Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae) Paulsen, M. J. Insecta Mundi 2019 2019-01-18 684 1 14 48DDF Paulsen Paulsen 2019 [192,680,1047,1071] Insecta Ochodaeidae Ochodaeus Animalia Coleoptera 3 2 Arthropoda species modopunctatus sp. nov.   Type material.  Holotypemale ( CASC; Fig. 1), labeled: a) “ MADAGASCAR: Province/ d’Antsiranana, Parc National/ Montagne d’Ambre/ el  1050 m,  25–29 Jan 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad of Sciences/ M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors / malaise trapMA-01-01C-02”; c) “CASENT / 8006869”; d) red paper “  Ochodaeus/  modopunctatus n. sp./ M.J. Paulsendet.”.   Paratypemale ( MJPC) labeled a–b as holotype; c) “CASENT / 8006868”.  One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre, / elev  960 m  26–29 Jan 2001”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ California Acad. of Sciences / M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors, / malaise trapMA-01-01A-02”; c) “CASENT / 8011532”.  Two paratypeslabeled: a) “ MADAGASCAR: Diego- / Suarez Prov., Parc National / Montagne d’Ambre, / elev  960 m/  26–29.i.2001” malaise”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala MA-01-01A-02 / Calif. Acad. Sci. Collection”; c) “CASENT / #” (# = 8000057 or 8000058 [³]).  Two paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre, / elev  960 m,  29 January 2001”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ California Acad. of Sciences / M.E. Irwincollector, netted / by hand MA-01-01A-05”; c) “CASENT / #” (# = 8011551 [ ♀] or 8011561).  Three paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre, / elev  960 m  4–19 Mar 2001”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala, / malaise trapMA-01-01A-08”; c) “CASENT / #” (# = 8016282, 8016293, or 8016298).  Four paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre, / elev  960 m  5–21 April 2001”; b) “ 12° 30′ 52″ S, 49° 10′ 53″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala, / malaise trapMA-01-01A-10”; c) “CASENT / #” (# = 8013900, 8013903, 8013914, or 8013922 [ ♀]).  One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre / elev  975 m  21–25 Jan 2001”; b) “ 12° 31′ S, 49° 11′ E/ California Acad. of Sciences / M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors / malaise trapMA-01-01B-01”; c) “CASENT / 8000118”.  Onefemale paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Parc National / Montagne d’Ambre  975 m”; b) “  25 Jan to 11 Feb 2001/ 12° 31′ S, 49° 11′ E/ California Acad. of Sciences / M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors / malaise trapMA-01-01B-04”; c) “CASENT / 8032442”.  One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre  975 m  4–19 Mar 2001”; b) “ 12° 31′ S, 49° 11′ E/ California Acad. of Sciences / M.E. Irwin, E.I. Schlinger/ & R. Harin’Hala collectors / malaise trapMA-01-01B-06”; c) “CASENT / 8013823”.  One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev  10 m,  6–20 Mar 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-04”; c) “CASENT / 8011513”.  Two paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev  10 m,  20 Mar–7 April 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-05”; c) “CASENT / #” (# = 8011542 [³] or 8011544).  Onefemale paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev  10 m,  27 April–2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-08”; c) “CASENT / 8013984”.  One paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Sakalava / Beach, dwarf littoral forest / elev  10 m,  20 Mar–7 April 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-09”; c) “CASENT / 8014033”.  One paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Sakalava / Beach, dwarf littoral forest / elev  10 m,  16–31 May 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-10”; c) “CASENT / 8014100”.  Onefemale paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev  10 m,  7–25 May 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-12”; c) “CASENT / 8016232”.  Two paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev  10 m,  25 June–6 July 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA-01-04B-13”; c) “CASENT / #” (# = 8013588 [ ♀] or 8013595).  Onemale paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Sakalava / Beach, dwarf littoral forest / elev  10 m,  28 July–6 August 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA- 01-04B-16”; c) “CASENT / 8032553˝.  One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Sakalava / Beach, dwarf littoral forest / elev  10 m,  13–20 August 2001”; b) “ 12° 15′ 46″ S, 49° 23′ 51″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trap- across sandy / trail MA- 01-04B-17”; c) “CASENT / 8025601”.  One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre  1125 m/  29 Jan to 11 Feb 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-03”; c) “CASENT / 8016175”.  One paratypelabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Parc National / Montagne d’Ambre  1125 m/  11 Feb to 4 March 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-04”; c) “CASENT / 8014060”.  Three paratypeslabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Parc National / Montagne d’Ambre  1125 m/  19 March to 5 April 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-06”; c) “CASENT / #” (# = 8014203, 8014204 [ ♀], or 8014205 [ ♀]).  One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre  1125 m/  5–21 April 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-07”; c) “CASENT / 8006346”.  Onefemale paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre  1125 m/  21–26 April 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-08”; c) “CASENT / 8016143”.  One paratypelabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre  1125 m/  12–14 May 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-10”; c) “CASENT / 8016164”.  Twofemale paratypeslabeled: a) “ MADAGASCAR: Province / d’Antsiranana, Parc National / Montagne d’Ambre  1125 m/  14–30 May 2001”; b) “ 12° 31′ 13″ S, 49° 10′ 45″ E/ California Acad. of Sciences / Coll: M. Irwin, R. Harin’Hala / malaise trapMA-01-01D-11”; c) “CASENT / #” (# = 8016113 or 8016118).  Three paratypeslabeled: a) “ MADAGASCAR: Province / Diego-Suarez, Montaigne / Francais, elev  150 m/  15 Feb–6 March 2001”; b) “ 12° 19.5′ S, 49° 20′ E/ California Acad. of Sciences / coll: M. Irwin, R. Harin’Hala / malaise, forested limestone / ridge MA-01-06-06”; c) “CASENT / #” (# = 8032601, 8032602, or 8032603).  Onefemale paratypelabeled: a) “ MADAGASCAR: Province/ Diego- Suarez, dry forest /  7 kmN of Joffreville,  360 m”; b) “  6–20 March 2001/ 12° 20′ S, 49° 15′ E/ R. Harin’Halacollector / malaise trapMA-01-07-08”; c) “CASENT / 8014117”.  One paratypelabeled: a) “ MADAGASCAR: Province/ Diego-Suarez, dry forest /  7 kmN of Joffreville,  360 m”; b) “  7–27 April 2001/ 12° 20′ S, 49° 15′ E/ R. Harin’Halacollector / malaise trapMA-01-07-10”; c) “CASENT / 8025772”.  One paratypelabeled: a) “ MADAGASCAR: Province/ d’Antsiranana, dry forest /  7 kmN of Joffreville,  360 m”; b) “  27 April–13 May 2001/ 12° 20′ S, 49° 15′ E/ R. Harin’Halacollector / malaise trapMA-01-07-11”; c) “CASENT / 8011458”. All paratypes( n= 43) with label: on yellow paper, “  Ochodaeus/  modopunctatus/ Paulsen / PARATYPE”.   