Ommatius chiastoneurus (Speiser), 1910: 105 Hull 1962: 435 Oldroyd 1980: 347 Ommatius senex Bromley, 1936:127 Hull 1962: 436 Oldroyd 1980: 347 Afroestricus, a new Afrotropical Ommatine (Diptera: Asilidae) genus with twenty species and two species groups Scarbrough Table Of Contents, Aubrey G. Zootaxa 2005 2005-08-26 1041 1 1 76 (Speiser) Speiser [264,708,1406,1432] Insecta Asilidae Afroestricus Animalia Diptera 15 16 Arthropoda species chiastoneurus   Figs. 15–24, 193     Ommatius chiastoneurus(Speiser), 1910: 105 comb. n.  Hull 1962: 435;  Oldroyd 1980: 347.   Ommatius senex Bromley, 1936:127  syn. n.  Hull 1962: 436;  Oldroyd 1980: 347.  Redescription, male. Black. Body 11.4–15.0 mm; wing 7.1–9.3 mm. Head: Face yellowish to gray tomentose, 8 to 14 black bristles and scattered black setae present on dorsal 1/2; vestiture denser, longer, and yellowish ventrally, stout yellow bristles present medially; FHWR 1.0:4.5–1.0:5.0. Antenna brown to black; flagellum slightly longer than scape, FWLR 1.0:3.0–1.0:3.5. Frons yellow to brownish­yellow tomentose; longest setae as long as scape+pedicel combined. Occiput yellow to yellowish­gray tomentose with mostly white setae, sparse black setae sometimes present dorsally; 8 to 12 short, thick, black, postocular bristles present, two or three slightly arched forward, apex at or just before posterior margin of eye.   FIGURES 15–24.  Afroestricus chiastoneurus(Speiser). Male terminalia: 15–17. Lateral, dorsal, and ventral views. 18. Gonostylus. 19–20. Aedeagus, dorsal and lateral views. Female terminalia: 21–22. Dorsal and ventral views. 23. Spermatheca. 24. Genital fork with bulbus bursa and three spermathecal ducts.  Thorax: Pronotum dorsally brown tomentose, setae yellowish, bristles black. Mesonotum black, postalar callus sometimes partly yellowish to red; two narrow, brown, paramedial, tomentose stripes and three brown, lateral spots present, tomentum elsewhere yellow to yellowish­gray posteriorly; vestiture black, sparse whitish setae present laterally and between dorsocentrals posteriorly; anterior setae slightly longer than scape; four long, thick lateral and four to five dorsocentral bristles present, latter present at or just behind transverse suture. Scutellum yellowish­gray tomentose with yellow setae. Pleuron yellow tomentose anteriorly, yellowish­gray posteriorly, anepisternum sometimes brown with sparse black setae; setae and bristles usually yellow; anepimeral bristle yellow or black. Halter yellow to reddish­yellow.  Wing( Fig. 3a): Yellowish­brown microtrichose. Cell r 1wide apically, vein R 2+3strongly curved forward apically. Cell r 4with base just beyond apex of d cell; cell wide apically with vein R 4strongly curved anteriorly before wing margin. Crossvein r­m before or just beyond middle of cell d, much longer than CuA 1+M 3; cu­m sometimes present. Cell m 1WR 1.0:3.0:2.8–1.0:4.7:4.8.  Leg: Coxae, trochanters and femora black. Coxae white tomentose with abundant white to yellow setae. Femora mostly short yellowish setose, long ventrally, latter sometimes black; fore and middle femora black setose dorsally; bristles usually black, often white to yellow in specimens in northern part of range, especially ventrally. HFWLR 1.0:4.0–1.0:5.0. Tibiae yellow with apices of first two blackish; hind tibia usually black on apical 1/2 anteriorly; bristles mostly black, two or three lateral bristles on fore and middle tibiae yellow. Tarsi with fore basotarsomere mostly yellow, apex brown; tarsomeres otherwise mostly brown, base of tarsomere three and four narrowly yellow; fore and middle tarsi, sometimes hind tarsus, with three to six yellow bristles.  