 R, AMNH Fisher & E The Sinks Heard Island and McDonald Islands 341 33.73607 Limestone Canyon Regional Park 1 -117.67049 12 13 1 paratype 1999-07-20 1999-07-30 1999-07-20 R, AMNH Fisher Heard Island and McDonald Islands 453 33.716217 Limestone Canyon Regional Park 1 -117.65893 The Sinks 12 13 1 paratype 1998-06-01 1998-09-03 1998-06-01 R, AMNH Fisher & N Niguel Hill, NW Heard Island and McDonald Islands 378 33.722008 Laguna Niguel 1 -117.665794 12 13 1 paratype Aug. 27 - 31, 2001, Jan. 28 - Feb. 1, 2002 R, AMNH Fisher & N Niguel Hill, NW Heard Island and McDonald Islands 195 33.53621 Laguna Niguel 1 -117.730156 12 13 1 paratype 2000-07-10 2000-08-31 2000-07-10 R, AMNH Heard Island and McDonald Islands 132 33.53883 San Joaquin Hills 1 -117.72545 Laguna Canyon 12 13 1 paratype 2002-05-11 2002-09-13 2002-05-11 R, AMNH, WRI Fisher & W. Icenogle Heard Island and McDonald Islands Santa Ana Mountains 175 13 14 33.59469 Highway 1 -117.7753 Hot Spring Canyon 12 13 1 paratype 1955-11-20 R, AMNH A. Schick Heard Island and McDonald Islands Starr Ranch Silverado Canyon Santa Ana Mountains 13 14 1 paratype 1998-09-05 1998-10-14 1998-09-05 R, AMNH Fisher & N Temple Hill & W Aliso Wood Canyon Heard Island and McDonald Islands 297 33.630257 1 -117.553085 13 14 1 paratype 2000-07-10 2000-07-20 2000-07-10 R, AMNH Fisher & N Temple Hill & W Aliso Wood Canyon Heard Island and McDonald Islands 215 33.56991 1 -117.75634 13 14 1 paratype 2002-05-13 2002-05-17 2002-05-13 R, AMNH Fisher Heard Island and McDonald Islands 206 33.583847 Weir Canyon 1 -117.750305 13 14 1 paratype 1999-06-15 1999-06-25 1999-06-15 R, AMNH Fisher Heard Island and McDonald Islands 236 33.809948 Weir Canyon 1 -117.755646 13 14 1 paratype 1976-11-13 2000-09-29 1976-11-13 R, AMNH, MET Fihser & M. Long Heard Island and McDonald Islands Riverside Co. 232 33.81556 Bay Tree Springs 1 -117.74714 San Jacinto Mountains 13 14 1 paratype 1960-04-20 UCB J. Powell Heard Island and McDonald Islands Chino Canyon 13 14 1 paratype 2002-06-04 WRI S Dark Canyon Campground & W. Icenogle Heard Island and McDonald Islands Hwy San Jacinto Mountains 13 14 1 paratype 2002-06-05 WRI Icenogle Heard Island and McDonald Islands 13 14 1 paratype 2003-06-04 WRI Icenogle Heard Island and McDonald Islands 1757 13 14 1 paratype 1977-05-10 WRI W. Icenogle Heard Island and McDonald Islands 610 De Luz Road 13 14 1 paratype 1952-06-17 AMNH Gertsch Heard Island and McDonald Islands 13 14 1 paratype 1953-07-07 V, AMNH, UCR W. Gertsch & Roth Heard Island and McDonald Islands 1829 13 14 1 paratype 2002-06-29 CDU D., S. Ubick & M. Harvey Heard Island and McDonald Islands Highway 168 James Reserve San Jacinto Mountains 13 14 1 paratype UCR J. Kempf Heard Island and McDonald Islands Highway 1640 Lake Fulmor San Jacinto Mountains 13 14 1 paratype 1968-08 1968-09 1968-08 D. Bixler & Bixler & W Temecula Heard Island and McDonald Islands Santa Margarita Ecological Reserve 13 14 1 paratype 1999-09-07 1999-09-17 1999-09-07 R, AMNH Fisher & W Temecula Heard Island and McDonald Islands 393 33.445236 Santa Margarita Ecological Reserve 1 -117.16446 13 14 1 paratype 1998-05-11 1998-05-21 1998-05-11 R, AMNH Fisher & W Temecula