Pimpla croceipes Cresson 1874: 398 Coccygomimus Coccygomimus P . molesta Darwin wasps of the subfamily Pimplinae (Hymenoptera: Ichneumonidae) of Mexico: genera Apechthis Förster, Itoplectis Förster and Pimpla Fabricius Khalaim, Andrey I. Ruíz-Cancino, Enrique Zootaxa 2021 2021-11-25 5071 4 451 491 4HXD8 1070301 Cresson, 1874 Cresson 1874 [181,568,1591,1618] Insecta Ichneumonidae Pimpla Animalia Hymenoptera 17 468 Arthropoda species croceipes  ( Fig. 29)      Pimpla croceipes  Cresson 1874: 398[ ♀ ♂(ANSP), Mexico, Veracruz, Orizaba].  References. Cameron 1886: 266[ Mexico( Veracruz), Guatemala, Costa Rica, Panama]. Morley 1914: 63[remarks; Mexico( Guerrero), Guatemala, Panama, Costa Rica]. Cresson 1916:26[ lectotype“603”(ANSP) designated; Mexico, Veracruz]. Townes & Townes 1966: 25[  Coccygomimus; catalogue; Mexico]. Porter 1970a: 87[  Coccygomimus; description; Colombia, Ecuador]. Maes 1989: 33[catalogue; Nicaragua]. Gauld 1991: 523[description; Mexico( Guerrero), Guatemala, Costa Rica, Panama]. Gauld et al. 1998: 140[in key; Costa Rica]. Hernández-Aguilar et al. 1999: 36[ Mexico( Tamaulipas)]. Díaz 2000: 248[ Venezuela]. Ruíz-Cancino et al. 2002: 645[checklist; Mexico]. Gauld et al. 2002: 60[ El Salvador]. Hernández-Aguilar et al. 2005: 470[ Mexico( Tamaulipas)]. Ruíz-Cancino et al. 2010: 66[checklist; Mexico( Tamaulipas)]. Veijalainen et al. 2011[DNA barcoding; separated from  P. molesta; Guatemala, Honduras, Costa Rica, Venezuela]. Sánchez-García et al. 2015: 828[checklist; Mexico( Oaxaca)]. Khalaim et al. 2018: 23[ Mexico( Veracruz)].   Remarks.The species was described from syntypefemale(s) and male(s) (Cresson 1974: 398); later the specimen under number “603” was designated as a lectotype, but its sex was not mentioned ( Cresson 1916: 26); and only in 1966, Townes ( Townes & Townes 1966: 25) recorded that the lectotypeis female.   FIGURES 28–31.  Pimpla chamela  sp. nov., female: 28—habitus (without head and wings), lateral.  Pimpla croceipes, female: 29—habitus (without antennae and wing apices), lateral.  Pimpla croceiventris, male: 30—habitus (without antennae and wing apices), lateral; 31—mesosoma, dorsal. Unfortunately, we could not confidently separate  P. croceipesand  P. molestaon the basis of morphological characters given by Veijalainen et al. (2011). Almost all examined Mexican specimens have subtegular ridge black or only dark reddish brown, fore coxa black or black and yellow, second tergite sparsely to closely punctate (in females), and transverse groove on the second tergite situated slightly behind its middle. Only one femalefrom Lagos de Montebello, Chiapas, has subtegular ridge conspicuously yellow marked and fore coxa 50×50 yellow and black. All examined specimens from Mexicoare considered here as  P. croceipes(s. lato) but further investigation of this material is required. See also Remarkssection under  P. molesta.   