Description. Holotypemale ( Fig. 1a). Coleoptera: Scarabaeoidea: Ochodaeidae. Length: 9.9 mm. Width: 4.1 mm. Color:Everywhere uniformly dark reddish brown; surface nitid. Head:Surface lacking tubercles, irregularly punctate; punctures moderately large, setose, separated by ~ 4 puncture diameters. Eyes large, globose; lacking ventral projection of canthus. Antennal club subequal to eye in size. Mandibles broadly scooped, basal angle prominent, apices falcate; left mandible with large rectangular internal tooth; right mandible with complimenting tooth reduced, second tooth large, obtuse. Frons weakly convex. Clypeus flat with disc short (at most 1/5 as long as wide), anterior margin strongly beaded; bead thick, rounded, almost as long as disc. Mentum ( Fig. 4) slightly wider than long, tumid (not flat); most strongly tumid near base, decreasing anteriorly; surface with 2 lateral and 1 median depression near base, median depression contiguous with oval fovea on disc; fovea continuing anteriorly. Pronotum:Form convex. Surface densely punctate, appearing smooth between punctures, lacking distinct tubercles (at 40× minute tubercle anterior to seta); punctation mixed, with large setose punctures and smaller naked punctures interspersed. Elytra:Form convex, relatively elongate, elytra together longer than wide. Surface with striae moderately impressed, punctate, surface smooth between punctures (except sutural stria with connecting groove); strial punctures separated by> 5 diameters, lacking setae. Intervals irregularly punctate; punctures small, setose, with minute tubercle anteriorly. Legs:Protibia tridentate externally, with short, acute pollex directed anteriorly ( Fig. 1b). Profemur and mesofemur with apical tooth reduced to rounded lobe. Metafemur with apical fin-like projection with weak tooth in distal fourth. Venter/Abdomen:Metasternum and abdomen sparsely punctate; punctures large with long, erect, golden setae. Pygidium punctate, setose. Stridulatory peg with narrow neck and rounded apex ( Fig. 10). Male genitalia:Sclerotized patch on internal sac between parameres broadly oblong with bifurcate end ( Fig. 7).   Paratypevariation.The paratypesshow little variation from the holotype, and negligible sexual dimorphism. The clypeus may appear more trapezoidal (truncate medially), and although this is a common sexual dimorphism in ochodaeids this does not appear to be sexually diagnostic in  O. modopunctatus. The subapical tooth on the metafemur is present in the holotypeand the three largest males but is obsolete in all smaller males and females.   Remarks.Due to the lack of any useful sexual dimorphism in the specimens that had open pygidia with visible genitalia, the majority of the remaining specimens were not subjected to potential damage during genitalic dissection. The sex is indicated only for paratypeswith everted genitalia.   Etymology.The name is constructed from the Latin adverb ‘modo’ meaning ‘only’ and the adjective for ‘punctate’. This refers to the apparent lack of tubercles on the pronotum of this species, which is unique among the Madagascan species. The other species treated below have at least weak indications of tubercles between punctures, and the known taxa all have a distinctly tuberculate pronotum.   Distribution.This species is thus far known only from the extreme northern tip of the island in the vicinity of the typelocality, Montagne d’Ambre ( Fig. 19). 