Heard Island and McDonald Islands 786 33.455048 Santa Margarita Ecological Reserve 1 -117.17124 13 14 1 paratype 2000-01-28 2001-08-12 2000-01-28 R, AMNH, WRI Fisher & W. Icenogle Heard Island and McDonald Islands San Jacinto Mountains 788 33.456898 Strawberry Creek 1 -117.16846 Highway 13 14 1 paratype 1997-02 UCR J. Klotz Heard Island and McDonald Islands San Bernardino Co. 1524 Crestline San Bernardino Mountains 13 14 1 paratype 2001-08-30 WRI W. Icenogle Heard Island and McDonald Islands San Bernardino Mountains 1280 Crestline area Highway 13 14 1 paratype 1999-07-06 WRI W. Icenogle Heard Island and McDonald Islands San Bernardino Mountains 1524 East Fork Mountain Home Creek Highway 13 14 1 paratype 1942-07-16 AMNH E. Schlinger Heard Island and McDonald Islands Fontana 13 14 1 paratype 1971-06-11 1985-08-10 1971-06-11 MCZ D. Bixler & D. Lowrie Heard Island and McDonald Islands Hill Creek 2134 Forest Falls Forest Home 13 14 1 paratype Heard Island and McDonald Islands Glass Road San Bernardino Mountains 13 14 1 paratype 1999-07-10 2003-04-01 1999-07-10 T, UCR, CBH East Fork Mountain, E & Prentice & B. Hebert Heard Island and McDonald Islands Lake Arrowhead Village 1820 38.34811 Highway Highway 13 14 1 paratype 1936-05-06 AMNH S. Bishop Heard Island and McDonald Islands Lake Arrowhead 13 14 1 paratype 1970-08-20 1992-05-14 1970-08-20 CBH D. Bixler & Manker Flats Campground & B. Hebert Heard Island and McDonald Islands San Gabriel Mountains 914 Lytle Creek Canyon Lytle Creek 13 14 1 paratype 1969-05-15 1969-05-28 1969-05-15 WRI W. Icenogle Heard Island and McDonald Islands San Bernardino Mountains 1829 Mill Creek Falls Picnic Area 13 14 1 paratype 1969-06-18 WRI Icenogle Heard Island and McDonald Islands 1829 13 14 1 paratype 1959-08-13 1995-07-21 1959-08-13 V, AMNH, UCR W. Gertsch & Summer & Silverwood Lake State Recreation Area, SE Heard Island and McDonald Islands Silverwood Lake Mountain Home Creek Silverwood Lake State Recreation Area 13 14 1 paratype 2001-06-10 2002-08-16 2001-06-10 R, AMNH, WRI Fisher & W. Icenogle Heard Island and McDonald Islands San Bernardino Mountains 1035 34.277767 Skinner Creek 1 -117.31695 Forest Falls 13 14 1 paratype 1983-12-28 1984-01-26 1983-12-28 UCR M. Narog Heard Island and McDonald Islands 1494 14 15 38.34807 Skinner Ridge Highway 13 14 1 paratype 1995-07-12 UCR Heard Island and McDonald Islands Smiley Park Running Springs 14 15 1 paratype 1995-07-10 UCR J. Wambaugh Heard Island and McDonald Islands Smiley Park Fredalba Road 14 15 1 paratype UCR Twin Peaks & K. Wormald Heard Island and McDonald Islands San Bernardino Mountains 14 15 1 paratype 1996-06 UCR K. Wormald Heard Island and McDonald Islands May 14 15 1 paratype UCR Wormald Heard Island and McDonald Islands 14 15 1 paratype 1996-08 1996-09-31 1996-08 UCR Wormald Heard Island and McDonald Islands 1676 14 15 1 paratype 1997-06 1997-06-11 1997-06 R, UCR Vetter & K. Wormald Heard Island and McDonald Islands May 14 15 1 paratype 1984-05-19 CGL De Luz Road & G. Lowe Heard Island and McDonald Islands San Diego Co. 1 mi N Fallbrook 14 15 1 paratype [124,544,592,613] 1984-08-25 CGL Lowe Heard Island and McDonald Islands 14 15 