Material examined.   MEXICO. Tamaulipas: 1 ♀( UAT) Cd. Victoria, Cañón de la Libertad,  800 m,  13.xii.1987, coll. R. MezaV.  1 ♀( UAT) same locality and collector,  1000 m,  5.iii.1988.  1 ♀( UAT) 15 kmSSW of Cd. Victoria, El Madroño,  16–18.xii.2008, coll. A.I. Khalaim.  1 ♂( UAT) same locality,  20.v.1984, coll. H. RuízC.  1 ♀( UAT) same locality and collector,  29.i.1985.  4 ♀( UAT) same locality,  23.xii.1987, coll. H. & E. RuízC., S. Arrambide.  3 ♀, 2 ♂( UAT) same locality,  1400 m,  20.i.2000, coll. E. RuízC. & J.A. Martínez Ramirez.  1 ♀( UAT) Tula,  1000 m,  7.xi.2007, coll. E. RuízC.  2 ♀( UAT) Tula, Ejido Acahuales,  1400 m, oak-forest, Malaise trap, 27.viii–10.ix and  29.x–12.xi.2016, coll. E. RuízC. & J.M. Coronado B.  1 ♂( UAT) Gómez Farías,  v.1992, coll. J.M. CoronadoB.  1 ♀( UAT) Gómez Farías, Altas Cimas, Malaise trap,  4–11.xi.2000, coll. D.R. Kasparyan.  3 ♂( UAT) Gómez Farías, Canindo, Malaise trap,  20–21.vii.1994, coll. J.B. Wooley.  2 ♂( UAT) same locality and collector,  28–30.vii.1993.  1 ♀( UAT) Gómez Farías, Río El Cielo,  3.iii.1989, coll. R. Thompson.   San Luis Potosí: 1 ♂( UAT) Xilitla, Las Pozas(park),  21–23.xii.2008, coll. A.I. Khalaim.   Querétaro: 2 ♀( UAQ) Jalpan de Serra,  4.x.2013.  1 ♀( UAQ) Pinal de Amoles,  2.x.2013.   Veracruz: 3 ♀, 6 ♂( UAT) SW of Xalapa, park near Institute de Ecología, 19°30.768’N, 96°56.349’,  1260 m,  17–18.ii.2009, coll. A.I. Khalaim.  5 ♀, 4 ♂( 3 ♀and 2 ♂in UAT, 2 ♀and 2 ♂in ZISP) same data, but  12–14.iv.2014.  1 ♂( EMEC) Fortínde las Flores,  25.xii.1963, coll. C.A. Toschi& M.J. Tauber.   Morelos: 1 ♀( UAT) N of Cuernavaca, Chamilpa,  20.ix.1991, coll. G. Peña.  1 ♀( UAT) Tepoztlán, 18°59’51”N, 99°05’55”W,  1490 m,  5.iii.2010, coll. E. RuízC.  2 ♀( CIUM) Tlaquiltenango,  5 kmW of Santiopa, 18.44701°N, 98.98758°W,  1005 m,  5.x.2013, coll. J.A. Hernández et al.   Oaxaca: 1 ♀, 2 ♂( UNAM) Santiago Comaltepec, La Esperanza, N17.02661º, W 96.36950º,  1600 m, Malaise trap,  1.vii–12.viii.2008, coll. A. Lopez García.  2 ♀( UNAM) Santiago Comaltepec, N 17.58429º, W 96.49398º,  2332 m,  6.vi.2009, coll. H. Clebsch& A. ZaldívarR.  3 ♂( TAMU)  15.1 mi.N of San Gabriel Mixtepec,  3850 ft.(=  1175 m), Highway131,  11.vii.1987, coll. R. Wharton.   Chiapas: 10 ♀( UANL, UAT) Reserva El Triunfo,  15 kmS of La Trinitaria, Palo Gordo,  20–22.vii.1997, coll. A. GonzálezH.  1 ♀( UAT)  50 kmSSE of Comitán, National Park Lagos de Montebello, 16°06.223’N, 91°44.072’W,  1450 m, mixed forest with pines,  5–6.iii.2013, coll. A.I. Khalaim.  1 ♂( UAT)  50 kmESE of Comitán, National Park Lagos de Montebello, 16.106551, -91.705158,  1500 m,  8.viii.2021, coll. A.I. Khalaim.  1 ♀( UAT)  40 kmE of Comitán, National Park Lagos de Montebello, 16.132282, -91.729243,  9.viii.2021, coll. A.I. Khalaim.  