Abdomen: Mostly yellowish­gray tomentose with yellow to white setae; terga 4 to 5 medially with sparse black setae, abundant on terga 6–8; bristles yellow or black, none especially stout.  Terminalia: Black to reddish­brown, apical margins reddish to yellowish. Cercus 2.5 to 3X longer than wide. Epandrium with prominently notched dorsoapically; apex narrow, dorsal corner usually angular in dorsal view. Gonostylus distinctive. Epiphallus, narrow in dorsal view, extends vertically just beyond middle; distiphallus wide apically; ejaculatory apodeme thin. Gonocoxite short, apex well before apical angle of epandrium; apical margin obtusely rounded in ventral view, evenly contured, not raised, and significantly shorter than anteromedial corner. Hypandrium round apically. Female. Differs as follows. Body 9.7–15.3 mm; wing 7.0– 10.3 mm; FHWR 1.0:3.5– 1.0:3.7; FWLR 1.0:2.0–1.0:2.8; m 1WR 1.0:2.9:3.0–1.0:4.6:4.2; HFWLR 1.0:4.6–1.0:5.6. Head: Face with black setae extending ventrally to or beyond middle of face and 12 or more black bristles. Thorax:Setae between dorsocentral rows posteriorly often black. Hind tarsus usually with only black bristles. Abdomen: Terga usually black setose dorsally. Unusually stout yellow and black bristles present on apical segments. Terminalia( Figs. 21–24): Tergum 9 entirely sclerotized dorsally, about 1/3 as long as cercus. Sternum 8 without stout bristles. Spermatheca slightly enlarged and flattened apically, surface smooth. Genital fork distinctive.   Typesexamined.  TANZANIA: 1 M [ holotype  Ommatius chiastoneurusSpeiser], Kilimandjaro, Sjostedt1905–6, Kibonoto[Mkomazi Game Reserve]  1300–1900 m, 4 Dec( NHRS).   SOUTH AFRICA: 1 M [ holotype  O. senexBromley], New Hanover[ 29°23’S 30°32’E], Natal,  16.xii.1913, C.B. Hardenberg( NMSA); 1 M, [ paratype  O. senexBromley], New Hanover[ 29°23’S 30°32’E], Natal, 20.xii.14, C.B. Hardanberg, Bromleycoll. ( USNM). Additional material.   BURUNDI: 1 F, Urundi( Mosso)  1450 m, Makoronkwe  12.iii.1953, P. Basilewsky( MRAC).   DEMOCRATIC REPUBLIC OF CONGO: 1 M, Zimbéze, Nova Choupanga, Prés Chemba, P. Lesne1928 ( MNHN); 1 M, 1 F, Congobelge: P.N.A, Katanda,  950m  31.xi.1934, G.F. de Witte: 857 ( MRAC); 1 F, Mayya Moto,  950m,  15.xi.1934, G.F. de Witte: 758 ( MRAC).   ETHIOPIA: 2 F, Abyssinie, Mission de Conchamps, C.H. Micheland M. Potter, 1899 ( MNHN); 1 M, Abyssinia,  ix.1911, R. J. Stordy( BMNH).   KENYA: 1 F, A prey on B, vi, Nairobi Dt[ 01°17’S 36°49’E], 1911, T. J. Anderson, 1911­384 ( BMNH); 1 M, Nairobi[ 01°17’S 36°49’E], B.E.A ~ G. Babault,  iii.1943( MNHN); 1 M, Nairobi[ 01 °17S 36 °49E], B.E.A,  vi–vii.1912, Dr. A.D. Milne( MRAC); 1 F, Brit. E. Africa, Nairobi District[ 01°17’S 36°49’E],  3.v.1911, T. J. Anderson( MRAC); 1 F, Ruwenzori, Mt Karangna,  8­9000 ft,  xii.1934–  i.1935, F.W. Edwards( BMNH).   MALAWI: 1 M, Nyasaland, Chikala District[ 14°38’S 35°12’E],  i.1913, H. S. Stannus[male terminalia, slide 51] ( BMNH).   MOZAMBIQUE: 1 M, Prov. Gorongoza, Tendos du Sungoue, et Riy Vunduzi(alt  40 m), G. Vasse1907 ( MNHN).   