Scolytodes Ferrari (Coleoptera, Scolytinae) from Ecuador: 40 new species, and a molecular phylogenetic guide to infer species differences Jordal, Bjarte H. Smith, Sarah M. Zootaxa 2020 2020-07-14 4813 1 1 67 Jordal and Smith Jordal and Smith 2020 [151,605,1125,1152] Insecta Curculionidae Scolytodes Animalia Coleoptera 55 56 Arthropoda species sparsus sp. nov.  ( Figs 120, 123, 126)   Type material.   Holotype, female: Ecuador: Napo province, Cosanga, Yanayacu station,  2100 m, GIS: -0.594, - 77.877, HG-vapor light, #756,  3iii2018, R. Osborn, leg. The holotypeis deposited in QCAZ.   Diagnosis.Interstriae 10 sharply carinate to near apex. Protibiae without additional mesal tooth, tooth 1 slightly larger than tooth 2. Elytra with large punctures in irregular rows. Related to  S. nitidus(Eggers, 1928), but distinguished by the larger strial punctures placed in less regular rows, and by the female frons having a smaller impressed area with fine setae.   Description, female.Length 2.4 mm, 2.2 × as long as wide; colour dark brown. Head.Eyes slightly sinuate, separated above by c. 2.5–3 × their width (not fully exposed). Frons convex above, impressed on a semi-circular area from epistoma to just above level of antennal insertion; surface with large, deep punctured above, and much denser and smaller punctures in impressed area, except for a longitudinal impunctate field dividing impressed area in two halves; vestiture consisting of short, fine setae in impressed area. Antennal club pilose, sutures obscure. Funiculus possibly 6-segmented. Pronotumreticulate, dull, punctures spaced by 0.5– 1 × their diameter, reaching anterior margin; anterior half with low asperities or soft wrinkles, laterally forming near contiguous lines; lateral carinae anteriorly turned slightly medially before termination. Glabrous (0-0-0). Elytrashiny, surface slightly rugose; striae more or less regular, not impressed, punctures large, deep, separated by 1–1.5 × their diameter; interstriae 3–4 × broader than striae, punctures almost as large as strial punctures, in part confused. Interstriae 10 sharply raised to near apex. Glabrous. Legs.Procoxae and mesocoxae separated by the width of one procoxa. Protibiae narrow, distal teeth 1 larger than 2, with 3–4 additional small spines or transverse rugae towards tibial base; protibial mucro short, blunt, curved laterally. Meso- and metatibiae with 8–9 and 7–8 socketed teeth on distal two-thirds and half, respectively. Ventral vestiture. Setae on mesanepisternum plumose, on metanepisternum simple to trifid, on metasternum long, largely bifid or simple.   FIGURES 118–126.Dorsal, lateral and frontal views of the female holotype of  Scolytodes arcuatus(118, 121, 124),  Scolytodes validus(119, 122, 125), and  Scolytodes sparsus(120, 123, 126).  Male.Not known.   Key ( Wood 2007).Keys to couplet 33,  S. ovalis(Eggers, 1940), but differs by the larger strial and interstrial punctures. More closely related to  Scolytodes nitiduswhich is erroneously placed in couplet 44; females of that species have no setae reaching upper level of eyes.   Etymology.The Latin name  sparsusis a masculine participle, meaning spotted or freckled, referring to irregular placement of large punctures on the elytra.   Biology and distribution. Only known from the high-altitude typelocality in Ecuador. A single specimen was collected a mercury-light trap. 2018-03-03 QCAZ R. Osborn & The Ecuador 2100 -0.594 Yanayacu station 78 -77.877 Cosanga 55 56 2 Napo holotype Ecuador Ecuador 57 58 1