1 ♂( TAMU) Lagunas de Montebello[National Park], Laguna Cinco Lagos,  11.viii.1990, coll. J.C. Schaffner.  1 ♂( TAMU) Reserva El Triunfo, Sendero Palo Gordo,  6400 ft.(=  1950 m), screen sweep,  21.vii.1977, coll. J.B. Woolley.  1 ♀, 4 ♂( TAMU) Reserva El Triunfo,  6620 ft.(=  2020 m),  14–23.vii.1997, coll. J.B. Woolley.  1 ♀( TAMU)  4 kmW of San Cristóbal[de las Casas], SanFelipe,  2200 m, oak-grass woodlands,  24–28.viii.1990, coll. R.W. Jones.  1 ♀( TAMU)  4 kmW of San Cristóbal[de las Casas], Reserva Huitepec,  7800 ft.(=  2380 m), cloud forest, Malaise trap,  5.x–25.xi.1990, coll. R.W. Jones.  2 ♀, 2 ♂( TAMU) Municipio San Cristóbal, Reserva Huitepec, Malaise trap,  8–28.viii.1990, coll. R.W. Jones.   PANAMA. 1 ♂( TAMU)  Chiriquí Prov., Lagunas del Volcán,  5 kmSW of Volcán, 8°45’52”N, 82°40’33”W,  4220 ft.(=  1285 m),  30.vii.1999, coll. J.B. Woolley.   Distribution. Mexico( Tamaulipas, San Luis Potosí, Querétaro, Veracruz, Morelos, Guerrero, Oaxaca, Chiapas), Guatemala, Honduras, El Salvador, Costa Rica, Panama, Colombia, Ecuador, Venezuela.   Biology. Gauld (1991)noticed that the colour pattern of  P. croceipesis the same as that of some very common sympatric species of  AuplopusSpinola( Hymenoptera: Pompilidae) and, in flight,  P. croceipesmay easily be confused with these species. Moreover,  P. croceipes, as far as  P. molesta, seems to share a mimicry pattern with a  Lissonotaspecies( Ichneumonidae: Banchinae) being involved in mimicry complexes characterized by black colouration with lower body parts being bright yellow ( Veijalainen et al. 2011: 66). 3411841452 [333,523,1696,1720] ANSP Mexico Orizaba 17 468 2 1 1 Veracruz de Ignacio de la Llave 3411841464 1987-12-13 UAT R. Meza Mexico 800 Cd 19 470 1 1 Tamaulipas 3411841408 [337,985,294,321] 1988-03-05 UAT R. Meza Mexico 1000 Cd 19 470 1 1 Tamaulipas 3411841417 2008-12-16 2008-12-18 2008-12-16 UAT A. I. Khalaim. Mexico Cd. 19 470 1 1 Tamaulipas 3411841373 [705,1270,331,357] 1984-05-20 2008-12-18 2008-12-16 UAT H. Ruiz Mexico Cd. 19 470 1 1 Tamaulipas 3411841430 1985-01-29 2008-12-18 2008-12-16 UAT H. Ruiz Mexico Cd. 19 470 1 1 Tamaulipas 3411841477 [589,1231,367,393] 1987-12-23 2008-12-18 2008-12-16 UAT H. & E. Ruiz Mexico Cd. 19 470 4 4 Tamaulipas 3411841305 [151,853,403,429] 2000-01-20 2008-12-18 2008-12-16 UAT E. Ruiz Mexico 1400 Cd. 19 470 5 3 2 Tamaulipas 3411841331 2007-11-07 UAT E. Ruiz Mexico 1000 Tula 19 470 1 1 Tamaulipas 3411841333 2016-10-29 2016-11-12 2016-10-29 UAT E. Ruiz Mexico 1400 Ejido Acahuales Tula 19 470 2 2 Tamaulipas 3411841439 [767,1381,475,501] 1992-05 UAT J. M. Coronado Mexico Gomez Farias 19 470 1 1 Tamaulipas 3411841392 2000-11-04 2000-11-11 2000-11-04 UAT D. R. Kasparyan. Mexico Altas Cimas Gomez Farias 19 470 1 1 Tamaulipas 3411841475 1994-07-20 1994-07-21 1994-07-20 UAT J. B. Wooley. Mexico Canindo Gomez Farias 19 470 3 3 Tamaulipas 3411841338 [809,1433,547,573] 