1 paratype 1984-08-04 UCR D. Boe Heard Island and McDonald Islands Fry Creek Mount 14 15 1 paratype 1956-07-30 V, AMNH W. Gertsch & Roth Heard Island and McDonald Islands Lake Henshaw Henshaw Cleveland National Forest 14 15 1 paratype 2004-05-16 AMNH Palmer & P. Paquin & T. Prentice & C. Vink Heard Island and McDonald Islands 308 33.24439 1 -116.764114 14 15 1 paratype 1997-06-03 1997-07-02 1997-06-03 UCR J. George Heard Island and McDonald Islands Mendenhall Valley 14 15 1 paratype 1997-07-02 1997-08-07 1997-07-02 UCR George Heard Island and McDonald Islands 1829 14 15 1 paratype 1931-07-26 1967-07-04 1931-07-26 R, AMNH, V, WRIK Chamberlin & W. Gertsch & B. Kaston Heard Island and McDonald Islands 1219 Mount Palomar Roth 14 15 1 paratype 2004-05-16 AMNH Palmer & P. Paquin & T. Prentice & C. Vink Heard Island and McDonald Islands 1500 33.34588 1 -116.87948 14 15 1 paratype 1953-07-13 AMNH W., J. Gertsch Heard Island and McDonald Islands State Park 14 15 1 paratype 1953-07-17 2003-02-20 1953-07-17 R, UCR, V, AMNH Vetter & Cold Spring Trail & W. Gertsch & Roth & W., J. Gertsch Heard Island and McDonald Islands Santa Barbara Co. 1414 State Park Santa Barbara 14 15 1 paratype UCR May & D. Boe Heard Island and McDonald Islands Refugio Rd. Apr. 14 15 1 paratype 1950-11 1950-12-31 1950-11 AMNH H. Shantz Heard Island and McDonald Islands Santa Barbara 14 15 1 paratype 1953-07-17 AMNH Shantz Heard Island and McDonald Islands 14 15 1 paratype 1970-07-27 CDFA Cravens Heard Island and McDonald Islands 14 15 1 paratype 1934-07-12 AMNH Ivie Heard Island and McDonald Islands 10 mi W Santa Barbara 14 15 1 paratype 1997-05-11 1997-06-11 1997-05-11 CAS E., M. Schlinger Heard Island and McDonald Islands 427 Sedgwick Ranch Reserve 14 15 1 paratype 1997-06-24 1997-07-21 1997-06-24 CAS Schlinger Heard Island and McDonald Islands 351 14 15 1 paratype 1997-05-28 CAS E. M. Schlinger Heard Island and McDonald Islands 610 Aug. Nov. 14 15 1 paratype 1975-04-26 1975-05-18 1975-04-26 WRI F. Hovore Heard Island and McDonald Islands Upper Oso Campground Santa Ynez River 14 15 1 paratype 1982-07-03 CAS D. Burdick Heard Island and McDonald Islands Ash Mountain Kaweah Power Station 14 15 1 paratype 1983-03-12 CAS Burdick Heard Island and McDonald Islands 14 15 1 paratype 1983-06-19 1983-06-21 1983-06-19 CAS Haines Heard Island and McDonald Islands 14 15 1 paratype 1983-07-06 1983-07-10 1983-07-06 CDU Burdick Heard Island and McDonald Islands 14 15 1 paratype [778,1225,352,373] 1983-07-17 CAS Burdick Heard Island and McDonald Islands 14 15 1 paratype 1983-07-31 1995-09-03 1983-07-31 R, CDU, CDFA, CAS D. Burdick Heard Island and McDonald Islands 14 15 1 paratype 1996-07-24 CAS Burdick Heard Island and McDonald Islands 14 15 1 paratype 1956-07-05 1958-07-09 1956-07-05 V, AMNH W. Gertsch & Roth & NE Case Mountain Peak Heard Island and McDonald Islands Mountain Peak 1128 Ash Mountain Case Mountain 14 15 1 paratype 2003-07-14 2003-07-18 2003-07-14 R, AMNH Fisher & NE Case Mountain Peak Heard Island and McDonald Islands 1691 36.417908 1 -118.7933 14 15 1 paratype 2003-07-14 2003-07-18 2003-07-14 R, AMNH Fisher & NE Case Mountain Peak Heard Island and McDonald