Revision of the Psilota Meigen, 1822 flower flies (Diptera: Syrphidae) of Australia Young, Andrew D. Skevington, Jeffrey H. Steenis, Wouter Van Zootaxa 2020 2020-02-17 4737 1 1 127 4PJPH Curran, 1926 Curran. A 1926 [151,541,1269,1296] Insecta Syrphidae Psilota Animalia Diptera 25 26 Arthropoda species auricauda   Diagnosis: Sexually dimorphic. Both sexes with cuticle entirely shining black, scutum and pleuron with faint blue sheen. Scutellum densely white pilose, with at most a few scattered dark pile on the apical rim. Halter orange. Calypter pale, with brown rim, calypter pile pale. Hind leg simple. Female only with bright yellow pile on tergites 3, 4 and 5. Yellow pile on apical half of tergite 4 long and dense. Body length: 12.0– 15.4 mm.   Description: MALE: Head:Head black, white pollinose. Face white pilose, frons and vertex black pilose. Antenna brown, postpedicel oval, approximately 1.5 times as long as wide. Eye densely pilose, pile white on lower half and dull orange to brown on upper half.  Thorax:Thorax black, faintly reflective blue. Femora dark brown, tibiae and tarsi dull orange to brown. Hind femur unmodified, with only a few weak, black setulae apicoventrally. Pleuron white pollinose. Pile on pleuron and scutellum entirely pale, with at most a few black pile on apical rim of scutellum. Pile on scutum mixed black and pale yellow. Wing mostly colourless, with only stigma and cell sc brown. Cells bc, c, and bm entirely bare. Anterior half of cells br, cua, m 4, and dm bare. Alula entirely microtrichose. Halter orange. Calypter pale with a dark brown border.  Abdomen:Abdomen black. Tergite 2 mostly pale pilose. Tergite 3 with some dark pile, mostly pale pile laterally. Tergite 4 mostly dark pilose.  Genitalia:Epandrium compact, about as long as tall. Cercus approximately triangular. Outer lobe of surstylus inserted near the top of the inner lobe, thin, and abruptly curved downwards at about one-third its length. Inner lobe of surstylus long, thin, and straight, approximately 5 times as long as wide (at base), slightly swollen at base, otherwise parallel sided over entire length, somewhat blunt tipped. Phallapodeme with a small, downward facing spur on ventral side. Postgonite slightly darkened.   FIGURE 8.  P. auricauda  sp. nov., A) male dorsal habitus, B) male anterolateral face, C) male lateral habitus. Scale bar =1 mm.   FIGURE 9.  P. auricaudaCurran. A) female dorsal habitus, B) female anterolateral face, C) female lateral habitus. Scale bar =1 mm.  FEMALE:Similar to male, differing as follows: Abdomen:Tergites 3, 4 and 5 bright yellowish-orange pilose. Pile of tergite 3 and anterior half of tergite 4 sparse. Pile on posterior half of tergite 4 and all of tergite 5 long and dense.  Specimens examined:  HOLOTYPE: AUSTRALIA:  Queensland: Brisbane, -27.527 153.040,  15.ix.1915, D.H. Colless, ANIC29014845( 1♀, ANIC); Other material: Australian Capital Territory: Canberra, -35.282000 149.128686,  23.xi.1959, K.R. Norris, ANIC29034797( 1♀, ANIC);   New South Wales: Sydney, -33.868819 151.168819,  14.x.1915, Jeff_ Skevington_ Specimen30411 ( 1♀, MCZ); Cheltenham, -33.758389 151.075689, xixii.1949, N.W. Rodd, AustralianMuseumK407023 ( 1♀, AM); Dorrigo, -30.365542 152.738950, W. Heron, Jeff_ Skevington_Specimen45813 ( 1♀, USNM);  Gibraltar Ra. N.P., Waratah track, -29.5 152.333333, on  Leptospermumsp. Blossom,  10-12.xi.1984, D.K. Yeates, UQIC_Reg94618 ( 1♀, QM);  Queensland:  8 kmWest of Tyalgum, -28.369994 153.208767,  