1993-07-28 1993-07-30 1993-07-28 UAT J. B. Wooley. Mexico Canindo Gomez Farias 19 470 2 2 Tamaulipas 3411841315 [151,960,583,609] 1989-03-03 UAT R. Thompson. Mexico Rio El Cielo Gomez Farias 19 470 1 1 Tamaulipas 3411841397 2008-12-21 2008-12-23 2008-12-21 UAT A. I. Khalaim. Mexico Las Pozas Xilitla 19 470 1 1 San Luis Potosi 3411841448 [716,1291,618,645] 2013-10-04 UAQ Mexico Jalpan de Serra 19 470 2 2 Queretaro de Arteaga 3411841327 2013-10-02 UAQ Mexico Pinal de Amoles 19 470 1 1 Queretaro 3411841435 2009-02-17 2009-02-18 2009-02-17 UAT A. I. Khalaim. Mexico 1260 19.5128 Institute de Ecologia 19 470 9 3 6 Veracruz de Ignacio de la Llave 3411841443 2014-04-12 2014-04-14 2014-04-12 ZISP A. I. Khalaim. Mexico 1260 19.5128 Institute de Ecologia 19 470 18 10 8 Veracruz 3411841456 [429,1323,726,753] 1963-12-25 EMEC C. A. Toschi & M. J. Tauber. Mexico Flores Fortin 19 470 1 1 Veracruz 3411841335 1991-09-20 UAT G. Pena. Mexico Chamilpa Cuernavaca 19 470 1 1 Morelos 3411841343 2010-03-05 UAT E. Ruiz Mexico 1490 18.9975 Tepoztlan 21 -99.09861 19 470 1 1 Morelos 3411841349 2013-10-05 CIUM J. A. Hernandez Mexico 1005 18.44701 Tlaquiltenango 1 -98.98758 19 470 2 2 Morelos 3411841450 2008-07-01 2008-08-12 2008-07-01 UNAM La Esperanza, N & A. Lopez Garcia. Mexico 1600 Santiago Comaltepec 19 470 3 1 2 Oaxaca 3411841436 2009-06-06 UNAM H. Clebsch & A. Zaldivar Mexico 2332 Santiago Comaltepec 19 470 2 2 Oaxaca 3411841471 1987-07-11 TAMU R. Wharton. Mexico 1173 Highway 19 470 3 3 Oaxaca 3411841391 1997-07-20 1997-07-22 1997-07-20 UANL, UAT Reserva El Triunfo & A. Gonzalez Mexico Palo Gordo 19 470 10 10 Chiapas 3411841440 2013-03-05 2013-03-06 2013-03-05 UAT A. I. Khalaim. Mexico 1450 16.103716 National Park Lagos de Montebello 1 -91.734535 19 470 1 1 Chiapas 3411841445 2021-08-08 UAT A. I. Khalaim. Mexico 1500 16.10655 National Park Lagos de Montebello 1 -91.705154 19 470 1 1 Chiapas 3411841355 2021-08-09 UAT A. I. Khalaim. Mexico 16.132282 National Park Lagos de Montebello 1 -91.72924 19 470 1 1 Chiapas 3411841421 1990-08-11 TAMU J. C. Schaffner. Mexico Laguna Cinco Lagos Lagunas de Montebello 19 470 1 1 Chiapas 3411841395 1977-07-21 TAMU Reserva El Triunfo & J. B. Woolley. Mexico 1951 Sendero Palo Gordo 19 470 1 1 Chiapas 3411841393 1997-07-14 1997-07-23 1997-07-14 TAMU J. B. Woolley. Mexico 2018 Reserva El Triunfo 19 470 5 1 4 Chiapas 3411841314 1990-08-24 1990-08-28 1990-08-24 TAMU R. W. Jones. Mexico 2200 San 19 470 1 1 Chiapas 3411841407 1990-10-05 1990-11-25 1990-10-05 TAMU R. W. Jones. Mexico 2377 Reserva Huitepec 19 470 1 1 Chiapas 3411841323 1990-08-08 1990-08-28 1990-08-08 TAMU R. W. Jones. Mexico Reserva Huitepec San Cristobal 19 470 4 2 2 Chiapas 3411841390 [352,641,1374,1401] TAMU Panama PANAMA 19 470 1 1 Chiapas 3411841462 1999-07-30 J. B. Woolley. Panama 1286 8.764444 Lagunas del Volcan 21 -82.67583 19 470 1 Chiriqui