Islands 1646 36.421837 Mountain Grove 1 -118.79371 14 15 1 paratype 2002-07-14 2002-07-22 2002-07-14 R, AMNH Fisher & Case Mountain Peak, E Heard Island and McDonald Islands 1810 36.413177 Salt Creek 1 -118.79191 14 15 1 paratype 2002-07-19 2002-07-22 2002-07-19 R, AMNH Fisher Heard Island and McDonald Islands 1644 36.420357 1 -118.793915 14 15 1 paratype 2002-07-19 2002-07-22 2002-07-19 R, AMNH Fisher & NW Case Mountain Peak, S Heard Island and McDonald Islands 1819 36.42109 Salt Creek 1 -118.79977 14 15 1 paratype 2002-07-14 2002-07-22 2002-07-14 R, AMNH Fisher & NW Case Mountain Peak, S Heard Island and McDonald Islands 1855 36.41443 Salt Creek 1 -118.81382 14 15 1 paratype 2003-07-14 2003-07-18 2003-07-14 R, AMNH Fisher Heard Island and McDonald Islands 1709 36.417816 Case Mountain 1 -118.819305 Mountain Grove 14 15 1 paratype 2003-07-14 2003-07-18 2003-07-14 R, AMNH Fisher Heard Island and McDonald Islands 1678 36.40373 Case Mountain 1 -118.781906 Mountain Grove 14 15 1 paratype 2002-07-19 2002-07-22 2002-07-19 R, AMNH Fisher & SE Case Mountain Peak Heard Island and McDonald Islands 1805 36.405117 Salt Creek 1 -118.18983 14 15 1 paratype 2002-07-19 2002-07-22 2002-07-19 R, AMNH Fisher Heard Island and McDonald Islands 1801 36.397408 1 -118.77598 14 15 1 paratype 1952-07-17 2002-07-22 1952-07-17 R, AMNH Fisher & W. Gertsch Heard Island and McDonald Islands 1848 36.398876 Kaweah River 1 -118.78779 14 15 1 paratype 1956-07-03 2004-04-28 1956-07-03 V, AMNH, CDU W. Gertsch & D. Ubick Heard Island and McDonald Islands Kaweah Cave 914 15 16 Mineral King Road Sequoia National Park 14 15 1 paratype 1958-07-11 2004-05-15 1958-07-11 V, AMNH, UCR Soda Creek, W & W. Gertsch & Ojai & D. Boe Heard Island and McDonald Islands Ventura Co. Soda Creek Camp Nelson 15 16 1 paratype 1980-05-06 UCB C. Griswold Heard Island and McDonald Islands 1219 Reyes Creek Woods 15 16 1 paratype 1986-07-04 CBH N Santa Paula & B. Hebert Heard Island and McDonald Islands 198 Stecker Park Highway 15 16 1 paratype 1958-07-02 1983-06-26 1958-07-02 V, AMNH, CAS, CGL W. Gertsch & Roth & G. Lowe Heard Island and McDonald Islands Wheeler Springs Yerba Buena Road 15 16 1 paratype 1946-01-24 1946-01-25 1946-01-24 J. Linsdale Hastings Natural History State Reserve 15 16 2 California holotype 1992-09-10 CDU Ubick & J. Boutin 110 16 17 1 Arroyo Seco & Indians Road Los Padres National Forest 16 17 1 1995-05-07 CDU Ubick & W. Savary 335 16 17 1 1995-07-08 CDU Ubick & S. Fend & W. Savary 16 17 1 1996-08-24 CDU Ubick 16 17 1 1999-07-03 CDU Fend & J. Ledford & D. Ubick 16 17 1 2000-04-15 CDU Ubick 16 17 1 2001-04-28 CDU Ubick 16 17 1 2001-04-27 2001-08-19 2001-04-27 CDU Ubick 335 16 17 1 2004-06-18 2004-10-24 2004-06-18 Arroyo Seco Campground Los Padres National Forest 16 17 1 1980-04-26 CDU Ubick 16 17 1 1994-05-13 CDU Ubick & J. Boutin 290 16 17 1 1994-05-13 1994-05-15 1994-05-13 CDU Boutin 274 16 17 1 1999-04-24 CDU Ubick & W. Savary 16 17 1 Arroyo Seco Campground & S The Lakes Los Padres National Forest 16 17 1 1994-05-13 1994-05-15 1994-05-13 CDU Ubick & J. Boutin 274 16 17 1 1995-05-06 CDU Ubick & W. Savary 16 17 1 2001-04-27 2001-05-18 2001-04-27 Ubick 16 17 1 