Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae)
Paulsen, M. J.
Insecta Mundi
2019
2019-01-18
684
1
14
48DDF
Paulsen
Paulsen
2019
[192,680,1047,1071]
Insecta
Ochodaeidae
Ochodaeus
Animalia
Coleoptera
3
2
Arthropoda
species
modopunctatus
sp. nov.
Type material. Holotypemale ( CASC; Fig. 1), labeled: a) “ MADAGASCAR: Province/ d’Antsiranana, Parc National/ Montagne d’Ambre/ el 1050 m, 25–29 Jan 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad of Sciences/ M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors / malaise trapMA-01-01C-02”; c) “CASENT / 8006869”; d) red paper “ Ochodaeus/ modopunctatus n. sp./ M.J. Paulsendet.”. Paratypemale ( MJPC) labeled a–b as holotype; c) “CASENT / 8006868”. One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre, / elev 960 m 26–29 Jan 2001”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ California Acad. of Sciences / M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors, / malaise trapMA-01-01A-02”; c) “CASENT / 8011532”. Two paratypeslabeled: a) “ MADAGASCAR: Diego- / Suarez Prov., Parc National / Montagne d’Ambre, / elev 960 m/ 26–29.i.2001” malaise”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala MA-01-01A-02 / Calif. Acad. Sci. Collection”; c) “CASENT / #” (# = 8000057 or 8000058 [³]). Two paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre, / elev 960 m, 29 January 2001”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ California Acad. of Sciences / M.E. Irwincollector, netted / by hand MA-01-01A-05”; c) “CASENT / #” (# = 8011551 [ ♀] or 8011561). Three paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre, / elev 960 m 4–19 Mar 2001”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala, / malaise trapMA-01-01A-08”; c) “CASENT / #” (# = 8016282, 8016293, or 8016298). Four paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre, / elev 960 m 5–21 April 2001”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala, / malaise trapMA-01-01A-10”; c) “CASENT / #” (# = 8013900, 8013903, 8013914, or 8013922 [ ♀]). One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre / elev 975 m 21–25 Jan 2001”; b) “ 12° 31′ S, 49° 11′ E/ California Acad. of Sciences / M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors / malaise trapMA-01-01B-01”; c) “CASENT / 8000118”. Onefemale paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Parc National / Montagne d’Ambre 975 m”; b) “ 25 Jan to 11 Feb 2001/ 12° 31′ S, 49° 11′ E/ California Acad. of Sciences / M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors / malaise trapMA-01-01B-04”; c) “CASENT / 8032442”. One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre 975 m 4–19 Mar 2001”; b) “ 12° 31′ S, 49° 11′ E/ California Acad. of Sciences / M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors / malaise trapMA-01-01B-06”; c) “CASENT / 8013823”. One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev 10 m, 6–20 Mar 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-04”; c) “CASENT / 8011513”. Two paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev 10 m, 20 Mar–7 April 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-05”; c) “CASENT / #” (# = 8011542 [³] or 8011544). Onefemale paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev 10 m, 27 April–2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-08”; c) “CASENT / 8013984”. One paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Sakalava / Beach, dwarf littoral forest / elev 10 m, 20 Mar–7 April 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-09”; c) “CASENT / 8014033”. One paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Sakalava / Beach, dwarf littoral forest / elev 10 m, 16–31 May 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-10”; c) “CASENT / 8014100”. Onefemale paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev 10 m, 7–25 May 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-12”; c) “CASENT / 8016232”. Two paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev 10 m, 25 June–6 July 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-13”; c) “CASENT / #” (# = 8013588 [ ♀] or 8013595). Onemale paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Sakalava / Beach, dwarf littoral forest / elev 10 m, 28 July–6 August 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA- 01-04B-16”; c) “CASENT / 8032553˝. One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev 10 m, 13–20 August 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA- 01-04B-17”; c) “CASENT / 8025601”. One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre 1125 m/ 29 Jan to 11 Feb 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-03”; c) “CASENT / 8016175”. One paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Parc National / Montagne d’Ambre 1125 m/ 11 Feb to 4 March 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-04”; c) “CASENT / 8014060”. Three paratypeslabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Parc National / Montagne d’Ambre 1125 m/ 19 March to 5 April 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-06”; c) “CASENT / #” (# = 8014203, 8014204 [ ♀], or 8014205 [ ♀]). One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre 1125 m/ 5–21 April 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-07”; c) “CASENT / 8006346”. Onefemale paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre 1125 m/ 21–26 April 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-08”; c) “CASENT / 8016143”. One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre 1125 m/ 12–14 May 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-10”; c) “CASENT / 8016164”. Twofemale paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre 1125 m/ 14–30 May 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-11”; c) “CASENT / #” (# = 8016113 or 8016118). Three paratypeslabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Montaigne / Francais, elev 150 m/ 15 Feb–6 March 2001”; b) “ 12° 19.5′ S, 49° 20′ E/ California Acad. of Sciences / coll: M. Irwin, R. Harin’Hala / malaise, forested limestone / ridge MA-01-06-06”; c) “CASENT / #” (# = 8032601, 8032602, or 8032603). Onefemale paratypelabeled: a) “ MADAGASCAR: Province/ Diego- Suarez, dry forest / 7 kmN of Joffreville, 360 m”; b) “ 6–20 March 2001/ 12° 20′ S, 49° 15′ E/ R. Harin’Halacollector / malaise trapMA-01-07-08”; c) “CASENT / 8014117”. One paratypelabeled: a) “ MADAGASCAR: Province/ Diego-Suarez, dry forest / 7 kmN of Joffreville, 360 m”; b) “ 7–27 April 2001/ 12° 20′ S, 49° 15′ E/ R. Harin’Halacollector / malaise trapMA-01-07-10”; c) “CASENT / 8025772”. One paratypelabeled: a) “ MADAGASCAR: Province/ d’Antsiranana, dry forest / 7 kmN of Joffreville, 360 m”; b) “ 27 April–13 May 2001/ 12° 20′ S, 49° 15′ E/ R. Harin’Halacollector / malaise trapMA-01-07-11”; c) “CASENT / 8011458”. All paratypes( n= 43) with label: on yellow paper, “ Ochodaeus/ modopunctatus/ Paulsen / PARATYPE”.