SOUTH AFRICA: 1 M, Mmbilo, Durban[ 29°48’S 31°00’E] Natal, 21.xi.26, A. L. Bevis~ Ommatius senex, Bromley 1945? det S. W. Bromley( BMNH); 1 M, Natal: Weenen[ 28°52’S 30°05’E],  i.1925, (termalilia slide 50), H.P. Thomasset( BMNH); 2 M, South Africa: Natal, Pietermaritzburg[ 29°36’S 30°23’E] 2930cb  19.iii.1978, JGH Londt( NMSA); 2 M, 1 F, Dargle, Grassveld,  4400 ft  20.vii.1987, P.E. Reavell( NMSA); 2 M, S.A. [=South Africa]: Natal: 2429AA, Transvaal,  20 kmNE of Potgietersrus rd.to Maccapans Mts, 27.i.78, JGH Londt, long grass under tree ( NMSA); 1 M, Sth[=South] Africa Transvaal, Johannesburg[ 26°09’34”S 28°04’21”E], 2628 Aa, Highlands North, R. Elferink  21.ii.1982( NMSA); 1 M, Sth[=South] Africa Transvaal, Johannesburg[ 26°09’34”S 28°04’21”E], 2628 Aa, Highlands North, R. Elferink  7.iii.1982( NMSA); 1 M, S.A. [=South Africa] Natal, Pietermaritzburg[ 29°36’S 30°23’E], Scottsville,  15– 21.iv.1986, R. M. Miller, Malaise Trap( NMSA); 1 M, S.A. [=South Africa] Natal, Durban[ 29°48’S 31°00’E] SE2931 CC, WW Strglingh Bluff, golf course 11.iv.87 ( NMSA); 1 M, S.A. [South Africa], Natalnr Itala[ 27°32’S 31°19’E], Game ReserveGraig Adam Dam 27°28’S 31°25’E,  ii.1991 R. M. Miller, Malaise Trap( NMSA); 1 M, S.A. [=South Africa]: Natal, Pietermaritzburg[ 29°36’S 30°23’E], Montrosein garden 2930 CB, JGH Londt,  12.xii.1981( NMSA); 1 M, 1 F, South Africa: Natal,  15 kmSE Ingwavuma,  21.ii.19792732AA, JGH Londt, bushy area with big trees ( NMSA).   UGANDA: 1 M, Mafunga Forests, Kiozi,  vi.1951, Van Someren. Com. Inst. Ent. Coll. No.12301 ( BMNH); 1 M, Entebbe[ 0°04’N 32°38’E],  1–15.v.1983, G.G.M. Schulten( ZMAN); 7 M, 7 F, Entebbe[ 0°04’N 32°38’E], Kisubi Forest,  9–10.v.1976, M.K. Paulus( CNCI); 4 M, 3 F, Entebbe[ 0°04’N 32°38’E], Kisubi Forest,  8–9.vi.1976, M. K. Paulus( CNCI); 1 F, Entebbe[ 0°04’N 32°38’E], Kisubi Forest,  24.iv.1976, M. K. Paulus( CNCI); 2 M, Nr. Entebbe[ 0°04’N 32°38’E],  1–13.ii.1973, 1160 m, H. Falke( CNCI); 19 M, 19 F, Entebbe[ 0°04’N 32°38’E],  10.iii.1972, in forest, H. Falke( CNCI); 6 M, Entebbe[ 0°04’N 32°38’E],  10.iii.1972, in forest, H. Falke( CNCI); 2 F, 7 mi. N. Entebbe[ 0°04’N 32°38’E], nr. Lk. Victoria,  2.iii.1972, 1135 m, H. Falke( CNCI); 1 M, Entebbe[ 0°04’N 32°38’E],  10.iii.1972, in forest, H. Falke( CNCI); 1 F, Uganda: Entebbe,  25.vi.1972, H. Falke( FSCA).   ZIMBABWE: 1 M, among grass, Salisbury[= Harare 17°50’S 31°03’E], S. Rhodesia, Dept. Agric,  3.i.1937, A. Cuthbertsoncoll, Bromleycoll. ( USNM); 1 M, S. Rhodesia, Dept. Agric,  28.i.1936, Bromleycoll ( USNM). Distribution ( Tables 1, 2; Fig. 193). Wide spread in eastern Africa; collected up to 2647 meters from October to June Remarks.  Afroestricus chiastoneurusis distinguished from congeners by the combination of black femora and characters of the terminalia ( Figs. 15–24).  Afroestricus chiastoneurus, A. macroscelis,  A. insectaris,and  A. varipeshave black femora. However, the femora are entirely black in  A. chiastoneurusand  A. macrosceliswhereas they are only partly black in the latter two species, i.e. brownish­yellow to brownish dorsally in  A. varipesor yellow apically in  A. insectaris. Females of  A. chiastoneurusand  A. macroscelisare essentially identical externally but each has distinctive genital forks ( Figs. 24, 80).  Bromley (1936), uncertain of the identity of  O. variabilis, described  O. senex,a black South African species with black femora and terminalia. Apparently, he was not aware of  A. chiastoneurus( Speiser 1910). My examination of the typesof the two species reveals that they are conspecific. Since  A. chiastoneuruswas described earlier than  O. senex, the former name has priority and is retained for the species.  