2004-06-18 2004-10-24 2004-06-18 Same Big Sur 16 17 1 1937-07 AMNH Ivie 16 17 1 1952-06-25 R, AMNH M. Cazier & W. Gertsch & Schrammel 16 17 1 1953-07-18 AMNH Gertsch 16 17 1 1960-04-02 R, AMNH W. Gertsch & W. Ivie & Schrammel 14 mi S Big Sur 16 17 1 1953-12-22 V, AMNH Roth Carmel 16 17 1 1953-11-12 AMNH 16 17 1 [293,648,1451,1472] 1954-04-05 1954-05-23 1954-04-05 AMNH 16 17 1 [124,386,1477,1499] 1 mi N Carmel 16 17 1 1953-12-21 V, AMNH Roth Carmel Valley 16 17 1 1979-04-09 UCR Boe 16 17 1 1996-05 CDU Stadille 16 17 1 1991-06-01 CDU Ubick 914 16 17 1 Hastings Natural History State Reserve 16 17 1 1939-05-19 1939-05-29 1939-05-19 CAS Linsdale 16 17 1 1940-05-30 CAS Linsdale 16 17 1 [246,602,1720,1741] 1941-02-22 CAS Linsdale 16 17 1 [700,1075,192,213] 1942-05-27 CAS Linsdale 16 17 1 1944-07-18 CAS Linsdale 16 17 1 1946-01-24 1946-01-25 1946-01-24 AMNH Linsdale 16 17 1 1946-04-10 CAS Linsdale 16 17 1 1946-05-17 AMNH Linsdale 16 17 1 1946-11-18 AMNH Linsdale 16 17 1 1946-12-08 AMNH Linsdale 16 17 1 [123,544,192,213] CAS Linsdale 17 18 1 1947-06 AMNH Linsdale 17 18 1 1948-05-24 AMNH Linsdale 17 18 1 1950-05-17 CAS Linsdale 17 18 1 1950-06-24 1999-07-23 1950-06-24 AMNH, CDU J. Linsdale & S. Fend & Ledford & Ubick 17 18 1 1999-08-02 1999-08-04 1999-08-02 CAS D. Ubick & S. Fend & Ledford 17 18 1 2004-01-09 AMNH Stewart 578 36.388996 Hastings Natural History State Reserve 1 -121.548615 Big Creek 17 18 1 1979-04-10 UCR Highway G & Boe 17 18 1 1982-06-04 1982-06-06 1982-06-04 UCB Griswold Hill Big Creek Reservoir 17 18 1 2002-09-06 UCR Edwards 17 18 1 1991-05-31 CDU Ubick 305 Nacimiento Road Kirk Creek 17 18 1 1991-06-01 CDU Ubick 472 Nacimiento Road Fork Mill Creek 17 18 1 1931-08-15 AMNH Ivie Pacific Grove 17 18 1 1964-05-03 CAS Craig 91 Pfeifer Big Sur State Park 17 18 1 1987-08-19 CAS Gross 17 18 1 1960-04-02 1961-09-25 1960-04-02 R, AMNH Gertsch & Ivie & Schrammel & W. Gertsch & W. Ivie Redwood Canyon Gorda 17 18 1 1978-09-13 2002-05-27 1978-09-13 R, AMNH, UCB Doty & Lew Whale Point Cabin 200 Tassajara Spring Buddhist Big Creek Reserve 17 18 1 2002-10-24 UCB Lew 17 18 1 2003-05-12 UCB Lew 17 18 1 1958-07-03 V, AMNH W. Gertsch & Roth San Benito Co. Pinnacles National Monument June 17 18 1 1972-01-22 A. Jung Inyo Co. Titus Canyon Cave Male 17 18 2 California holotype San Bernardino Co. Medicine Cave 18 19 1 California 1972-01-22 1988-09-07 1972-01-22 R, CAS, AMNH Aalbu & Death Valley National Monument & A. Jung Inyo Co. 1219 Stone Canyon Defense Cave 18 19 1 California [124,469,1224,1247] Inyo Co. Inyo Co. 18 19 1 California 1979-03-17 1979-06-16 1979-03-17 R San Bernardino Co. 1311 Mitchell Caverns State Park Park Headquarters 18 19 1 California holotype 1988-06-20 1988-09-27 1988-06-20 R, CAS Mountains State Recreation Area 19 20 1 San Bernardino Co. San Bernardino Co. 19 20 1 California 1961-04-21 J. Beatty Pinal Co. Peppersauce Cave Canyon Male 19 20 1 Arizona holotype 1952-07-25 AMNH M. Cazier Coconino Co. Oak Creek Canyon Manzanita Camp 19 20 1 OTHER MATERIAL EXAMINED 1956-07-14 V, AMNH W. Gertsch & Roth Mexico Graham Co. Mount Graham Lower Canyon 19 20 1 ARIZONA 1973-09-11 V, CAS Roth Mexico Cajon Bonito 19 20 1 Sonora