Description. Holotypemale ( Fig. 1a). Coleoptera: Scarabaeoidea: Ochodaeidae. Length: 9.9 mm. Width: 4.1 mm. Color:Everywhere uniformly dark reddish brown; surface nitid. Head:Surface lacking tubercles, irregularly punctate; punctures moderately large, setose, separated by ~ 4 puncture diameters. Eyes large, globose; lacking ventral projection of canthus. Antennal club subequal to eye in size. Mandibles broadly scooped, basal angle prominent, apices falcate; left mandible with large rectangular internal tooth; right mandible with complimenting tooth reduced, second tooth large, obtuse. Frons weakly convex. Clypeus flat with disc short (at most 1/5 as long as wide), anterior margin strongly beaded; bead thick, rounded, almost as long as disc. Mentum ( Fig. 4) slightly wider than long, tumid (not flat); most strongly tumid near base, decreasing anteriorly; surface with 2 lateral and 1 median depression near base, median depression contiguous with oval fovea on disc; fovea continuing anteriorly. Pronotum:Form convex. Surface densely punctate, appearing smooth between punctures, lacking distinct tubercles (at 40× minute tubercle anterior to seta); punctation mixed, with large setose punctures and smaller naked punctures interspersed. Elytra:Form convex, relatively elongate, elytra together longer than wide. Surface with striae moderately impressed, punctate, surface smooth between punctures (except sutural stria with connecting groove); strial punctures separated by> 5 diameters, lacking setae. Intervals irregularly punctate; punctures small, setose, with minute tubercle anteriorly. Legs:Protibia tridentate externally, with short, acute pollex directed anteriorly ( Fig. 1b). Profemur and mesofemur with apical tooth reduced to rounded lobe. Metafemur with apical fin-like projection with weak tooth in distal fourth. Venter/Abdomen:Metasternum and abdomen sparsely punctate; punctures large with long, erect, golden setae. Pygidium punctate, setose. Stridulatory peg with narrow neck and rounded apex ( Fig. 10). Male genitalia:Sclerotized patch on internal sac between parameres broadly oblong with bifurcate end ( Fig. 7). Paratypevariation.The paratypesshow little variation from the holotype, and negligible sexual dimorphism. The clypeus may appear more trapezoidal (truncate medially), and although this is a common sexual dimorphism in ochodaeids this does not appear to be sexually diagnostic in O. modopunctatus. The subapical tooth on the metafemur is present in the holotypeand the three largest males but is obsolete in all smaller males and females.
Remarks.Due to the lack of any useful sexual dimorphism in the specimens that had open pygidia with visible genitalia, the majority of the remaining specimens were not subjected to potential damage during genitalic dissection. The sex is indicated only for paratypeswith everted genitalia.
Etymology.The name is constructed from the Latin adverb ‘modo’ meaning ‘only’ and the adjective for ‘punctate’. This refers to the apparent lack of tubercles on the pronotum of this species, which is unique among the Madagascan species. The other species treated below have at least weak indications of tubercles between punctures, and the known taxa all have a distinctly tuberculate pronotum.
Distribution.This species is thus far known only from the extreme northern tip of the island in the vicinity of the typelocality, Montagne d’Ambre ( Fig. 19).
