Frolov & Scholtz, 2005 : 385 An update to the taxonomy of the genus Macroderes Westwood 1842 (Coleoptera: Scarabaeidae: Scarabaeinae) with descriptions of new species from South Africa Abdalla, Ishtiag H. Deschodt, Christian M. Scholtz, Clarke H. Sole, Catherine L. Zootaxa 2018 2018-10-23 4504 1 41 75 3X4TF Abdalla & Deschodt Abdalla & Deschodt & Scholtz & Sole 2018 [151,743,1829,1856] Insecta Scarabaeidae Macroderes GBIF Animalia Coleoptera 7 48 Arthropoda species cederbergensis sp. nov.  ( Figs. 20, 29, 76, 83, 94)  urn:lsid:zoobank.org:act: 9FED0DC0-CFF6-424C-A3AE- 092656B24972     Macroderes minutus  Frolov & Scholtz, 2005: 385(partim).   Type locality.Cederberg Mts, Jeep Track, 32°26'S 19°13'E.  Type material.  Holotype, ♂( TMSA): “ Cederberg Mts/ Jeep Track/ 32° 26' S 19° 13' E/  1.IX.1981/ groundtraps with faeces and meat bait / 63 days / Endrödy – Youngaleg [printed] // HOLOTYPE/  Macroderes cederbergensis/Abdalla & Deschodt / 2018 [red label, printed]”.  Paratypes: 2 ♀♀( TMSA): “ Cederberg Mts/ Jeep Track/ 32° 26' S 19° 13' E/  1.IX.1981/ groundtraps with faeces and meat bait / 63 days / Endrody – Youngaleg [printed] // PARATYPE/  Macroderes cederbergensis/Abdalla & Deschodt / 2018 [red label, printed]”;  1 ♂: “ Cederberg Mts./ Jeep Track/ 32° 23' S 19° 24' E/  1.IX.1981/ groundtrap with meat bait / 63 days / Endrödy— Youngaleg [printed] // PARATYPE/  Macroderes cederbergensis/Abdalla & Deschodt / 2018 [red label, printed]”;  1 ♂and 1 ♀( TMSA): “ Cederberg Mts/ Jeep Track/ 32° 24' S 19° 10' E/  1.IX.1981/  Redunca rufo[name could not be traced, probably  Pelea capreolus(Forster, 1790)or Grey Rhebok] dung / Endrödy—Youngaleg [printed] // PARATYPE/  Macroderes cederbergensis/Abdalla & Deschodt / 2018 [red label, printed]”;  2 ♂♂( TMSA) with the same data but collected from groundtraps with meat and feaces bait set up for 63 days.   Holotypedescription. Holotype, ♂( Fig. 20). Body length 9.4 mm, width 6.8 mm.  Head.Genae right–angled. Frontal suture visible, not raised, strongly curved, no tubercle. Genal sutures barely discernible. Dorsal surface of clypeus behind frontal suture punctate punctures small, irregular and separated by 1– 3 puncture diameters, area between punctures slightly rugose proximally of frontal suture. Dorsal part of eyes clearly visible.   Pronotum.Slightly convex. Anterior lateral angles almost right–angled, posterior lateral angles fairly rounded. No excavation antero-laterally. Lateral border not punctate. Dorsal surface smooth, with regular punctures separated by 0.5–1.5 puncture diameters ( Figs. 76, 83).  Elytra.Elytral interstriae 1–8 flat, shagreened, irregularly punctate, punctures irregularly sized, separated by 0.5–3 puncture diameters, margins of punctures not carinate ( Fig. 94). Striae punctate with punctures separated by 2–4 puncture diameters. Sutural interstria narrow with single row of irregularly spaced punctures. Elytral stria 9 2/ 3 the length of elytron; interstria 10 about 5 times narrower than interstria 9 inthe middle of elytron.  Aedeagus.The sclerite of internal sac is curved, with small lateral process apically ( Fig. 29).  Size range.Males length: 7.5–9.0 mm, width: 5.5–6.8 mm; females length: 7.8–9.5 mm, width: 6.0– 6.3 mm.  Female.Female differs from male by having elytral interstriae 1–8 less shagreened; otherwise variation between male and female is very slight.   Etymology.The specific epithet is derived from the Cederberg Range where the species occurs.   Differential diagnosis.It is close to  M. soleiana new speciesbut can be separated by having flatter elytral intervals ( Figs. 20, 94) and the striae being slightly arcuate and not straight as in  M. soleiana new species.   Habitat and distribution.This species was collected on the highest plateau in the Cederberg range ( Fig. 2), above 1100 ma.s.l. (metres above sea level), characterised by clay soils and classified as Northern Inland Shale Band Vegetation ( Mucina & Rutherford 2006).  Macroderes soleiana new speciesoccurs below 1100 ma.s.l.   Remark.See  M. minutusfor an explanation of the origin of the typeseries. 2028137725 1981-09-01 TMSA Younga -32.433334 Cederberg Mts 1211 19.216667 Jeep Track 8 49 1 1 holotype 2028137711 1981-09-01 TMSA Younga -32.433334 Cederberg Mts 1211 19.216667 Jeep Track 8 49 2 2 paratype 2028137719 1981-09-01 Younga -32.383335 Cederberg Mts. 1211 19.4 Jeep Track 8 49 1 1 paratype 2028137718 1981-09-01 TMSA Endrody-Younga -32.4 Cederberg Mts 1211 19.166666 Jeep Track 8 49 2 1 1 paratype 2028137724 1981-09-01 TMSA Endrody-Younga -32.4 Cederberg Mts 1211 19.166666 Jeep Track 8 49 2 2 paratype