2564843961 2001-01-25 2001-01-29 2001-01-25 CASC, R M. E. Irwin & E. I. Schlinger Madagascar California Acad of Sciences M. J. Paulsen 1050 -12.520278 Province 21 49.17917 Montagne d'Ambre 3 2 1 holotype 2564843948 [232,1124,1225,1250] MJPC 3 2 2 paratype 2564843920 2001-01-26 2001-01-29 2001-01-26 R M. E. Irwin & E. I. Schlinger Madagascar 960 -12.514444 One 21 49.18139 3 2 1 paratype 2564843929 2001-01-26 2001-01-29 2001-01-26 R M. E. Irwin & E. I. Schlinger Madagascar 960 -12.514444 Two 21 49.18139 3 2 1 paratype 2564843925 2001-01-29 M. E. Irwin Madagascar 960 -12.514444 Two 21 49.18139 3 2 1 1 paratype 2564843922 2001-03-04 2001-03-19 2001-03-04 R M. Irwin Madagascar 960 -12.514444 Three 21 49.18139 3 2 1 paratype 2564843953 2001-04-05 2001-04-21 2001-04-05 R M. Irwin Madagascar 960 -12.514444 Four 21 49.18139 3 2 1 1 paratype 2564843913 2001-01-21 2001-01-25 2001-01-21 R M. E. Irwin & E. I. Schlinger Madagascar 975 -12.516666 One 1293 49.183334 3 2 1 paratype 2564843966 2001-01-25 2001-02-11 2001-01-25 R M. E. Irwin & E. I. Schlinger Madagascar 975 -12.516666 One 1293 49.183334 3 2 1 paratype 2564843933 2001-03-04 2001-03-19 2001-03-04 R M. E. Irwin & E. I. Schlinger Madagascar 975 4 3 -12.516666 One 1293 49.183334 3 2 1 paratype 2564843910 2001-03-06 2001-03-20 2001-03-06 R M. Irwin Madagascar 10 -12.262777 One 21 49.397503 4 3 1 paratype 2564843919 2001-03-20 2001-04-07 2001-03-20 R M. Irwin Madagascar 10 -12.262777 Two 21 49.397503 4 3 1 paratype 2564843914 2001-04-27 R M. Irwin Madagascar 10 -12.262777 One 21 49.397503 4 3 1 paratype 2564843950 2001-03-20 2001-04-07 2001-03-20 R M. Irwin Madagascar 10 -12.262777 One 21 49.397503 4 3 1 paratype 2564843947 2001-05-16 2001-05-31 2001-05-16 R M. Irwin Madagascar 10 -12.262777 One 21 49.397503 4 3 1 paratype 2564843918 2001-05-07 2001-05-25 2001-05-07 R M. Irwin Madagascar 10 -12.262777 One 21 49.397503 4 3 1 paratype 2564843945 2001-06-25 2001-07-06 2001-06-25 R M. Irwin Madagascar 10 -12.262777 Two 21 49.397503 4 3 1 1 paratype 2564843943 2001-07-28 2001-08-06 2001-07-28 R M. Irwin Madagascar 10 -12.262777 One 21 49.397503 4 3 1 paratype 2564843964 2001-08-13 2001-08-20 2001-08-13 R M. Irwin Madagascar 10 -12.262777 One 21 49.397503 4 3 1 paratype 2564843923 2001-01-29 2001-02-11 2001-01-29 R M. Irwin Madagascar 1125 -12.520278 One 21 49.17917 4 3 1 paratype 2564843944 2001-02-11 2001-03-04 2001-02-11 R M. Irwin Madagascar 1125 -12.520278 One 21 49.17917 4 3 1 paratype 2564843958 2001-03-19 2001-04-05 2001-03-19 R M. Irwin Madagascar 1125 -12.520278 Three 21 49.17917 4 3 2 2 paratype 2564843951 2001-04-05 2001-04-21 2001-04-05 R M. Irwin Madagascar 1125 -12.520278 One 21 49.17917 4 3 1 paratype 2564843909 2001-04-21 2001-04-26 2001-04-21 R M. Irwin Madagascar 1125 -12.520278 One 21 49.17917 4 3 1 paratype 2564843935 2001-05-12 2001-05-14 2001-05-12 R M. Irwin Madagascar 1125 -12.520278 One 21 49.17917 4 3 1 paratype 2564843936 2001-05-14 2001-05-30 2001-05-14 R M. Irwin Madagascar 1125 -12.520278 Two 21 49.17917 4 3 1 paratype 2564843957 2001-02-15 2001-03-06 2001-02-15 R M. Irwin Madagascar 150 5 4 -12.325 Three 909 49.333332 4 3 1 paratype 2564843912 2001-03-06 2001-03-20 2001-03-06 R Harin'Hala Madagascar Suarez 360 -12.333333 One 1293 49.25 Province 5 4 1 paratype 2564843940 2001-04-07 2001-04-27 2001-04-07 R Harin'Hala Madagascar Diego-Suarez 360 -12.333333 One 1293 49.25 Province 5 4 1 paratype 2564843915 2001-04-27 2001-05-13 2001-04-27 R Harin'Hala Madagascar d'Antsiranana 360 -12.333333 One 1293 49.25 Province 5 4 1 paratype