2564843961
2001-01-25
2001-01-29
2001-01-25
CASC, R
M. E. Irwin & E. I. Schlinger
Madagascar
California Acad of Sciences
M. J. Paulsen
1050
-12.520278
Province
21
49.17917
Montagne d'Ambre
3
2
1
holotype
2564843948
[232,1124,1225,1250]
MJPC
3
2
2
paratype
2564843920
2001-01-26
2001-01-29
2001-01-26
R
M. E. Irwin & E. I. Schlinger
Madagascar
960
-12.514444
One
21
49.18139
3
2
1
paratype
2564843929
2001-01-26
2001-01-29
2001-01-26
R
M. E. Irwin & E. I. Schlinger
Madagascar
960
-12.514444
Two
21
49.18139
3
2
1
paratype
2564843925
2001-01-29
M. E. Irwin
Madagascar
960
-12.514444
Two
21
49.18139
3
2
1
1
paratype
2564843922
2001-03-04
2001-03-19
2001-03-04
R
M. Irwin
Madagascar
960
-12.514444
Three
21
49.18139
3
2
1
paratype
2564843953
2001-04-05
2001-04-21
2001-04-05
R
M. Irwin
Madagascar
960
-12.514444
Four
21
49.18139
3
2
1
1
paratype
2564843913
2001-01-21
2001-01-25
2001-01-21
R
M. E. Irwin & E. I. Schlinger
Madagascar
975
-12.516666
One
1293
49.183334
3
2
1
paratype
2564843966
2001-01-25
2001-02-11
2001-01-25
R
M. E. Irwin & E. I. Schlinger
Madagascar
975
-12.516666
One
1293
49.183334
3
2
1
paratype
2564843933
2001-03-04
2001-03-19
2001-03-04
R
M. E. Irwin & E. I. Schlinger
Madagascar
975
4
3
-12.516666
One
1293
49.183334
3
2
1
paratype
2564843910
2001-03-06
2001-03-20
2001-03-06
R
M. Irwin
Madagascar
10
-12.262777
One
21
49.397503
4
3
1
paratype
2564843919
2001-03-20
2001-04-07
2001-03-20
R
M. Irwin
Madagascar
10
-12.262777
Two
21
49.397503
4
3
1
paratype
2564843914
2001-04-27
R
M. Irwin
Madagascar
10
-12.262777
One
21
49.397503
4
3
1
paratype
2564843950
2001-03-20
2001-04-07
2001-03-20
R
M. Irwin
Madagascar
10
-12.262777
One
21
49.397503
4
3
1
paratype
2564843947
2001-05-16
2001-05-31
2001-05-16
R
M. Irwin
Madagascar
10
-12.262777
One
21
49.397503
4
3
1
paratype
2564843918
2001-05-07
2001-05-25
2001-05-07
R
M. Irwin
Madagascar
10
-12.262777
One
21
49.397503
4
3
1
paratype
2564843945
2001-06-25
2001-07-06
2001-06-25
R
M. Irwin
Madagascar
10
-12.262777
Two
21
49.397503
4
3
1
1
paratype
2564843943
2001-07-28
2001-08-06
2001-07-28
R
M. Irwin
Madagascar
10
-12.262777
One
21
49.397503
4
3
1
paratype
2564843964
2001-08-13
2001-08-20
2001-08-13
R
M. Irwin
Madagascar
10
-12.262777
One
21
49.397503
4
3
1
paratype
2564843923
2001-01-29
2001-02-11
2001-01-29
R
M. Irwin
Madagascar
1125
-12.520278
One
21
49.17917
4
3
1
paratype
2564843944
2001-02-11
2001-03-04
2001-02-11
R
M. Irwin
Madagascar
1125
-12.520278
One
21
49.17917
4
3
1
paratype
2564843958
2001-03-19
2001-04-05
2001-03-19
R
M. Irwin
Madagascar
1125
-12.520278
Three
21
49.17917
4
3
2
2
paratype
2564843951
2001-04-05
2001-04-21
2001-04-05
R
M. Irwin
Madagascar
1125
-12.520278
One
21
49.17917
4
3
1
paratype
2564843909
2001-04-21
2001-04-26
2001-04-21
R
M. Irwin
Madagascar
1125
-12.520278
One
21
49.17917
4
3
1
paratype
2564843935
2001-05-12
2001-05-14
2001-05-12
R
M. Irwin
Madagascar
1125
-12.520278
One
21
49.17917
4
3
1
paratype
2564843936
2001-05-14
2001-05-30
2001-05-14
R
M. Irwin
Madagascar
1125
-12.520278
Two
21
49.17917
4
3
1
paratype
2564843957
2001-02-15
2001-03-06
2001-02-15
R
M. Irwin
Madagascar
150
5
4
-12.325
Three
909
49.333332
4
3
1
paratype
2564843912
2001-03-06
2001-03-20
2001-03-06
R
Harin'Hala
Madagascar
Suarez
360
-12.333333
One
1293
49.25
Province
5
4
1
paratype
2564843940
2001-04-07
2001-04-27
2001-04-07
R
Harin'Hala
Madagascar
Diego-Suarez
360
-12.333333
One
1293
49.25
Province
5
4
1
paratype
2564843915
2001-04-27
2001-05-13
2001-04-27
R
Harin'Hala
Madagascar
d'Antsiranana
360
-12.333333
One
1293
49.25
Province
5
4
1
paratype