24.ix.1983, N.W. Rodd, AustralianMuseumK407296 ( 1♀, AM);  Amiens State Forestnear. Stanthorpe, -28.600522 151.799608,  24-26.xi.1981, UQIC_Reg94619 ( 1♀, QM);  Beerburrum, -26.970639 152.968256,  16.ix.1971, E.C. Dahms, QM_ REG._NO._T223163 ( 1♀, QM); Brisbane, -27.526975 153.040322,  15.ix.1915, D.H. Colless, ANIC29014845( 1♀, ANIC);   23.x.1911, H. Hacker, ANIC29034792( 1♀, ANIC);   24.ix.1914, H. Hacker, QM_ REG._NO._T223161 ( 1♀, QM);  25.ix.1911, H. Hacker, QM_ REG._NO._T223159 ( 1♀, QM);  25.ix.1916, H. Hacker, QM_ REG._NO._T223160 ( 1♀, QM);  25.ix.1919, H. Hacker, CNC_  Diptera255064 ( 1♀, ANIC);  7.ix.1927, J. Mann, UQIC_ Reg94822 ( 1♀, QM);  7.ix.1941, C.F. Ashby, ANIC29034790( 1♀, ANIC);   ix.1929, G.H. Hardy, AustralianMuseumK407241, K410316 ( 1♂, 1♀, AM);  xi.1928, G.H. Hardy, ANIC29034795, UQIC_ Reg94810 ( 1♂, UQIC, 1♀, ANIC); Camiranear Ipswich, -27.634853 152.915925,  12.ix.1981, G. Daniels, GDCBReg_19612 ( 1♀, GDCB);  Canungra, -28.017142 153.164708,  30.iii.1937, R.F. Langdon, UQIC_ Reg94620 ( 1♀, QM);  Cimara, nr. Ipswich, -27.493986 152.944925,  12.ix.1981, G. Daniels, GDCBReg_19613 ( 1♀, QM);  Clarence, Blue Mountains, -33.482308 150.224264,  21.xi.1977, N.W. Rodd, AustralianMuseumK407297 ( 1♀, AM);  Cleveland, -27.533333 153.266667,  viii.1924, G.H. Hardy, ANIC29034796( 1♀, ANIC);  Dayboro,  0.5 kmSouth, -27.200000 152.816667,  18-19.ix.1999, S.G. Evans, QM_ REG._NO._T223162 ( 1♀, QM); Lansdowne, - 31.758753 152.534258, Drysclerophyll,  19.i.1981, G. Williams, AustralianMuseumK407022 ( 1♀, AM);  Mt. Marlay, near Stanthorpe, S.E.Q., -28.655786 151.947126,  6.xi.1984, D.K. Yeates, UQIC_Reg94621 ( 1♂, QM);  Mt. Tinbeerwahnear Cooroy, -26.384206 152.966639,  6.ix.1980, G. Daniels, GDCBReg_19610-19611 ( 2♀♀, GDCB); One Tree Hill, Brisbane, -27.526975 153.040322,  12.xii.1925, A. Musgrave, AustralianMuseumK407024 ( 1♀, AM);  Rockpool Gorge Mt. Walsh Nat. Park, Biggenden, -25.605889 152.027519,  4.x.1976, H. Frauca, CNC_  Diptera253687 ( 1♂, ANIC); Rockpool Gorge, Mount Walsh National Park, Biggenden, -25.605889 152.027519,  4.x.1976, H. Frauca, ANIC29034794, CNC_  Diptera255065-255066, USNM_ ENT249821-249822( 1♂, 4♀♀, ANIC);  S.Q., on  Leptospermum,  21.ix.1959, Mackerras, UQIC_ Reg94564 ( 1♂, QM); Stanthorpe, -28.657928 151.930344, 1924, Dateis ambiguous reads ”1.10.24”, ANIC29034791( 1♀, ANIC);  Stockyard Creek, SE of Ca- palaba, -27.550000 153.166667, 27°33’S 153°10’E  22.ix.1985, G. and A. Daniels, GDCBReg_19561 ( 1♂, QM);  Sunnybank, -27.578997 153.059006,  27.viii.1950, I.M. Mackerras, ANIC29034793( 1♀, ANIC);  The Blunder, Brisbane, -27.526975 153.040322,  18.ix.1966, C.F. Ashby, CNC_  Diptera255062 ( 1♀, ANIC);  2.x.1966, C.F. Ashby, CNC_  Diptera255061 ( 1♂, ANIC);  8.ix.1962, C.F. Ashby, CNC_  Diptera255063 ( 1♀, ANIC);  26.ix.1966, C.F. Ashby, ANIC29034788( 1♀, ANIC);   4.x.1968, C.F. Ashby, ANIC29034787( 1♀, ANIC);  Bluff Range S.[South] Slope, Biggenden, -25.600000 152.050000,  1000m,  15.viii.1973, H. Frauca, ANIC29034789 ( 1♀, ANIC).   