Ommatius senexis here declared a new synonmynof  A. variabilis. NHRS Sjostedt & Kibonoto & Dec Tanzania 1600 Types 18 19 1 Kilimanjaro holotype 1913-12-16 NMSA Natal & C. B. Hardenberg South Africa -29.383333 New Hanover 1227 30.533333 18 19 1 holotype USNM Natal & C. B. Hardanberg & Bromley South Africa -29.383333 New Hanover 1227 30.533333 18 19 1 paratype 1953-03-12 MRAC P. Basilewsky Burundi Urundi 1450 Makoronkwe Mosso 18 19 1 MNHN Pres Chemba, P & Lesne Democratic Republic of the Congo Nova Choupanga Zimbeze 18 19 1 1934-11-31 MRAC G. F. de Witte Democratic Republic of the Congo 950 Katanda Congo 18 19 1 1934-11-15 MRAC G. F. de Witte Democratic Republic of the Congo 950 Moto May 18 19 1 MNHN C. H. Michel & M. Potter Ethiopia Mission de Conchamps Abyssinie 18 19 1 1911-09 R, BMNH J. Stordy Ethiopia Abyssinia 18 19 1 T, BMNH J. Anderson Kenya -1.2833333 Nairobi Dt 1308 36.816666 18 19 1 1943-03 MNHN G. Babault Kenya -1.2833333 Nairobi 1308 36.816666 18 19 1 1912-06 1912-07-31 1912-06 MRAC Kenya -1.2833333 Dr. A. D. Milne 1308 36.816666 Nairobi 18 19 1 1911-05-03 T, MRAC Brit. E. Africa & J. Anderson Kenya -1.2833333 Nairobi District 1308 36.816666 18 19 1 1934-12 1935-01 1934-12 BMNH F. W. Edwards Kenya 1373 Mt Karangna Ruwenzori 18 19 1 1913-01 BMNH H. S. Stannus Malawi -14.633333 Chikala District 1287 35.2 Nyasaland 18 19 1 MNHN Tendos du Sungoue & G. Vasse Mozambique 40 Riy Vunduzi Prov. Gorongoza 18 19 1 BMNH Natal & A. L. Bevis & S. W. Bromley South Africa -29.8 Durban 1225 31.0 Mmbilo 18 19 1 1925-01 BMNH H. P. Thomasset South Africa -28.866667 Weenen 1230 30.083334 Natal 18 19 1 1978-03-19 JGH, NMSA Londt South Africa South Africa -29.6 Pietermaritzburg 1226 30.383333 Natal 18 19 1 [386,1250,1244,1270] 1987-07-20 NMSA P. E. Reavell South Africa 1341 Grassveld Dargle 18 19 1 JGH, NMSA Londt South Africa Natal Maccapans Mts Transvaal 18 19 1 1982-02-21 R, NMSA Aa & Highlands North South Africa Sth -26.159445 Johannesburg 20 28.0725 Africa Transvaal 18 19 1 1982-03-07 R, NMSA Aa & Highlands North South Africa Sth -26.159445 Johannesburg 20 28.0725 Africa Transvaal 18 19 1 1986-04-15 1986-04-21 1986-04-15 R, NMSA Scottsville & M. Miller South Africa -29.6 Pietermaritzburg 1226 30.383333 Natal 18 19 1 CC, WW, NMSA Strglingh Bluff South Africa -29.8 Durban 1225 31.0 Natal 18 19 1 1991-02 R, NMSA M. Miller South Africa Natal -27.466667 Game Reserve 1237 31.416666 Itala 18 19 1 1981-12-12 CB, JGH, NMSA Montrose South Africa Natal -29.6 Londt 1226 30.383333 Pietermaritzburg 18 19 1 1979-02-21 JGH, NMSA Londt South Africa Natal South Africa 18 19 1 1951-06 BMNH Van Someren. Com. Inst. Ent. & Coll. No. Uganda Kiozi Mafunga Forests 18 19 1 1983-05-01 1983-05-15 1983-05-01 ZMAN G. G. M. Schulten Uganda 0.06666667 Entebbe 1308 32.633335 18 19 1 1976-05-09 1976-05-10 1976-05-09 CNCI M. K. Paulus Uganda 19 20 0.06666667 Kisubi Forest 1308 32.633335 Entebbe 18 19 1 1976-06-08 1976-06-09 1976-06-08 CNCI M. K. Paulus Uganda 0.06666667 Kisubi Forest 1308 32.633335 Entebbe 19 20 1 1976-04-24 CNCI M. K. Paulus Uganda 0.06666667 Kisubi Forest 1308 32.633335 Entebbe 19 20 1 1973-02-01 1973-02-13 1973-02-01 CNCI M, Nr. Entebbe & Falke Uganda 0.06666667 1308 32.633335 19 20 1 1972-03-10 CNCI H. Falke Uganda 0.06666667 Entebbe 1308 32.633335 19 20 1 1972-03-10 CNCI H. Falke Uganda 0.06666667 Entebbe 1308 32.633335 19 20 1 1972-03-02 CNCI N. Entebbe & Lk. Victoria & H. Falke Uganda 0.06666667 1308 32.633335 19 20 1 1972-03-10 CNCI H. Falke Uganda 0.06666667 Entebbe 1308 32.633335 19 20 1 1972-06-25 FSCA H. Falke Uganda Entebbe Uganda 19 20 1 1937-01-03 USNM S. Rhodesia & A. Cuthbertson & Bromley Zimbabwe -17.833334 Dept. Agric 1277 31.05 Salisbury 19 20 1 Harare 1936-01-28 USNM M, S. Rhodesia & Bromley Zimbabwe Dept. Agric 19 20 1