Discussion:This is the first time the male of this species has been described, as the original description by Curran was from female specimens only. 2565883828 1915-09-15 ANIC Colless Australia -27.527 Brisbane 74 153.04 28 29 ANIC29014845 1 1 Queensland holotype 2565883817 1959-11-23 ANIC Norris Australia -35.282 Canberra 1 149.1287 28 29 ANIC29034797 1 1 Australian Capital Territory 2571544671 1915-10-14 MCZ Australia -33.86882 151.16882 28 29 1 1 New South Wales 2571544677 AM N.W. Rodd Australia -33.75839 151.07568 28 29 1 1 New South Wales 2571544662 USNM W. Heron Australia -30.365541 152.73895 28 29 1 1 New South Wales 2571544670 1984-11-10 QM Gibraltar Ra. N.P. & D.K. Yeates Australia 28 29 1 1 New South Wales 2571544666 1983-09-24 AM N.W. Rodd Australia -28.369993 153.20877 28 29 1 1 Queensland 2571544664 1981-11-24 QM Australia -28.600521 151.7996 28 29 1 1 Queensland 2571544669 1971-09-16 QM Beerburrum & E.C. Dahms Australia -26.970638 152.96826 28 29 Queensland 2565883836 1915-09-15 QM, ANIC D. H. Colless Australia -27.526976 Brisbane 1 153.04033 28 29 ANIC29014845 2 2 New South Wales 2565883826 [820,1432,621,647] 1911-10-23 ANIC Hacker Australia 28 29 ANIC29034792 1 1 New South Wales 2565883831 [151,451,657,683] 1914-09-24 QM Hacker Australia 28 29 1 New South Wales 2565883811 [465,1128,656,683] 1911-09-25 QM Hacker Australia 28 29 1 1 New South Wales 2565883816 1916-09-25 QM Hacker Australia 28 29 1 1 New South Wales 2565883825 [550,1254,692,719] 1919-09-25 QM, CNC Hacker Australia 28 29 1 1 New South Wales 2565883822 1927-09-07 1941-09-07 1927-09-07 ANIC, QM Mann & Reg & Ashby Australia 28 29 ANIC29034790 3 3 New South Wales 2565883832 [151,1433,764,791] 1928-11 1929-09 1928-11 AM Hardy Australia 28 29 ANIC29034795 2 1 1 New South Wales 2571544684 1981-09-12 UQIC G. Daniels Australia -27.634853 152.91592 28 29 2 1 1 2571544673 1937-03-30 QM R.F. Langdon Australia -28.017141 153.1647 28 29 1 1 2571544676 [285,1432,872,899] 1981-09-12 QM G. Daniels Australia -27.493986 152.94493 28 29 1 1 2571544682 1977-11-21 AM N.W. Rodd Australia -33.482307 150.22426 28 29 1 1 2571544686 [222,1230,944,971] 1924-08 ANIC G.H. Hardy Australia -27.533333 153.26666 28 29 ANIC29034796 1 1 2565883809 1999-09-18 1999-09-19 1999-09-18 QM Evans Australia -27.2 Dayboro 1 152.81667 28 29 1 New South Wales 2571544655 1981-01-19 QM G. Williams Australia -31.758753 152.53426 28 29 1 1 2571544650 1984-11-06 QM Stanthorpe, S.E.Q. & D.K. Yeates Australia -28.655787 151.94713 28 29 1 1 2571544679 1980-09-06 GDCB G. Daniels Australia -26.384207 152.96664 28 29 2 2 2571544681 1925-12-12 AM A. Musgrave Australia -27.526976 153.04033 28 29 1 1 2571544687 [277,1422,1160,1187] 1976-10-04 CNC Biggenden & H. Frauca Australia -25.605888 152.02751 28 29 2565883829 1976-10-04 ANIC, CNC H. Frauca Australia Rockpool Gorge -25.605888 Biggenden 1 152.02751 Mount Walsh National Park 28 29 ANIC29034794 1 1 New South Wales 2565883824 USNM, ANIC Australia 28 29 ENT249821-249822 5 4 1 New South Wales 2565883830 1959-09-21 QM, ANIC Stanthorpe, S.E.Q. & Mackerras & Reg & Date Australia -28.657928 Stanthorpe 1 151.93034 28 29 ANIC29034791 2 1 1 New South Wales 2571544663 1985-09-22 QM Daniels Australia -27.55 153.16667 28 29 1 1 2571544683 [151,1270,1376,1403] 1950-08-27 ANIC Mackerras Australia -27.578997 153.059 28 29 ANIC29034793 1 1 2565883814 1966-09-18 CNC Ashby Australia -27.526976 Brisbane 1 153.04033 The Blunder 28 29 1 New South Wales 2565883819 1966-10-02 ANIC, CNC Ashby Australia 28 29 1 1 New South Wales 2565883820 [311,963,1448,1475] 1962-09-08 ANIC, CNC Ashby Australia 28 29 1 1 New South Wales 2565883812 1966-09-26 ANIC Ashby Australia 28 29 ANIC29034788 2 2 New South Wales 2565883823 [647,1244,1485,1511] 1968-10-04 ANIC Ashby Australia 28 29 ANIC29034787 1 1 New South Wales 2565883810 1973-08-15 ANIC Bluff Range S. & Frauca Australia 1000 -25.6 Biggenden 1 152.05 Slope 28 29 ANIC29034789 1 New South Wales 2565883818 [1294,1433,1521,1547] ANIC Australia 28 29 1 1 New South Wales